A catalogue of Japanese Trichoptera
Family Hydropsychidae Curtis
Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last updated: 23 Feb. 2024
Subfamily Arctopsychinae Martynov
Two genera, Arctopsyche McLachlan and Parapsyche Betten, are known from Japan.
Genus Arctopsyche McLachlan
Only two named species have been known from Japan, but four unnamed species
accompanied with an alphabetic code have been described based only on the
larval stage. Arctopsyche hirayamai described by Matsumura (1931) was transfered to the genus Plectrocnemia (Polycentropodidae) by Ohkawa and Ito (2007).
Arctopsyche amurensis Martynov 1934
Arctopsyche amurensis Martynov 1934, 247, 250, male, Russia (Amur).
Schmid 1968, 59-60, Male.
Yamanouchi 1980, 7, Hokkaido (Tokachi).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Muramatsu 2008, 5, Hokkaido (Okhotsk).
Ito 2013, 55, Hokkaido (Kushiro).
Ijima and Tsuchiya 2014, 29, Hokkaido (Okhotsk, Kushiro).
Ito and Kuhara 2016, 16-17, Hokkaido (Okhotsk, Kushiro).
Tanida 2016, 94.
Muramatsu 2022, 21, male, female.
Ito and Kuhara 2023, 54, Hokkaido (Kamikawa,
Type locality. Amur, Russia.
Distribution. Japan (Hokkaido), Russia.
Remarks. Tanida (2016) mistakenly listed Honshu, Shikoku and Tsushima as the distribution
area of this species (Tanida 2018).
Arctopsyche spinifera Ulmer 1907
Arctopsyche spinifera Ulmer 1907a, 70-71, pl. 4, male, Honshu (Gifu).
Matsumura 1931, 1132, adult (spinigera).
Matsumura 1933, 7, pl. 2, adult (spinigera).
Fischer 1963, 222.
Kobayashi 1968, 3, Honshu (Niigata).
Schmid 1968, 50-51, male, Honshu (Fukushima, Nagano).
Tani 1978, 4-5, male, Honshu (Niigata).
Morita 1988, 5, Honshu (Mie).
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 39, Honshu (Kanagawa).
Morita 1989, 39, Honshu (Mie).
Tanida et al. 1991, 46, adult, pupa, larva.
Morita 1993, 4, Honshu (Mie).
Moriya 1994, 71, Honshu (Yamanashi).
Kuranishi and Ohtaka 1996, 14, Honshu (Aomori).
Moriya 1997, 62, Honshu (Yamanashi).
Nozaki 1997a, 35, Honshu (Kanagawa).
Kagaya et al. 1998, 26, 50-51, 155, Honshu (Tokyo, Yamanashi).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338, Honshu (Mie).
Morita 2003, 25, Honshu (Mie).
Nozaki 2004, 1231, Honshu (Kanagawa).
Katsuma 2005a, 4, Honshu (Ibaraki).
Katsuma 2005b, 13, Honshu (Gumma).
Nozaki 2005, 108, adult, larva.
Mey and Nozaki 2006, 24, Honshu (Nagano).
Torii and Hattori 2006, 37, Honshu (Shizuoka).
Morita 2007a, 14, Honshu (Mie).
Tanida 2008, 249, pl. 92, adult, Honshu (Iwate).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Morita 2009a, 140, Honshu (Mie).
Katsuma 2010b, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 41, Honshu (Shiga).
Inazu and Nishida 2011, 189, male, Honshu (Hyogo).
Katsuma 2011c, 67, Honshu (Ibaraki).
Katsuma 2012b, 65, Honshu (Ibaraki).
Kawase and Morita 2014, 5, Honshu (Shiga).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Nozaki 2016, 75, 345-347, 435, Honshu (Yamanashi).
Tanaka 2016a, 22, 24, male, Honshu (Akita).
Tanida 2016, 94.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kawase et al. 2017, 30, Honshu (Shiga).
Nojima 2017, 118, Honshu (Okayama).
Tanaka 2017, 2, Honshu (Akita).
Ohira and Tsutsumi 2018, 24, Honshu (Fukushima).
Tanida 2018, 567, 570, male.
Tanida et al. 2018, 30, Honshu (Toyama).
Tanaka 2019a 27, Honshu (Akita).
Tanaka 2019b, 12, Honshu (Akita).
Morita 2020d, 81, Honshu (Mie).
Tanaka 2022, 35, Honshu (Akita).
Tanaka 2023, 41, Honshu (Akita).
Tanaka 2024, 26, Honshu (Akita).
Type locality. Honshu (Gifu), Japan.
Distribution. Japan (Honshu).
Remarks. Tanida (2016) included Hokkaido, Shikoku and Tsushima as the distribution area of this species, but no reliable records from these areas are found.
Arctopsyche sp. AA
Arctopsyche sp. A: Tsuda 1937, 302-303, larva, Honshu (Nagano, Kyoto) (larval record).
Akagi 1956a, 22-23, larva, Honshu (Tochigi, Shiga) (larval record).
Tsuda and Akagi 1962, 132, larva.
Arctopsyche sp. AA: Tanida, 1985, 178-179, larva.
Tanida 2005, 479-480, larva.
Inazu and Nishida 2011, larva.
Distribution. Japan (Honshu).
Remarks. This species and Arctopsyche sp. AD are probably the larval stage of Arctopsyche spinifera (Kagaya et al. 1998; Tanida 2018).
Arctopsyche sp. AB
Arctopsyche sp. AB: Kim 1974, 70-71, male and female genitalia, Honshu (Nara).
Distribution. Japan (Honshu).
Arctopsyche sp. AD
Arctopsyche sp. D: Akagi 1956a, 20, 22-23, larva, Honshu (Tochigi, Shiga, Kyoto, Nara).
Tsuda and Akagi 1962, 132, larva.
Arctopsyche sp. AD: Tanida, 1985, 177-178, larva.
Tanida 2005, 479-480, larva.
Inazu and Nishida 2011, 189, larva, Honshu (Hyogo).
Distribution. Japan (Honshu).
Remarks. This species and Arctopsyche sp. AA are probably the larval stage of Arctopsyche spinifera (Kagaya et al. 1998).
Arctopsyche sp. AE
Arctopsyche sp. E: Tani 1967, 12, 14, larva, Hokkaido (Okhotsk, Tokachi, Kamikawa) (larval record).
Arctopsyche sp. AE: Tanida 1985, 178-179, larva.
Tanida 2005, 479-480, larva.
Distribution. Japan (Hokkaido).
Remarks. This species is probably the larval stage of Arctopsyche amurensis (Tanida 2018).
Arctopsyche sp. C
Arctopsyche sp. C: Akagi 1956a, 20-211, larva, Honshu (Tochigi: larval record).
Distribution. Japan (Honshu).
Remarks. Tanida (1985) suggested that this species is probably an early instar
larva of Arctopsyche sp. AA.
Genus Parapsyche Betten
Four named species are recognized for Japanese fauna, but the revisional
study is needed for Japanese species (Nozaki et al. 2014).
Parapsyche aureocephala Schmid 1964
Parapsyche aureocephla Schmid 1964, 826-827, male, Honshu (Nagano).
Schmid 1968, 74, 77, 79, female.
Nozaki 1988b, 39, Honshu (Kanagawa).
Morita 1989, 39, Honshu (Mie).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Kuranishi 1991, 149, 152, oviposition, Honshu (Ibaraki).
Morita 1993, 3, Honshu (Mie).
Moriya 1994, 71, Honshu (Kanagawa).
Moriya 1997, 62, Honshu (Yamanashi).
Nozaki 1997a, 35, Honshu (Kanagawa)
Kagaya et al. 1998, 26, 51, 157, Honshu (Tokyo, Yamanashi).
Uchida 2002, 77, Honshu (Tochigi).
Kawase et al. 2004, 111, Honshu (Ibaraki).
Katsuma 2005b, 13, Honshu (Gumma).
Tanida 2005, 478, 481, male, female.
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Morita and Kawase 2008, 55, Honshu (Ishikawa).
Morita 2009a, 140, Honshu (Mie).
Kawase and Hayashi 2010b, 85, Honshu (Shimane).
Kawase and Morita 2010, 41, Honshu (Shiga).
Mano 2010, 22-23, Honshu (Aichi).
Katsuma 2011b, 49, 51, Honshu (Ibaraki).
Inazu 2012, 7, Honshu (Hyogo).
Morita and Kawase 2012, 14, Honshu (Ishikawa).
Inazu 2013b, 72, Honshu (Hyogo).
Morita 2013, 63, Honshu (Mie).
Morita and Kawase 2016, 21, Honshu (Ishikawa).
Tanida 2016, 94.
Kawase et al. 2017, 30, Honshu (Shiga).
Nojima 2017, 119, Honshu (Okayama).
Tanida 2018, 567, 570, male.
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu).
Remarks. Tanida (2016) listed Hokkaido and Shikoku as the distribution area of
this species, but no reliable records from these areas are found.
Parapsyche kurosawai (Kobayashi 1956)
Arctopsyche kurosawai Kobayashi 1956, 83-85, male, Honshu (Yamanashi).
Parapsyche kurosawai: Schmid 1968, 62, 79, combination.
Kobayashi 1984c, 4-5, Sado.
Tanida 2016, 94.
Type locality. Honshu (Yamanashi), Japan.
Distribution. Japan (Honshu, Sado).
Parapsyche maculata (Ulmer 1907)
Arctopsyche maculata Ulmer 1907a, 71-72, pl. 4, male, Oiwake (Nagano?).
Matsumura 1931, 1131-1132, adult.
Uéno 1931, 109, Honshu (Nagano).
Kuwayama 1932, 1512, adult.
Matsumura 1933, 7, pl. 2, adult.
Uéno 1935, 82, Honshu (Nagano).
Tsuda 1937, 301-302, larva, Honshu (Nagano, Kyoto).
Esaki et al. 1938, 135, pl. 61, adult.
Tsuda 1942a, 276-277, Honshu (Nagano, Kyoto).
Tsuda 1943, 164-165, 169, Honshu (Kyoto).
Tsuda 1950, 416, adult.
Ishihara et al. 1953, 37, Shikoku (Ehime).
Takeuchi 1955, 76, pl. 28, adult, Kyushu (Oita).
Akagi 1956a, 19-21, larva, Honshu (Kyoto, Nara).
Gyotoku 1957, 10, Kyushu (Fukuoka).
Tsuda 1959, 139, larva.
Tsuda and Akagi 1962, 131-132, larva.
Fischer 1963, 221-222.
Kuwada 1965, 164, pl. 82, adult, Shikoku (Ehime)..
Kobayashi 1968, 3, Honshu (Niigata).
Parapsyche nigrocephala Schmid 1964, 826-828, male, Honshu (Nara: Yoshino). Synonymized by Schmid (1968).
Parapsyche maculata: Schmid 1968, 73, 79.
Tani 1977, 195, pl. 43, adult, Honshu (Nara).
Tani 1978, 4, Honshu (Niigata).
Tanida 1980a, 136, Honshu (Ishikawa).
Tanida 1982a, 357, Honshu (Gumma).
Tanida 1982b, 18, 27, Honshu (Ishikawa).
Tanida 1985, 177-178, larva.
Togashi and Tanida 1985, Honshu (Ishikawa).
Ohtsuka and Tanida 1986, 5, Kyushu (Kumamoto).
Gyotoku and Nozaki 1991, 74, pl. 9, adult, Kyushu (Fukuoka).
Nishiharu 1997a, 60, Honshu (Fukui).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Kagaya et al. 1998, 26, 51-52, 158, Honshu (Tokyo, Yamanashi).
Nozaki and Nakamura 2002, 171, Honshu (Hiroshima).
Tanida 2005, 478-481, male, larva.
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Tanida 2008, 249, pl. 92, adult, Shikoku (Ehime).
Inazu and Nishida 2011, male, larva, Honshu (Hyogo).
Inazu 2012, 7, Honshu (Hyogo).
Tanida 2016, 94 (maculate).
Nojima 2017, 119-120, Honshu (Okayama).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Type locality. Honshu (Nagano?), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. Katsuma (2005a) recorded a species as this species from Ibaraki Prefecture,
but Katsuma (2011c) treated it as a possible undescribed species. His specimens
had clear orange colored wings (Katsuma 2011c).
Parapsyche shikotsuensis (Iwata 1927)
Diplectrona shikotsuensis Iwata 1927a, 223, 228, pl. 7, larva, Hokkaido (Ishikari).
Iwata 1927b, 207, pl. 3, larva, Hokkaido (Ishikari).
Fischer 1963, 149.
Parapsyche sp. PC: Tanida, 1985, 177-178, 215, larva. Mentioned by Kuranishi (1989).
Tanida 2005, 479-480, larva.
Parapsyche shikotsuensis: Kuranishi 1989, 813, combination.
Kuranishi 1991, 149-151, adult, oviposition, Hokakido (Iburi).
Kuhara 1992, 45, Hokkaido (Shiribeshi).
Kuranishi and Kuhara 1994, 1225, Hokkaido (Kushiro).
Kuhara 1997a, 59, Hokkaido (Shiribeshi).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Ito 1999, 163-173, life history, Hokkaido (Ishikari).
Ohkawa 1999, 40-42, Hokkaido (Tokachi).
Ito et al. 2000a, 24, Hokkaido (Ishikari).
Kuhara et al. 2000, 246, Hokkadio (Iburi).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Kuranishi and Satake 2004, 35, Hokkaido (Okhotsk).
Kuhara et al. 2006, 45-46, 49-50, Hokkaido (Ishikari).
Ito et al. 2007, 148, Hokkaido (Kushiro).
Kuhara et al. 2007, app. 70, Hokkaido (Ishikari).
Muramatsu 2008, 5, Hokkaido (Okhotsk).
Ito et al. 2010, 61, Hokkaido (Shiribeshi, Hiyama, Oshima).
Tsuji et al. 2010, 5, Hokkaido (Okhotsk).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Kuhara 2011, 51, 55, 59, 70, Hokkaido (Sorachi, Ishikari, Hidaka, Iburi).
Ito 2013, 57, Hokkaido (Nemuro, Okhotsk, Kushiro).
Ijima and Tsuchiya 2014, 30, Hokkaido (Okhotsk, Kushiro).
Ito and Nagasaka, 2014, 11, Hokkaido (Okhotsk).
Tanida 2016, 94.
Ito 2017b, 42-43, male, female, Hokkaido (Kamikawa).
Tanida 2018, 567, 569-570, larva, male, female.
Nakatani 2019, 49, Hokkaido (Kushiro).
Muramatsu 2022, 20, male, female.
Ito and Kuhara 2023, 55, Hokkaido (Soya, Kamikawa).
Type locality. Hokkaido (Ishikari), Japan.
Distribution. Japan (Hokkaido).
Remarks. Kuranishi (1989) found that the larva described as D. shikotsuensis by Iwata (1927a) is coincident with the 3rd instar larva of P. sp. PC.
Parapsyche sp. PB
Diplectrona felix McLachlan: Iwata 1927a, 227-228, pl. 7, larva. Mentioned by Akagi (1956b).
Iwata 1927b, 207, pl. 3, larva.
Arctopsyche sp. B: Tsuda 1937, 303-303, larva, Honshu (Shiga, Kyoto) (larval record).
Akagi 1956a, 20-21, larva, Honshu (Kyoto: larval record).
Tsuda and Akagi 1962, 131-132, larva.
Parapsyche sp. PB: Tanida, 1985, 177-178, 215, larva.
Tanida 2005, 479-480, larva.
Inazu and Nishida 2011, 190, larva, Honshu (Hyogo: larval record).
Tanida 2018, 567, 569, larva.
Distribution. Japan (Honshu).
Remarks. Akagi (1956b) pointed out that the larva recorded as D. felix by Iwata (1927a) must be coincident with Arctopsyche sp. B. Tanida (1985) redescribed Arctopsyche sp. B as Parapsyche. sp. PB.
'Parapsyche maculata species complex'
Parapsyche sp.: Kuranishi 2013, 188-190, female, larva, Mitochondrial COI gene, Honshu (Kanagawa).
Kawase & Kato 2023, 4, Honshu (Niigata).
Parapsyche maculata species complex: Nozaki et al. 2014, 61, Honshu (Nagano).
Yamamoto and Ito 2014, 13, Shikoku (Ehime).
Nozaki 2016, 75, 347-348, 450, adult, Honshu (Nagano), Shikoku (Kochi).
Yamamoto and Ito 2016, 40, 43, Shikoku (Ehime).
Tanaka 2022, 36, 39, male, Honshu (Aomori, Akita).
Yamamoto 2021, 74, Shikoku (Ehime).
Tanaka 2024, 26, Honshu (Akita).
Parapsyche sp. 1: Tanaka 2016a, 23, 26, male, Honshu (Akita).
Parapsyche sp. 2: Tanaka 2016a, 23, 26, male, Honshu (Akita).
Parapsyche maculata gr.: Tanida 2018, 567, 569, 570, male, larva.
Remarks. Records of unidentified adults related to P. maculata are tentatively listed here for future study.
Subfamily Diplectroninae Ulmer
Two genera, Diplectrona and Homoplectra, has been recorded from Japan.
Genus Diplectrona Westwood
Six named species are known from Japan. Diplectrona albomarginata Ulmer 1907 and Diplectrona tohokuensius Kobayashi 1973 were transferred to the genus Homoplectra (Nozaki 2009, 2021). Diplectrona japonica (Banks, 1906), originally described as a member of the genus Arctopsyche, was also transferred to the genus Homoplectra (Nozaki 2021).
Diplectrona aiensis Kobayashi 1987
Diplectrona aiensis Kobayashi 1987, 19, 32, male, Honshu (Shimane).
Morita 2001c, 175, Honshu (Mie).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Katsuma 2009, 35, Honshu (Ibaraki).
Katsuma 2010b, 85, Honshu (Ibaraki).
Kawase and Hayashi 2011b, 192, Honshu (Shimane).
Kawase and Hayashi 2012, 256, Honshu (Shimane).
Morita 2012, 250, Honshu (Mie).
Morita 2013, 62-63, Honshu (Mie).
Inazu 2013a, 56, Honshu (Hyogo).
Nozaki 2016, 348, 435, male.
Tanaka 2016b, 2, 5, male, Honshu (Akita).
Tanida 2016, 97.
Kobayashi et al. 2017, 17, Honshu (Shiga).
Nojima 2017, 119, Honshu (Okayama).
Tanaka 2018, 21, Honshu (Akita).
Nozaki 2021, 250-251, 256, male, female, Honshu (Akita, Ibaraki, Aichi, Mie, Shiga, Okayama), Kyushu (Saga).
Tanaka 2023, 41, Honshu (Akita).
Kochi et al. 2024, 63-78, larva, pupa, habitat, ecology, Honshu (Nara).
Tanaka 2024, 26, Honshu (Akita).
Type locality. Honshu (Shimane), Japan.
Distribution. Japan (Honshu, Kyushu).
Diplectrona albofasciata (Ulmer 1913)
Diplectrona albofasciata Ulmer 1913, 49-50, male, Taiwan.
Ito and Nozaki 2018, 548-549, male, female, Ishigaki-jima, Yonaguni-jima.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima, Yonaguni-jima), Taiwan.
Diplectrona burha Schmid 1961
Diplectrona burha Schmid 1961, 200-201, male, Pakistan.
Malicky 2002, 1205-1206, 1218, 1233, pl. 14, male, Pakistan, India, Nepal, Thailand, Vietnam, Taiwan.
Ito and Nozaki 2018, 548-549, male, female, Amami-o-shima, Okinawa-jima, Ishigaki-jima, Iriomote-jima.
Nozaki 2021, 249-250, 256, male, Shikoku (Kochi).
Type locality. Pakistan.
Distribution. Japan (Shikoku, Amami-o-shima, Okinawa-jima, Ishigaki-jima, Iriomote-jima),
China (Hubei), Taiwan, Pakistan, India, Nepal, Thailand, Vietnam.
Remarks. Malicky (2002) reported geographic variations of the male genitalia in
this species and provided illustrations of males collected from Nepal,
Thailand, Vietnam and Taiwan. On the other hand, Oláh et al. (2021) recognized each population of the latter three countries as a distinct
species based on the illustrations by Malicky (2002).
Diplectrona kibuneana Tsuda 1940
Diplectrona kibuneana Tsuda 1940, 24-25, 33, male, Honshu (Kyoto).
Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942a, 281, Honshu (Nagano, Kyoto).
Tsuda 1943, 163, 165, 170, Honshu (Kyoto).
Fischer 1963, 146.
Kobayashi 1983, 32, 34, Honshu (Kanagawa).
Morita 1988, 5, Honshu (Mie).
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 39, Honshu (Kanagawa).
Morita 1989, 39, Honshu (Mie).
Kugo and Nozaki 1990, 61, Honshu (Hyogo).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Moriya 1992, 41, Honshu (Kanagawa).
Morita 1993, 3, Honshu (Mie).
Moriya 1994, 71, Hnshu (Kanagawa).
Ito et al. 1997, 23, 27, Hokkaido (Hiyama).
Kagaya et al. 1998, 25, 47, 144, Honshu (Tokyo, Yamanashi).
Nakano and Tanida 1999, 276, Honshu (Nara).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Morita 2001b, 338, Honshu (Mie).
Kawase et al. 2004, 110, Honshu (Ibaraki).
Nozaki 2004, 1230, Honshu (Kanagawa).
Torii and Hattori 2006, 37, Honshu (Shizuoka).
Inazu 2008, 76, 92, 102, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Morita 2009a, 140, Honshu (Mie).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito et al. 2010, 61, Hokkaido (Oshima).
Katsuma 2010b, 85, Honshu (Ibaraki).
Kawase and Hayashi 2010b, 85, Honshu (Shimane).
Kawase and Morita 2010, 42, Honshu (Shiga).
Inazu and Nishida 2011, 191, male, larva, Honshu (Hyogo).
Katsuma 2011b, 49-51, Honshu (Ibaraki).
Kuhara 2011, 51, 70, Hokkaido (Ishikari).
Morita 2011, 216, Honshu (Mie).
Katsuma 2012b, 65, Honshu (Ibaraki).
Moriya and Kubota 2012, 312, Honshu (Kanagawa).
Ito and Nagasaka 2014, 11, Hokkaido (Sorachi).
Kawase and Morita 2014, 5, Honshu (Shiga).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Yamamoto and Ito 2014, 13, Shikoku (Ehime).
Nozaki 2016, 75, 300, 346, 348, 435, 450, male, Shikoku (Kochi).
Tanaka 2016b, 2, 5, male, Honshu (Akita).
Tanida 2016, 98.
Hayashi et al. 80, Kyushu (Miyazaki).
Imasaka et al. 2017, 61, Kysuhu (Fukuoka).
Ito 2017a, 5, Honshu (Fukui).
Katsuma 2017, 39-41, Honshu (Chiba).
Kawase et al. 2017, 32, Honshu (Shiga).
Kuhara and Ito, 2017, 14-15, Yaku-shima.
Nojima 2017, 119, Honshu (Okayama).
Tanida 2018, 571, 577, larva.
Morita 2020d, 81, Honshu (Mie).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Nozaki 2021, 246-248, 256, male, female, Hokkaido (Kushiro, Oshima), Honshu (Iwate, Akita, Fukushima, Gumma, Ibaraki,
Tokyo, Kanagawa, Niigata, Toyama, Yamanashi, Nagano, Gifu, Shizuoka, Mie,
Shiga, Hyogo, Hiroshima), Shikoku (Ehime, Kochi), Kyushu (Fukuoka, Nagasaki,
Kumamoto, Miyazaki, Kagoshima).
Shimura and Imasaka 2021, 4, Kamikoshiki-jima, Shimokoshiki-jima.
Tezuka et al. 2021, 43, Honshu (Kanagawa).
Yamamoto et al. 2021, 73, Shikoku (Ehime).
Tanaka 2023, 41, Honshu (Akita).
Ito and Kuhara 2023, 54, Hokkaido (Kamikawa).
Tanaka 2024, 26, Honshu (Akita).
Diplectrona sp. DB: Akagi 1956b, 24-25, larva, (Ibaraki, Mie, Nara). Associated with the adult by Kagaya et al. (1998).
Tsuda and Akagi 1962, 133-134, larva.
Tanida 1985, 176-177, larva.
Tanida 2005, 487, larva.
Hydropsyche difficultata Kobayashi 1984a, 5-6, male, Honshu (Kanagawa). Synonymized by Nozaki (2021).
Kobayashi 1985a, 12-13, Tsushima.
Moriya 1994, 72, Honshu (Kanagawa).
Diplectrona difficultata: Nozaki 1997a, 34, Honshu (Kanagawa).
Moriya 1997, 62, Honshu (Kanagawa, Yamanashi).
Morita 2008c, 723, Honshu (Mie).
Morita 2009b, 6, Honshu (Mie).
Tanida 2016, 98.
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Koshiki-jima, Yaku-shima), Korea.
Remarks. Shimura (2015) recorded this species from Fukue-jima based on identification
of the larval stage.
Diplectrona takaii Nozaki 2021
Diplectrona sp. DC: Akagi 1956b, 24-25, larva, Honshu (Nara: larval record). Mentioned by Nozaki (2021). The larval
and adult association made by Kagaya et al. (1998).
Tsuda and Akagi 1962, 133-134, larva.
Tanida 1985, 176-177, larva.
Tanida 2005, 487, larva.
Tanida 2018, 577, larva.
Diplectrona difficultata: Kagaya et al. 1993, 75. Misidentification of Diplectrona sp. DC mentioned by Kagaya et al. (1998).
Diplectrona sp. 1: Torii & Hattori 2006, 37, Honshu (Shiga). Mentioned by Nozaki (2021).
Diplectrona sp. (aiensis gr.): Kawase & Morita 2010, 42, Honshu (Shiga). Mentioned by Nozaki (2021).
Diplectrona sp. (aff. aiensis): Nojima 2017, 119, Honshu (Okayama). Mentioned by Nozaki (2021).
Diplectrona sp.: Nozaki 2016, 75, 348, adult, Shikoku (Kochi). Mentioned by Nozaki (2021).
Diplectrona takaii Nozaki 2021, 251-254, 256, male, female, Honshu (Tokyo, Shizuoka, Gifu, Shiga, Okayama), Shikoku (Ehime, Kochi).
Kochi et al. 2024, 67-68, larva.
Type locality. Shikoku (Kochi), Japan.
Distribution. Japan (Honshu, Shikoku).
Diplectrona uken Ito and Nozaki 2018
Diplectrona uken Ito and Nozaki 2018, 549-551, male, Amami-o-shima, Ishigaki-jima.
Type locality. Japan (Amami-o-shima).
Distribution. Japan (Amami-o-shima, Ishigaki-jima).
Diplectrona sp. DA
Diplectrona sp. DA: Akagi 1956b, 23-25, larva, Honshu (Nagano, Shiga, Nara) (larval record).
Tsuda and Akagi 1962, 133-134, larva.
Tanida 1985, 176-177, larva.
Tanida 2005, 487, larva.
?Diplectrona sp. DA: Tanida 2018, 577, larva.
Diplectrona sp.
Diplectrona sp. ?DC: Tanida 1997, 448, Okinawa-jima, Kume-jima, Yaeyama Islands, (larval record).
Diplectrona sp.: Tanida 2003, 376, 381, larva, Okinawa-jima, (larval record).
Remarks. This larva is very similar to that of Diplectrona sp. DC (Tanida, 1997, 2003). Although Tanida (1997) examined both larval
and adult specimens, the specific status has not yet been clarified.
Diplectrona sp.
Diplectrona sp.: Shimura 2020, 67-68, larva, Kikai-jima (larval record).
Remarks. Shimura (2020) recorded a larva similar to Diplectrona sp. DC, but the larva bears distinct markings on it's head.
Genus Homoplectra Ross
Ten named species belonging to this genus are known from Japan.
Homoplectra albomarginata (Ulmer 1907)
Diplectrona albomarginata Ulmer 1907a, 74, female, Japan.
Tanida 2016, 97.
Homoplectra albomarginata: Nozaki 2021, 254-256, female, combination, Honshu (Gifu, Shiga).
Type locality. Japan (precise locality unknown).
Distribution. Japan (Honshu).
Remarks. Nozaki (2021) mistakenly wrote that the type specimen of this species
was collected from Gifu, but Ulmer (1907) did not mention precise locality
other than Japan.
Homoplectra crassa Nozaki 2019
Diplectrona sp. 3: Torii & Hattori 2006, 38, Honshu (Shizuoka). Mentioned by Nozaki (2019).
Homoplectra sp.: Katsuma 2012a, 53, Honshu (Ibaraki). Mentioned by Nozaki (2019).
Tsuruda 2018, 27, 31, Honshu (Tokyo).
Homoplectra crassa Nozaki 2019, 330-333, 341-342, male, female, larva, Honshu (Ibaraki, Tokyo, Kanagawa, Yamanashi, Shizuoka, Aichi).
Type locality. Honshu (Tokyo), Japan.
Distribution. Japan (central Honshu).
Homoplectra crassicornea Nozaki and Ito 2018
Homoplectra sp.: Kuhara and Ito 2017, 15, Yaku-shima. Mentioned by Ito and Nozaki (2018).
Homoplectra crassicornea Nozaki and Ito 2018: Ito and Nozaki 2018, 551-552, male, female, Yaku-shima.
Type locality. Yaku-shima, Japan.
Distribution. Japan (Yaku-shima).
Homoplectra gracilis Nozaki 2019
Diplectroninae gen. sp.: Kugo and Nozaki 1991, 327, Honshu (Hyogo). Mentioned by Nozaki (2019).
Nozaki and Nakamura 2002, 171, Honshu (Hiroshima).
Diplectroninae gen. sp. 1: Inazu 2008, 76, Honshu (Hyogo). Mentioned by Nozaki (2019).
Inazu 2012, 8, Honshu (Okayama).
Homoplectra sp. A: Nojima 2017, 119, Honshu (Okayama). Mentioned by Nozaki (2019).
Homoplectra gracilis Nozaki 2019, 339-340, 342-343, male, female, Honshu (Hyogo, Okayama, Hiroshima).
Nokima 2022, 38, Honshu (Okayama).
Kochi et al. 2023, 286-294, male, female, larva, pupa, biology, Honshu (Nara).
Type locality. Honshu (Okayama), Japan.
Distribution. Japan (western Honshu).
Remarks. Kawase & Kato (2023) recorded males and females possiblly belonging
to this species as Homoplectra sp. cf. gracillis from Niigata Prefecture, eastern Japan.
Homoplectra hattorii Nozaki 2019
Homoplectra sp.: Ito et al. 2010, 61, Hokkaido (Oshima). Mentioned by Nozaki (2019).
Homoplectra hattorii Nozaki 2019, 551-552, male, Hokkaido (Sorachi, Hidaka, Oshima)..
Type locality. Hokkaido (Sorachi), Japan.
Distribution. Japan (Hokkaido).
Homoplectra ise Nozaki 2019
Homoplectra sp.: Morita 2008b, Honshu (Mie). Mentioned by Nozaki (2019).
Homoplectra sp. 1: Yamamoto and Ito 2014, 13, Shikoku (Ehime). Mentioned by Nozaki (2019).
Homoplectra ise Nozaki 2019, 333-335, 342, male, female, Honshu (Mie), Shikoku (Ehime).
Type locality. Honshu (Mie), Japan.
Distribution. Japan (Honshu (central), Shikoku).
Homoplectra japonica (Banks, 1906)
Arctopsyche japonica Banks 1906, 111, 113, color pattern of forewing, Kyushu (Fukuoka).
Diplectrona japonica: Ulmer 1907a, 72-74, pl. 4, male, Honshu (Gifu);
Ulmer 1907b, 177, pl. 38, adult.
Matsumura 1931, 1132, adult, Honshu (Nagano).
Kuwayama 1932, 1511, adult.
Esaki et al. 1938, 135, pl. 61, adult.
Tsuda 1940, 24, 33, Honshu (Nagano, Kyoto).
Kobayashi 1942, 220, Honshu (Tokyo).
Tsuda 1942a, 281, Honshu (Nagano, Gifu, Kyoto, Osaka).
Tsuda 1943, 164-165, Honshu (Kyoto).
Kuwayama 1949, 309, adult.
Tsuda 1950, 418, adult.
Ishihara et al. 1953, 38, Shikoku (Ehime).
Takeuchi 1955, 74, pl. 28, adult, Honshu (Kyoto).
Fischer 1963, 146.
Kuwada 1965, 164, pl. 82, adult, Shikoku (Ehime).
Kobayashi 1968, 3, Honshu (Niigata).
Tani 1977, 197, pl. 43, adult, Honshu (Nara).
Tanida 1980a, 136, Honshu (Ishikawa).
Tanida and Takemon 1981, 206, Honshu (Kyoto).
Morita 1990, 31, Honshu (Mie).
Gyotoku and Nozaki 1991, 74-75, pl. 9, adult, Fukuoka (Kyushu).
Hatta and Nozaki 1991, 200, 201, adult, Honshu (Aichi).
Morita 1993, 3, Honshu (Mie).
Nishiharu 1997a, 60, Honshu (Fukui).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Uenishi and Kurumi 2000, 275, 279, Honshu (Shiga).
Morita 2000, 60, Honshu (Mie).
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Sasaji et al. 2003, 275, Honshu (Fukui). (Diplectorona)
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Morita and Kawase 2004, 17, Honshu (Ishikawa).
Tanida 2005, 482, 487, adult.
Mey and Nozaki 2006, 24, Honshu (Nagano).
Morita 2008c, 723, Honshu (Mie).
Tanida 2008, 249-250, pl. 92, adult, Shikoku (Ehime).
Kawase and Hayashi 2010b, 85, Honshu (Shimane).
Inazu and Nishida 2011, 191, male, female, Honshu (Hyogo).
Inazu 2012, 8, Honshu (Hyogo).
Inazu 2013b, 72, Honshu (Hyogo).
Tanida 2016, 98.
Tanida 2018, 571, 577, adult.
Homoplectra japonica: Nozaki 2021, 254-256, male, conbination, Kyushu (Fukuoka).
Type locality. Kyushu (Fukuoka), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. This species has distinct yellow markings on the forewings (Banks 1906) and many authors recorded as this species based on this characteristic markings. However, Kawase and Morita (2014) and Nozaki et al. (2014) pointed out that multiple species have this character. This taxonomic
problem need to be solved.
Homoplectra occidens Nozaki 2019
Homoplectra sp.: Kawase et al. 2017, 32, Honshu (Shiga). Mentioned by Nozaki (2019).
Homoplectra sp. B: Nojima 2017 119, Honshu (Okayama). Mentioned by Nozaki (2019).
Homoplectra occidens Nozaki 2019, 338-339, 342, male, female, Honshu (Shiga, Hyogo, Okayama).
Nojima 2022, 38, Honshu (Okayama).
Type locality. Honshu (Hyogo), Japan.
Distribution. Japan (central to western).
Homoplectra tohokuensis (Kobayashi 1973)
Diplectrona tohokuensis Kobayashi, 1973, 31-32, pl. 7, male, Honshu (Yamagata).
Tanida 2016, 98.
Homoplectra tohokuensis: Nozaki 2019, 336-338, 341, 342, male, female, larva, Honshu (Aomori, Iwate, Miyagi, Akita, Yamagata).
Tanaka 2022, 35, 38, male, Honshu (Akita).
Aoya and Tanaka 2023, 27, Honshu (Akita).
Type locality. Honshu (Yamagata), Japan.
Distribution. Japan (North east Honshu).
Homoplectra trifurcata Nozaki and Ito 2018
Homoplectra trifurcata Nozaki and Ito 2018: in Ito and Nozaki 2018, 552-553, male, female, Amami-o-shima.
Type locality. Japan (Amami-o-shima).
Distribution. Japan (Amami-o-shima).
Homoplectra japonica species complex
Diplectrona sp. (af. japonica): Morita 1995, 96, Honshu (Mie).
Diplectrona sp. 2: Torii and Hattori 2006, 37-38, Honshu (Shizuoka).
Diplectroninae Gen. sp.: Morita 2008b, 91-92, Honshu (Mie).
Morita and Kawase 2010, 26, Honshu (Ishikawa).
Kawase and Morita 2014, 5, Honshu (Shiga).
Gen. japonica species complex: Nozaki et al. 2014, 61, Honshu (Nagano).
Yamamoto and Ito 2014, 13, Shikoku (Ehime).
Nozaki 2016, 75, 348, adult, Shikoku (Kochi).
Nojima 2017, 119, Honshu (Okayama).
Yamamoto et al. 2021, 73, Shikoku (Ehime).
Diplectroninae Gen. spp.: Kawase et al. 2017, 32, Honshu (Shiga).
Diplectrona japonica species complex: Watanabe 2021, 41-43, adult, mating behavior, Honshu (Hyogo).
Remarks. Records of unidentified species with yellow markings on the forewings like H. japonica are tentatively listed under this name.
Subfamily Hydropsychinae Curtis
Four genera have been recorded from Japan, but two species recorded as
members of the genus Hydromanicus have not yet been identified to species level.
Genus Cheumatopsyche Wallengren
The genus Hydropsychodes Ulmer was synonymized with this genus by Kimmins (1963). Eleven species
belonging to this genus have been recorded from Japan. Tanida (2003, 2005)
provided illustrations of an unidentified larva from Ryukyu Islands.
Cheumatopsyche aira Ito and Nozaki 2018
Cheumatopsyche aira Ito and Nozaki 2018, 556-558, male, female, Iriomote-jima.
Type locality. Iriomote-jima, Japan.
Distribution. Japan (Iriomote-jima).
Cheumatopsyche addita (Navás 1916)
Hydropsychodes addita Navás 1916, 88-90, male, Japan.
Fischer 1963, 100.
Tanida 2016, 95.
Type locality. Japan (precise locality unknown).
Distribution. Japan.
Remarks. Oláh et al. (2008) treated this species as "incertae sedis".
Cheumatopsyche brevilineata (Iwata 1927)
Hydropsyche brevilineata Iwata 1927a, 223, 226-227, larva, Honshu (Kyoto, Hyogo).
Iwata 1927b, 207, larva, Honshu (Kyoto).
Hydropsychodes brevilineata: Tsuda 1940, 28-30, 33, male, wings, Hokkaido (Omu), Honshu (Saitama, Nagano, Hyogo, Kyoto), Shikoku (Tokushima),
Kyushu (Kumamoto).
Tsuda 1942a, 280, Honshu (Shiga, Kyoto).
Tsuda 1942b, 263-267, Honshu (Kyoto).
Tsuda 1943, 166-168, 170, Honshu (Kyoto).
Tsuda 1950, 417, male.
Tsuda and Tatsumi 1951, 28-29, 31, Honshu (Niigata).
Akagi 1955, 20, larva.
Chihara 1955, 24, pupa.
Tsuda and Kawai 1956, 73-75, Honshu (Kyoto).
Tsuda 1959, 141, larva.
Tsuda and Akagi 1962, 132-133, larva.
Fischer 1963, 101-102.
Kobayashi 1964, 87, Hokkaido (Tokachi).
Kuwada 1965, 163, male, pl. 82, Shikoku (Ehime).
Kobayashi 1973, 31, Honshu (Yamagata).
Tani 1976, 392, Tsushima.
Tani 1977, 196, male.
Tani 1978, 6, Honshu (Niigata).
Tanida 1980b, 98-99, Honshu (Kyoto).
Tanida and Takemon 1981, 206, Honshu (Kyoto).
Kobayashi 1983, 32-33, Honshu (Kanagawa).
Kobayashi 1984c, 3-4, Sado.
Kobayashi 1984d, 28-31, Honshu (Kanagawa).
Adachi et al. 1985, 82-88, early instar larvae.
Kobayashi 1985b, 68-70, 72-73, Honshu (Kanagawa).
Kobayashi 1987, 18-19, Honshu (Shimane).
Cheumatopsyche brevilineata: Tanida 1985, 215, convination.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Adachi et al. 1986, 86-89, 91-92, larval mandibles.
Adachi et al. 1987, 139-142, adult, egg, laboratory rearing,
Nozaki 1988a, 45, Honshu (Kanagawa).
Nozaki 1988b 39, Honshu (Kanagawa).
Itou 1989, 55, Hokkaido (Kushiro).
Yagi et al. 1989, peculiar organ of female, 177-181, Honshu (Kyoto).
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Nozaki and Gyotoku 1990, 13, 16, Kyushu (Fukuoka).
Tanida 1990, 55, Honshu (Hyogo).
Gyotoku and Nozaki 1991, 76, Kyushu (Fukuoka).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Hatta and Nozaki 1991, 201, 208, adult, Honshu (Gifu).
Tanida 1991a, 34, Honshu (Nara).
Tanida et al. 1991, 58, adult, pupa, larva.
Yagi and Sasakawa 1992, 4-9, life history, swarming behavior, Honshu (Kyoto).
Moriya 1992, 42, Honshu (Kanagawa).
Moriya 1994, 71, Honshu (Kanagawa).
Itou and Muramatsu 1995, 39, Hokkaido (Okhotsk).
Kuranishi and Ohtaka 1996, 14, Honshu (Aomori).
Tani and Nakamura 1997, 59-60, Honshu (Hiroshima).
Hayashi 1998, 177-181, enzyme polymorphism, Honshu (Tokyo, Kyoto)..
Kadowaki 1998, 4, Honshu (Shimane).
Sekimoto et al. 1998, 360, Honshu (Hiroshima).
Yodoe et al. 1998, 68, Honshu (Shimane).
Kuhara et al. 2000, 246, Hokkaido (Iburi).
Hayashi et al. 2001, 51-57, reach scale distribution, Honshu (Tokyo).
Morita 2001c, 175, Honshu (Mie).
Nishimoto and Nishimoto 2001, 72, 76, adult, Honshu (Aichi).
Nishimoto, 2002, 206-209, cuticular hydrocarbons, Honshu (Aichi).
Nozaki and Nakamura 2002, 168-169, Honshu (Hiroshima).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshyu (Hiroshima).
Tanida 2003, 382, wings.
Ito et al. 2004, 11-12, Hokkaido (Kushiro).
Kawase et al. 2004, 109-110, Honshu (Ibaraki).
Nozaki 2004, 1230, Honshu (Kanagawa).
Uenishi and Yoshitaka 2004, 16, Honshu (Okayama).
Ishiwata et al. 2005, 110, 242-244, Honshu (Kanagawa).
Katsuma 2005a, 4, Honshu (Ibaraki).
Nozaki and Nakamura 2005, 221, Honshu (Shimane).
Tanida 2005, 483, 488, wings.
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Kimura et al. 2006, 7, Honshu (Nagano).
Mochizuki et al. 2006, 132-134, larval growth and development, Honshu (Gifu).
Morita 2006, 200, Honshu (Mie).
Nakamura and Fukui 2006, 243, Honshu (Hiroshima).
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Torii and Hattori 2006, 37, Honshu (Shizuoka).
Ito and Kosugi 2007, 53, Hokkaido (Kushiro).
Katsuma 2007, 18, Honshu (Ibaraki).
Nakamura and Hajiri 2007, 198, Hnshu (Shimane).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Nakamura 2007, 94, Honshu (Hiroshima).
Nozaki and Tanida 2007, 248, Honshu (Shizuoka).
Inazu 2008, 75, 92, 102, Honshu (Hyogo).
Katsuma 2008, 58, Honshu (Ibaraki).
Kimura and Hirabayashi 2008, 134-138, Honshu (Nagano).
Kimura et al. 2008, 264, Honshu (Nagano).
Morita 2008a, 17, Honshu (Mie).
Morita 2008c, 722-723, Honshu (Mie).
Oláh et al. 2008, 50-51, male, Honshu (Nagano).
Tanida 2008, 250, pl. 92, adult, Shikoku (Ehime).
Kameyama et al. 2009, 111, Honshu (Shimane).
Nozaki 2009, 5-6, larval head.
Yokoyama et al. 2009, laboratory rearing.
Ito 2010, 130, Hokkaido (Kushiro).
Ito et al. 2010, 61, Hokkaido (Oshima).
Katsuma 2010a, 84, Honshu (Ibaraki).
Kawase and Hayashi 2010a, 80, Honshu (Shimane).
Kawase and Morita 2010, 41, Honshu (Mie, Shiga).
Mano, 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Nozaki 2010, 216, Honshu (Kanagawa).
Inazu and Nishida 2011, 192, male, Honshu (Hyogo).
Katsuma 2011d, 69, Honshu (Ibaraki).
Kawase and Hayashi 2011a, 172, Honshu (Shimane).
Kawase and Hayashi 2011b, 192, Honshu (Shimane).
Morita 2011, 216, Honshu (Mie).
Nozaki 2011, 4, Honshu (Aichi).
Kawase and Hayashi 2012, 256, Honshu (Shimane).
Morita 2012, 250, Honshu (Mie).
Torii et al. 2012, 51, larval head.
Inazu 2013a, 56, Honshu (Hyogo).
Inazu 2013b, 72, Honshu (Hyogo).
Ito 2013, 55, Hokkaido (Kushiro, Tokachi).
Nozaki 2013, 18, Honshu (Aichi).
Takai et al. 2013, 84, adult.
Ijima and Tsuchiya 2014, 29, Hokkaido (Okhotsk, Kushiro).
Ito and Nagasaka, 2014, 10, Hokkaido (Okhotsk).
Katsuma 2014, 45, Honshu (Ibaraki).
Mochizuki et al. 2014, 138-145, generation, Honshu (Gifu).
Nishimoto and Nishimoto 2014, Honshu (Aichi).
Yamamoto and Ito 2015, 33, Shikoku (Ehime) (bravilineata).
Ito and Kuhara 2016, 16, Hokkaido (Kushiro).
Nozaki 2016, 76, 346, 435, 349, male, Shikoklu (Kochi).
Nozaki et al. 2016, 328, Honshu (Niigata).
Tanaka 2016a, 22, 24. male, Honshu (Akita).
Tanida 2016, 95.
Yamamoto and Ito 2016, 40, 43, Shikoku (Ehime).
Ito 2017a, 5, Honshu (Fukui).
Kawase et al. 2017, 30-31, Honshu (Shiga).
Kobayashi et al. 2017, 17, 19, 20, Honshu (Shiga, Kyoto).
Nojima 2017, 118, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Ohira and Tsutsumi 2018, 24, Honshu (Fukushima).
Tanaka 2018a, 21, Honshu (Akita).
Tanaka 2018b, 8, Honshu (Akita).
Tanida 2018, 573, 578, larva, wings.
Tsuruda 2018, 26-27, Honshu (Tokyo).
Kimura et al. 2019, 60-61, Shikoku (Kagawa).
Nozaki et al. 2019, 169, Honshu (Nagano).
Tanaka 2019b, 12, Honshu (Akita).
Yamamoto et al. 2019, 71, Shikoku (Ehime).
Kimura and Kuranishi 2020, 157, Honshu (Kyoto).
Tezuka et al. 2021, 43, Honshu (Kanagawa).
Yamamoto and Nozaki 2021, 128, Shikoku (Ehime).
Tanaka 2022, 35, Honshu (Akita).
Ito and Kuhara 2023, 54, Hokkaido (Soya).
Kuhara et al. 2023, 15-16, Kyushu (Kumamoto, Miyazaki, Kagoshima).
Tanaka 2023, 41, Honshu (Akita).
Tanaka 2024, 26, Honshu (Akita).
Typelocality. Honshu (Kyoto, Hyogo), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki), Russia (Far East).
Remarks. Tanida et al. (1995) pointed out that this species has been confused with C. infascia and an undescribed species (now C. tanidai). Nishiharu (1997a) recorded this species from Fukui Prefecture, but Nishiharu
(1997b) revised the record as an unidentified species. Oláh et al. (2008) synonymized a continental species, Cheumatopsyche daurensis Ivanov 1996 in Arefina et al. (1996). Shimura (2015) recorded this species from Fukue-jima based on
identification of the larval stage.
Cheumatopsyche clavalis Martynov 1930
Cheumatopsyche clavalis Martynov 1930, 82, male, Taiwan.
Ito and Nozaki 2018, 554-556, male, female, Ishigaki-jima, Iriomote-jima.
Cheumatopsyche uchidai Kobayashi 1987, 40, 46, male, Taiwan. Synonymized by Ito and Nozaki (2018).
Oláh et al. 2008, 66-67, holotype male, Taiwan.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima, Iriomote-jima), Taiwan.
Cheumatopsyche galloisi (Matsumura 1931)
Hydromanicus galloisi Matsumura 1931, 1132, adult, Shikoku (Tokushima).
Tateishi 1934, 76, Kyushu (Fukuoka).
Tsuda 1940, 28, 33, Honshu (Kyoto, Osaka).
Gyotoku 1957, 10, Kyushu (Fukuoka).
Kobayashi 1959, 349, Kyushu (Fukuoka).
Fischer 1963, 95.
Tani 1977, 196, male.
Tani 1978, 6, Honshu (Niigata).
Tanida 1980b, 98-99, Honshu (Ishikaawa).
Kobayashi 1983, 32-34, Honshu (Kanagawa).
Kobayashi 1984d, 28-29, Honshu (Kanagawa) (gallosis, galloisis).
Kobayashi 1985b, 68, 70-72, Honshu (Kanagawa).
Kobayashi 1987, 18, Honshu (Shimane).
Yagi et al. 1989, peculiar organ of female, 178-179, 181, Honshu (Kyoto).
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Gyotoku and Nozaki 1991, 76, pl. 9, adult, Kyushu (Fukuoka).
Hori et al. 1993, 388, 390, adult, Honshu (Aichi).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Moriya 1994, 71, Honshu (Kanagawa).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Kameyama et al. 2009, 112, Honshu (Shimane).
Cheumatopsyche galloisi: Tanida et al. 1995, 2, 4, Honshu (Nara).
Morita 1997, 41, Honshu (Mie).
Moriya 1997, 62, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 34, Honshu (Kanagawa).
Hayashi and Yun, 1999, 380-383, esterase, adult and larval association,
Honshu (Yamaguchi).
Morita 1999, 82, Honshu (Mie).
Nakano and Tanida 1999, 276, Honshu (Nara).
Niashimoto 2000, 26, Honshu (Aichi).
Morita 2001, 338, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nishimoto, 207-209, cuticular hydrocarbons, Honshu (Aichi).
Nozaki and Nakamura 2002, 169, Honshu (Hiroshima).
Nozaki 2004, 1230, Honshu (Kanagawa).
Katsuma 2005a, 4-5, Honshu (Ibaraki).
Nozaki and Nakamura 2005, 221, Honshu (Shimane).
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Mey and Nozaki 2006, 24, Honshu (Gifu).
Morita 2006, 200, Honshu (Mie).
Torii and Hattori 2006, Honshu (Shizuoka).
Nakamura and Hajiri 2007, 198, Honshu (Shimane).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 94, Honshu (Hiroshima).
Inazu 2008, 75, 75, 92, 102, Honshu (Hyogo).
Morita 2008a, 17, Honshu (Mie).
Oláh et al 2008, 15, 25-26, forewing, male, Honshu (Wakayama).
Tanida 2008, 250, pl. 92, adult, Honshu (Kyoto).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito et al. 2010, 61, Hokkaido (Oshima).
Kobayashi 2010, 140, Honshu (Tochigi).
Nakamura and Kita 2010, 88, Honshu (Hiroshima).
Inazu and Nishida 2011, 192, male, Honshu (Hyogo).
Morita 2011, 216, Honshu (Mie).
Torii et al. 2012, 51, larval head.
Moriya and Kubota 2012, 132, Honshu (Kanagawa).
Inazu 2013b, 72, Honshu (Hyogo).
Ito and Nagasaka, 2014, 10, Hokkaido (Sorachi).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Hayashi & Kadowaki 2016, 31, 68-69, adult, Oki.
Nozaki 2016, 76, 349, 435, 450, adult, Shikoku (Kochi).
Tanaka 2016a, 22, 25, adult, Honshu (Akita).
Tanida 2016, 95.
Ito 2017b, 42, Hokkaido (Kamikawa).
Kawase et al. 2017, 31, Honshu (Shiga).
Nojima 2017, 118, Honshu (Okayama).
Morita 2019b, 66, Honshu (Mie).
Tanaka 2019a, 27, Honshu (Akita).
Tanaka 2019b, 12, Honshu (Akita).
Morita 2020d, 81, Honshu (Mie).
Tanaka 2023, 41, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).
Tanaka 2024, 26, Honshu (Akita).
Hydropsychodes sp. HA: Chino 1975, 8-5, larva: Mentioned by Hayashi and Yun (1999).
Hydropsychidae Gen. sp.: Tanaka 2010a, 24, 27, adult, Honshu (Akita). Mentioned by Tanaka (2011).
Type locality. Shikoku (Tokushima), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki).
Cheumatopsyche guerneana (Navás 1916)
Hydropsychodes guerneana Navás 1916, 88-89, male, Honshu (Kyoto).
Fischer 1963, 103.
Tanida 2016, 95.
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu).
Remarks. Oláh et al. (2008) treated this species as "incertae sedis".
Cheumatopsyche infascia Martynov, 1934
Cheumatopsyche infascia Martynov 1934, 283-285, male, Russia (Southern Far East).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Morita 1995, 96, male, Honshu (Mie).
Tanida et al. 1995, 2, Honshu (Nara).
Nozaki 1997a, 34, Honshu (Kanagawa).
Nozaki 1997b, 28, Honshu (Tokyo).
Nishimoto and Nishimoto 1998, 150, Honshu (Aichi).
Ito et al. 1999, 18, 23, 25-27, Hokkaido (Kushiro).
Hayashi 1998, 177-181, enzyme polymorphism, Honshu (Tokyo, Kanagawa, Kyoto)..
Hayashi and Yun, 1999, 381-383, esterase, Honshu (Yamaguchi).
Nakano and Tanida 1999, 276, Honshu (Nara).
Ohkawa 1999, 40-42, 44, Hokkaido (Tokachi).
Ito et al. 2000a, 23, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-30, Hokkaido (Soya).
Kuhara et al. 246, Hokkaido (Iburi).
Nishimoto 2000, 26, Honshu (Aichi).
Hayashi et al. 2001, 51-57, reach scale distribution, Honshu (Tokyo).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Hayashi and Yun, 2002, 2-5, esteraze, Hokkaido (Hidaka), Honshu (Aomori,
Fukushima, Tokyo, Kanagawa, Shizuoka, Kyoto, Yamaguchi), Shikoku (Tokushima),
Kyushu (Oita).
Nishimoto 2002, 206-209, cuticular hydrocarbons, Honshu (Aichi).
Nozaki and Nakamura 2002, 169, Honshu (Hiroshima).
Nagayasu et al. 2003, 106, Hokkaido (Ishikari).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Ito et al. 2004, 11-12, Hokkaido (Kushiro).
Kawase et al. 2004, 110, Honshu (Ibaraki).
Morita and Kawase 2004, 16, Honshu (Ishikawa).
Nozaki 2004, 1230, Honshu (Kanagawa).
Uenishi and Yoshitaka 2004, 16, Honshu (Okayama).
Yoshitaka 2004, 28, Honshu (Okayama).
Ishiwata et al. 2005, 110, 242-244, Honshu (Kanagawa).
Katsuma 2005a, 5, Honshu (Ibaraki).
Katsuma 2005b, 13, Honshu (Gumma).
Nagayasu et al. 2005, 53, Hokkaido (Ishikari).
Nozaki 2005, 108, adult, larva.
Nozaki and Nakamura 2005, 222, Honshu (Shimane).
Tanida 2005, 483, 488, male, larva.
Kameyama and Nakamura 2006, 189, Honshu (Tottori).
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Kimura et al. 2006, 7, Honshu (Nagano).
Mey and Nozaki 2006, 24, Honshu (gifu).
Nakamura and Fukui 2006, 243, Honshu (Hiroshima).
Nakamura et al. 2006, 101-102, Honshu (Hiroshima).
Torii and Hattori 2006, 37, Honshu (Shizuoka).
Ito and Kosugi 2007, 53, Hokkaido (Kushiro).
Ito et al. 2007, 148, Hokkaido (Kushiro).
Katsuma 2007, 18, Honshu (Ibaraki).
Kuhara et al. 2007, app. 70, Hokkaido (Ishikari).
Nakamura and Hajiri 2007, Honshu (Shimane).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nakano et al. 2007, 39-40, 42, Hokkaido (Nemuro).
Nozaki and Nakamura 2007, 94-95, Honshu (Hiroshima).
Nozaki and Tanida 2007, 248, Honshu (Shizuoka).
Inazu 2008, 75, 92, 102, Honshu (Hyogo).
Kimura et al. 2008, 264, Honshu (Nagano).
Morita 2008a, 17, Honshu (Mie).
Muramatsu 2008, 5, Hokkaido (Okhotsk).
Nakatani et al. 2008, 53, Hokkaido (Okhotsk).
Oláh et al 2008, 27-28, male, Russia.
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Kameyama et al. 2009, 112, Honshu (Shimane).
Nozaki 2009, 5-6, larva.
Tanaka 2009a, 45, adult, Honshu (Akita).
Tanaka 2009c, 18, Honshu (Akita).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito et al. 2010, 61, Hokkaido (Shiribeshi, Hiyama, Oshima).
Katsuma 2010a, 84, Honshu (Ibaraki).
Katsuma 2010b, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 41-42, Honshu (Mie, Shiga).
Kawase et al. 2010, 108, Honshu (Shimane).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Nozaki 2010, 216, Honshu (Kanagawa).
Tanaka 2010a, 24, 26, 28, male, Honshu (Akita).
Tanaka 2010b, 17, Honshu (Akita).
Inazu and Nishida 2011, 192, male, Honshu (Hyogo).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Katsuma 2011b, 49, Honshu (Ibaraki).
Katsuma 2011c, 67, Honshu (Ibaraki).
Kawase and Hayashi 2011a, 173, Honshu (Shimane).
Kawase and Hayashi 2011b, 192, Honshu (Shimane).
Kuhara 2011, 51, 70, Hokkaido (Sorachi, Ishikari, Iburi).
Moriya 2011, 216, Honshu (Mie).
Tanaka and Okura 2011, 24, Honshu (Akita).
Torii et al. 2012, 51, larval head.
Katsuma 2012b, 65, Honshu (Ibaraki).
Kawase and Hayashi 2012, 256, Honshu (Shimane), Oki (Dogo).
Moriya and Kubota 2012, 132, Honshu (Kanagawa).
Inazu 2013a, 56, Honshu (Hyogo).
Inazu 2013b, 72, Honshu (Hyogo).
Ito 2013, 55-56, Hokkaido (Nemuro, Kushiro, Tokachi).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Ijima and Tsuchiya 2014, 29, Hokkaido (Okhotsk, Kushiro, Tokachi, Ishikari).
Ito and Nagasaka, 2014, 10-11, Hokkaido (Okhotsk).
Kawase and Morita 2014, 5, Honshu (Shiga).
Kuranishi and Kikuchi 2014, 363, Honshu (Yamagata).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Shimura 2015, 51, Fukue-jima.
Hirabayashi et al. 2016, 208, 210, Honshu (Nagano).
Ito and Kuhara 2016, 16, Hokkaido (Okhotsk, Kushiro, Tokachi, Ishikari).
Katsuma 2016, 47, Honshu (Ibaraki).
Kimura et al. 2016, 246, Honshu (Yamanashi).
Nakatani 2016, 240, Hokkaido (Nemuro).
Nozaki 2016, 76, 349, 435, 450, male, Shikoku (Kochi).
Tanaka 2016a, 22, 25, male, Honshu (Akita).
Tanida 2016, 95.
Yamamoto and Ito 2016, 40, 43, Shikoku (Ehime).
Ito 2017b, 42, Hokkaido (Kamikawa).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kawase et al. 2017, 31, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Kuhara and Ito 2017, 13-14, Yaku-shima.
Matsuoka 2017, 8, 11, adult, Hokkaido (Tokachi). (nfascia)
Nojima 2017, 118, Honshu (Okayama).
Saito et al. 2017 183, Honshu (Chiba).
Ohira and Tsutsumi 2018, 24, Honshu (Fukushima).
Tanaka 2018a, 21, Honshu (Akita).
Tanaka 2018b, 8, Honshu (Akita).
Tanida 2018, 573, 578, male, larva.
Tsuruda 2018, 27, 31, Honshu (Tokyo).
Nakatani and Ito 2019, 163, Hokkaido (Kushiro).
Nozaki et al. 2019, 169, Honshu (Nagano).
Tanaka 2019a, 27, Honshu (Akita).
Tanaka 2019b, 12, Honshu (Akita).
Katsuma 2020, 15, Kyushu (Oita).
Yamamoto et al. 2019, 72, Shikoku (Ehime).
Yashiro et al. 2020, 185, Kyushu (Kamikoshiki-jima, Shimokoshiki-jima).
Shimura and Imasaka 2021, 4, Kamikoshiki-jima, Shimokoshiki-jima.
Tezuka et al. 2021, 43, Honshu (Kanagawa).
Yamamoto and Nozaki 2021, 128, Shikoku (Ehime).
Muramatsu 2022, 21, male, female.
Tanaka 2022, 35, Honshu (Akita).
Kawase & Kato 2023, 3, Honshu (Niigata).
Tanaka 2023, 41, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).
Ito and Kuhara 2023, 54, Hokkaido (Soya, Kamikawa, Rumoi).
Tanaka 2024, 26, Honshu (Akita).
Cheumatopsyche brevilineata: Tanida 1985, 180-181, larva. Misidentification mentioned by Tanida et al. (1999).
Cheumatopsyche sp.: Kuhara 1997a, 59, Hokkaido (Shiribeshi). Mentioned by Kuhara (2001).
Tanaka 2010a, 27-28, male, Honshu (Akita). Mentioned by Tanaka (2011).
Type Locality. Russia.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Koshiki-jima), Korea, Russia
(Far East).
Remarks. Shimura (2015) recorded this species from Fukue-jima based on identification
of the larval stage.
Cheumatopsyche japonica (Navás 1916)
Hydropsychodes japonica Navás 1916, 87, 89, male, Honshu (Kyoto).
Fischer 1963, 103.
Tanida 2016, 95.
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu).
Remarks. Oláh et al. (2008) treated this species as "incertae sedis".
Cheumatopsyche okinawana Oláh & Johanson 2008
Cheumatopsyche okinawana Oláh & Johanson 2008, 181-182, male, Okinawa-jima.
Tanida 2016, 95.
Ito and Nozaki 2018, 553-555, male, female, Amami-o-shima, Kume-jima (okinawa).
Type locality. Okinawa-jima, Japan.
Distribution. Japan (Amami-o-shima, Okinawa-jima, Kume-jima).
Cheumatopsyche tanidai Oláh & Johanson 2008
Cheumatopsyche tanidai Oláh & Johanson 2008, 184-185, male, Honshu (Kyoto).
Kawase and Morita 2010, 42, Honshu (MIe, Shiga).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Nozaki 2010, 216, Honshu (Kanagawa).
Inazu and Nishida 2011, 192, male, Honshu (Hyogo).
Kawase and Hayashi 2011a, 173, Honshu (Shimane).
Morita 2011, 216, Honshu (Mie).
Morita 2012, 250, Honshu (Mie).
Inazu 2012, 7, Honshu (Hyogo).
Kawase and Hayashi 2012, 256, Honshu (Shimane).
Inazu 2013a, 56, Honshu (Hyogo).
Kawase and Morita 2014, 5, Honshu (Shiga).
Nozaki 2016, 349, 435, male.
Tanaka 2016, 22, 25, male, Honshu (Akita).
Tanida 2016, 95.
Ito 2017a, 5, Honshu (Fukui).
Kawase et al. 2017, 31, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Nojima 2017, 118-119, Honshu (Okayama).
Tanida 2018, 573.
Morita 2019a, 22, Honshu (Mie).
Morimune and Nakamura 2020, 23, Honshu (Okayama).
Morita 2020b, 43, Honshu (Mie).
Morita 2020d, 81, Honshu (Mie).
Kuhara et al. 2023, 16, Kyushu (Saga).
Tanaka 2024, 26, Honshu (Akita).
Cheumatopsyche sp.: Tanida et al. (1995).
Hayashi 1998, 177-181, enzyme polymorphism, Honshu (Kyoto)..
Hayashi 1999, 381-383, esterase, Honshu (Yamaguchi).
Hayashi et al. 2001, 52-53, Honshu (Tokyo).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Uenishi and Yoshitaka 2004, 16, Honshu (Okayama).
Nozaki and Nakamura 2005, 222, Honshu (Shimane).
Nakamura and Fukui 2006, 243, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Morita 2008c, 723, Honshu (Mie).
Kameyama et al. 2009, 112, Honshu (Shimane).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu, Kyushu), Korea.
Remarks.Tanida et al. (1995) pointed out that this species has been confused with C. infascia and C. brevilineata, and tentatively recorded as an unidentified species with a Japanese name "Sato-kogata-shima-tobikera". Records of unidetified species with this Japanese name are listed under this species name. Tanida (2016) listed Shikoku and Kyushu as the distribution area of this species without precise data, but Kuhara et al. (2023) recorded this species from Saga, Kyushu, for the first time..
Cheumatopsyche tokunagai (Tsuda 1940)
Hydropsychodes tokunagai Tsuda 1940, 30-31, 33, male, Taiwan.
Kobayashi 1985a, 12, Tsushima.
Tanida 2016, 95.
Cheumatopsyche davisi Oláh and Johanson 2008, 190-192, male, Taiwan. Synonymized by Malicky (2014).
Type locality. Taiwan.
Distribution. Japan (Tsushima), Taiwan.
Genus Hydromanicus Brauer
Tanida (1997a, 2003) recorded this genus from Okinawa-jima and Yaeyama
islands by the male genitalic morphology in mature pupae. Although he recognized
two species in the Ryukyus, precise information of their morphology has
not yet been provided.
Genus Hydropsyche Pictet
Thirteen species belonging to this genus have been recorded from Japan.
Hydropsyche albicephala Tanida 1986
Hydropsyche sp. HC: Tsuda and Akagi 1974, 133, larva. Mentioned by Tanida (1986a, 1991).
Tanida 1985, 179-180, larva.
Hydrpsyche sp.: Tanida 1982b, 18-20, Honshu (Ishikawa). Mentioned by Tanida (1986a).
Hydropsyche albicephala ms. (nomen nudum): Tanida 1984, 116-128, larval microhabitat.
Hydropsyche albicephala Tanida, 1986a, 473-476, male, female, larva, Hokkaido (Rumoi, Ishikari), Honshu (Kanagawa, Nagano, Ishikawa, Shiga,
Kyoto), Kyushu (Ohita).
Ohtsuka & Tanida 1986, 4, Kyushu (Kumamoto).
Yamanouchi 1987, 35-36, Hokkaido (Tokachi).
Morita 1988, 4, 5, Honshu (Mie).
Nozaki 1988b, 39, Honshu (Kanagawa).
Tanida 1988, 179-180, larva.
Morita 1989, 39, Honshu (Mie).
Gyotoku and Nozaki 1991, 75, Kyushu (Fukuoka).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Hatta and Nozaki 1991, 200, Honshu (Gifu, Aichi).
Tanida et al. 1991, 56, adult, pupa, larva.
Kuhara 1992, 45, Hokkaido (Shiribeshi).
Morita 1993, 4, Honshu (Mie).
Tanida and Takemon 1993, 142, Honshu (Kyoto).
Kuranishi and Kuhara 1994, 1225, Hokkaido (Kushiro).
Moriya 1994, 72, Honshu (Kanagawa, Yamanashi).
Itou and Muramatsu 1995, 39, Hokkaido (Okhotsk).
Kuranishi and Ohtaka 1996, 14, Honshu (Aomori).
Ito et al. 1997, 23, 27, Hokkaido (Hiyama).
Kuhara 1997a, 59, Hokkaido (Shiribeshi).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Moriya 1997, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nozaki 1997a, 35, Honshu (Kanagawa).
Nozaki 1997b, 28, Honshu (Tokyo).
Ito et al. 1998, 10, 12, 15, Hokkaido (Nemuro).
Kadowaki 1998, 5, Honshu (Shimane).
Kagaya et al. 1998, 25, 47-48, 147, Honshu (Tokyo).
Nishimoto and Nishimoto 1998, 150, Honshu (Aichi).
Ito et al. 1999, 23, 25-27, Hokkaido (Kushiro).
Ito et al. 2000a, 23, Hokkaido (Ishikari).
Kuhara et al. 2000, 246, Hokkaido (Iburi).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001b, 338, Honshu (Mie).
Uchida 2002, 77, Honshu (Tochigi).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Kawase et al. 2004, 110-111, Honshu (Ibaraki).
Kuranishi and Satake 2004, 35, Hokkaido (Okhotsk).
Morita and Kawase 2004, 17, Honshu (Ishikawa).
Nozaki 2004, 1230, Honshu (Kanagawa).
Ishiwata et al. 2005, 242-244, Honshu ((Kanagawa).
Katsuma 2005a, 5, Honshu (Ibaraki).
Nozaki and Nakamura 2005, 222, Honshu (Shimane).
Tanida 2005, 484, 486, 489-490, male, female, larva.
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Kuhara et al. 2006, 4545-46, Hokkaido (Ishikari).
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Kuhara et al. 2007, app. 70, Hokkaido (Ishikari).
Morita 2007a, 14, Honshu (Mie).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Morita and Kawase 2008, 55, Honshu (Ishikawa).
Muramatsu 2008, 5-6, Hokkaido (Okhotsk).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Morita and Kawase 2009, 28, Honshu (Ishikawa).
Tanaka 2009c, 18, Honshu (Akita).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito 2010, 131, Hokkaido (Kushiro).
Ito et al. 2010, 61, Hokkaido (Shiribeshi, Hiyama, Oshima).
Katsuma 2010b, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 42, Honshu (Mie, Shiga).
Kobayashi 2010, 140, Honshu (Tochigi).
Tanaka 2010a, 24, 27-28, male, Honshu (Akita).
Tanaka 2010b, 17, Honshu (Akita).
Tsuji et al. 2010, 5, Hokkaido (Okhotsk).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Katsuma 2011b, 50-51, Honshu (Ibaraki).
Katsuma 2011c, 67, Honshu (Ibaraki).
Kuhara 2011, 51, 55, 59, 70, Hokkaido (Sorachi, Ishikari, Iburi).
Katsuma 2012b, 65, Honshu (Ibaraki).
Kawase and Hayashi 2014, 256, Honshu (Shimane).
Moriya and Kubota 2012, 312, Honshu (Kanagawa).
Ito 2013, 56, Hokkaido (Nemuro, Kushiro).
Iijima and Tsuchiya 2014, 29, Hokkaido (Okhotsk, Kushiro, Iburi).
Ito and Nagasaka, 2014, 11, Hokkaido (Okhotsk).
Kawase and Morita 2014, 5-6, Honshu (Shiga).
Nozaki et al. 2014, 61, Honshu (Nagano).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Morita and Kawase 2016, 21-23, Honshu (Ishikawa).
Nozaki 2016, 76, 350-351, 435, 450, male, Shikoku (Kochi).
Tanaka 2016a, 22, 25, male. Honshu (Akita).
Tanida 2016, 95-96.
Imasaka et al. 2017, 61, Kyushu (Fukuoka).
Ito 2017b, 42, Hokkaido (Kamikawa).
Kawase et al. 2017, 31, Honshu (Shiga).
Nojima 2017, 119, Honshu (Okayama).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka).
Tanaka 2017, 2, Honshu (Akita).
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Tanida 2018, 572, 579, 580, male, fenale, larva.
Suzuki 2019a, 117, Honshu (Gifu).
Suzuki 2019b, 65, Honshu (Gifu).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Shimura and Imasaka 2021, 4, Shimokoshiki-jima.
Muramatsu 2022, 23, male, female, Hokkaido (Okhotsk).
Tanaka 2022, 35, Honshu (Akita).
Aoya and Tanaka 2023, Honshu (Akita).
Tanaka 2023, 41, Honshu (Akita).
Tanaka 2024, 26, Honshu (Akita).
Hydropsyche nakaharai: Kobayashi 1964, 87 (a part), Hokkaido (Okhotsk, Tokachi). Misidentificatiom
mentioned by Ito and Kuhara (2023).
Type locality. Honshu (Ishikawa), Japan.
Distribution. Japan (Hokakido, Honshu, Shikoku, Kyushu, Koshiki-jima).
Hydropsyche ancorapunctata Tanida 1986
Hydropsche sp. HA: Akagi 1955, 19, larva. Mentioned by Tanida (1986a, 1991).
Tsuda & Akagi 1962,
Tanida 1985, 180, larva.
Hydropsyche selysi: Tanida 1980, 98-109, size of larval head, life history, distribution,
Honshu (Kyoto). Misidentification mentioned by Tanida (1984).
Hydropsyche ancorapunctata ms (nomen nudum): Tanida 1984, 123-126.
Hydropsyche ancorapunctata Tanida 1986b, 629-630, larva, Honshu (Ishikawa, Kyoto)
Tanida 1988, 180, larva.
Morita 1990, 31, Honshu (Mie).
Hatta and Nozaki 1991, Honshu (Aichi).
Tanida et al. 1991, 54, adult, pupa, larva.
Morita 1993, 4, Honshu (Mie).
Moriya 1994, 72, Honshu (Yamanashi).
Moriya 1997, 63, Hosnhu (Yamanashi).
Nozaki 2004, 1230, Honshu (Kanagawa).
Tanida 2005, 484, 486, 489, larva.
Morita 2007b, 96, Honshu (Mie).
Inazu and Nishida 2011, larva, Honshu (Hyogo).
Morita and Kawase 2012, 15, Honshu (Ishikawa).
Tanida 2016, 96.
Tanida 2016, 573, 579, larva.
Yamamoto 2021, 74, Shikoku (Ehime).
Nojima 2022, 38, Honshu (Okayama).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu).
Remarks. This species was originally described based only on the larval stage (Tanida
1986). Although the adults have been recorded from several localities in
central Honshu, their genitalic morphology has not yet been described.
Hydropsyche buyssoni Ulmer 1907
Hydropsyche buyssoni Ulmer 1907c, 23-24, male, Honshu (Tochigi, Tokyo).
Fischer 1963, 23.
Tanida 2016, 96.
Type locality. Honshu (Tochigi, Tokyo).
Distribution. Japan (Honshu).
Remarks. Tanida (1987) treated this species as "incertae sedis".
Hydropsyche dilatata Tanida 1986
Hydropsyche gifuana: Tanida 1980, 98-109, size of larval head, life history, distribution,
Honshu (Kyoto). Misidentification mentioned by Tanida (1984).
Hydropsyche selysi: Tanida 1982b, 18, 20, 27-28, Honshu (Ishikawa). Misidentification mentioned
by Tanida (1984).
Hydropsyche dilatata ms. (nomen nudum): Tanida 1984, 116-128, larval microhabitat.
Hydropsyche dilatata Tanida 1986b, 627-629, male, female, Honshu (Nagano, Ishikawa, Kyoto, Osaka, Nara, Shimane).
Morita 1988, 4, Honshu (Mie).
Morita 1989, 39, Honshu (Mie).
Yagi et al. 1989, peculiar organ of female, 178-179, 181, Honshu (Kyoto).
Morita 1990, 31, Honshu (Mie).
Kugo and Nozaki 1990 62, Honshu (Hyogo).
Nozaki and Gyotoku 1990, 16, Kyushu (Fukuoka).
Gyotoku and Nozaki 1991, 75, Kyushu (Fukuoka).
Hatta and Nozaki 1991, 200, Honshu (Gifu, Aichi).
Tanida 1991a, 34, Honshu (Nara).
Morita 1993, 4, Honshu (Mie).
Tanida et al. 1995, 2, Honshu (Nara).
Kadowaki 1998, 5, Honshu (Shimane).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Morita 2001a, 220, Honshu (Mie).
Nozaki and Nakamura 2002, 170, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Nozaki and Nakamura 2005, 222, Honshu (Shimane).
Tanida 2005, 484, 486, 489, 492, male, female, larva
Mey and Nozaki 2006, 24, Honshu (Gifu).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Tanida 2007, 297-306, microdistribution.
Inazu 2008, 76-78, 93, 102, Honshu (Hyogo).
Morita 2008c, 723, Honshu (Mie).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Kameyama et al. 2009, 112, Honshu (Shimane).
Kawase and Morita 2010, 42, Honshu (Shiga).
Kawase et al. 2010, 108, Honshu (Shimane).
Mano 2010, 21-23, Honshu (Aichi).
Inazu and Nishida 2011, 193, male, larva, Honshu (Hyogo).
Inazu 2013b, 73, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nishimoto and Nishimoto 2014, 65, adult, Honshu (Aichi).
Imasaka et al. 2016, 34, Kyushu (Fukuoka).
Morita and Kawase 2016, 21, Honshu (Ishikawa).
Nozaki 2016, 76, 350-351, 435, 450, male, Shikoku (Kochi).
Tanida 2016, 96.
Ito 2017a, 5, Honshu (Fukui).
Kawase et al. 2017, 31, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Nojima 2017, 119, Honshu (Okayama).
Tanida 2018, 573, 579, 582, male, female, larva.
Tanida et al. 2018, 30, Honshu (Toyama).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Kuhara et al. 2023, 16, Honshu (Fukui), Kyushu (Oita).
Type locality. Honshu (Nara).
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. In Tanida (2005, 2018), figures of adults of this species and those of
H. selysi are mistakenly replaced each other (Kuhara et al. 2023). A larva described as Hydropsyche sp. HE by Tanida (1985) includes this species and H. selysi (Tanida, 1988, 1991b). Shimura (2015) recorded this species from Fukue-jima
based on identification of the larval stage.
Hydropsyche gifuana Ulmer 1907
Hydropsyche gifuana Ulmer 1907a, 68, male, Japan.
Tsuda 1940, 25-26, Honshu (Kyoto, Hyogo).
Tsuda 1942a, 279, Honshu (Kyoto).
Tsuda 1942b, 263-267, Honshu (Kyoto).
Tsuda 1943, 166-168, 170, Honshu (Kyoto).
Tsuda 1950, 417, male.
Tsuda and Tatsumi 1951, 35, male.
Miwa and Okawa 1958, 18, Honshu (Mie).
Kobayashi 1959, 349, Kyushu (Fukuoka)
Tsuda and Akagi 1962, 133, larva.
Fischer 1963, 33.
Kuwada 1965, 163, pl. 82, male, Honshu (Ehime).
Tani 1977, 196, male.
Tanida and Takemon 1981, 206, Honshu (Kyoto).
Kobayashi 1982, 96, Honshu (Kanagawa).
Tanida 1985a, 180, larva.
Adachi et al. 1986, 86-90, larval mandible.
Ohtsuka & Tanida 1986, 7, Kyushu (Kumamoto).
Tanida 1986b, 630-633, male, female, larva, Honshu (Kanagawa, Kyoto, Hyogo), Shikoku (Kochi).
Kobayashi 1987, 18, Honshu (Shimane).
Yagi et al. 1989, peculiar organ of female, 178, 181, Honshu (Kyoto).
Nozaki and Gyotoku 1990, 16, Kyusyu (Fukuoka).
Gyotoku and Nozaki 1991, 75, pl. 9, adult, Kyushu (Fukuoka).
Moriya 1992, 41, Honshu (Kanagawa).
Morita 1993, 4, Honshu (Mie).
Tani and Nakamura 1997, 60, Honshu (Hiroshima).
Kagaya et al. 1998, 25, 48, 150, Honshu (Tokyo).
Morita 1998, 72, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 170, Honshu (Hiroshima).
Nozaki 2004, 123-1231, Honshu (Kanagawa).
Yoshitaka 2004, 28, Honshu (Okayama).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Katsuma 2005a, 5, Honshu (Ibaraki).
Katsuma 2005b, 13, Honshu (Gumma).
Tanida 2005, 484, 486, 489, 492, male, female, larva.
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Kimura et al. 2006, 7, Honshu (Nagano).
Nakamura and Fukui 2006, 243, Honshu (Hiroshima).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nakamura et al. 2006, 101, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Tanida 2008, 250, pl. 92, adult, Shikoku (Ehime).
Kameyama et al. 2009, 112, Honshu (Shimane).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Inazu and Nishida 2011, 194, male, female, Honshu (Hyogo).
Kawase and Hayashi 2011a, 173, Honshu (Shimane).
Morita 2012, 250, Honshu (Mie).
Nozaki 2016, 76, 350-351, 435, 450, male, Shikoku (Kochi).
Tanida 2016, 96.
Nojima 2017, 119, Honshu (Okayama).
Tanida 2018, 573, 579, 582, male, female, larva.
Morita 2020a, 43, Honshu (Mie).
Kuhara et al. 2023, 16, Honshu (Akita, Fukushima).
Hydropsyche drouardi Navás 1933, 96, Honshu (Kofu). Synonymized by Tanida (1986b).
Fischer 1963, 28.
Hydropsyche gifuana?: Akagi 1955, 18-19, larva.
Type locality. Japan (precise locality unknown).
Distribution. Japan (Honshu, Shikoku, Kyushu).
Hydropsyche isip Arefina, Minakawa & Nozaki 2004
Hydropsyche isip Arefina et al, 2004, 210-211, male, Sakhalin.
Nakano et al. 2007, 39-40, Hokkaido (Nemuro).
Ito 2013, 56, Hokkaido (Nemuro, Kushiro).
Tanida 2016, 96.
Tanida 2018, 573, 583, male, female.
Ito and Kuhara 2023, 54, Hokkaido (Soya).
Hydropsyche orientalis: Ito et al. 2000b, 26, 28-30, Hokkaido (Soya). Misidentification mentioned by Ito
and Kuhara (2003).
Hydropsyche sp. a: Ito et al., 2000b, 22, 28-30, Hokkaido (Soya). Mentioned by Ito and Kuhara (2003).
Type locality. Sakhalin.
Distribution. Japan (Hokkaido), Sakhalin.
Hydropsyche kozhantschikovi Martynov 1924
Hydropsyche kozhantschikovi Martynov 1924, 87-88, 97, male, Russia (South Siberia).
Kuhara et al. 2006, 45-46, 48, male, Hokkaido (Ishikari).
Kuhara et al. 2007, app. 70, Hokkaido (Ishikari).
Tanida 2016, 96.
Tanida 2018, 573, 583, male.
Hydropsyche kawamurai Tsuda 1940, 27-28, 33, male, Korea. Synonymized by Oláh and Johanson (2008).
Kobayashi 1959, 349, Kyushu (Fukuoka).
Type locality. Russia.
Distribution. Japan (Hokkaido, Kyushu), Russia, Korea, China, .Mongolia, Kazakhstan,
Pakistan.
Remarks. Tanida (2016) included Kuril Islands and Honshu as dostribution areas,
no reliable records are found from there.
Hydropsyche newae Kolenati 1958
Hydropsyche newae Kolenati 1858, 278, male, Russia (South Siberia).
Tanida 1986a, 477-479, male, female, larva, Hokkaido (Okhotsk, Tokachi, Rumoi, Kamikawa, Ishikari) (nevae).
Yamanouchi 1987, 35, Hokkaido (Tokachi) (nevae).
Yagi et al. 1989, peculiar organ of female, 177, 181, Hokkaido (nevae).
Nishimoto 1993, 22, Honshu (Aichi) (nevae).
Kuhara and Ito 1994 20, Hokkaido (Ishikari) (nevae).
Kuranishi and Kuhara 1994, 1225, Hokkaido (Kushiro) (nevae).
Itou and Muramatsu 1995, 39, Hokkaido (Okhotsk) (nevae).
Tanida 1988, 179, larva (nevae).
Itou 1989, 55, Hokkaido (Kushiro) (nevae).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Ito et al. 1999, 23, 25, Hokkaido (Kushiro) (nevae)
Ito et al. 2000a, 23, Hokkaido (Ishikari) (nevae).
Nagayasu et al. 2003, 106, Hokkaido (Ishikari) (nevae).
Nagayasu et al. 2005, 53, Hokkaido (Ishikari) (nevae).
Tanida 2005, 484-485, 489-490, male, female, larva (nevae).
Ito et al. 2007, 148, Hokkaido (Kushiro).
Muramatsu 2008, 6, Hokkaido (Okhotsk).
Nakatani et al. 2008, Hokkaido (Okhotsk).
Ito 2013, 56-57, Hokkaido (Kushiro, Tokachi).
Ijima and Tsuchiya 2014, 29, Hokkaido (Okhotsk, Kushiro, Tokachi).
Ito and Nagasaka, 2014, 11, Hokkaido (Okhotsk).
Ito and Kuhara 2016, 16-17, Hokkaido (Okhotsk).
Tanida 2016, 96.
Ito 2017, 42, Hokkaido (Kamikawa).
Tanida 2018, 572, 579, 580, male, female, larva.
Muramatsu 2022, 22, 55, male, female, larva, Hokkaido (Okhotsk).
Ito and Kuhara 2023, 54-55, Hokkaido (Soya, Kamikwa, Rumoi).
Hydropsyche sp. HB: Akagi 1962, 42, larva, male genitalia, Hokkaido (Okhotsk). Mentioned by Tanida (1986a, 1991b).
Tsuda and Akagi 1974, 133, larva.
Tanida 1985, 179-180, larva.
Type locality. Russia.
Distribution. Japan (Hokkaido), Russia, Korea, China, Europe.
Hydropsyche orientalis Martynov 1934
Hydropsyche orientalis Martynov 1934, 276-277, male, Russia (Southeast Siberia).
Tanida 1982b, 18-21, 27, Honshu (Ishikawa).
Tanida 1985, 179-180, larva.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Adachi et al. 1986, 86-87, 90, larval mandibles.
Ohtsuka and Tanida 1986, 4-5, Kyushu (Kumamoto).
Tanida 1986a, 468-473, male, female, larva, Hokkaido (Okhotsk, Tokachi, Rumoi, Ishikari, Oshima), Honshu (Tochigi,
Gumma, Nagano, Ishikawa, Gifu, Shiga, Kyoto, Hyogo, Nara, Shimane), Shikoku
(Tokushima, Kochi), Amami-oshima, Okinawa-jima.
Kobayashi 1987, 17-18, Honshu (Shimane).
Nozaki 1987, 41, Honshu (Kanagawa).
Morita 1988, 4, 5, Honshu (Mie).
Nozaki 1988a, 45-46, Honshu (Kanagawa).
Nozaki 1988b, 39, Honshu (Kanagawa).
Morita 1989, 39, Honshu (Mie).
Yagi et al. 1989, peculiar organ of female, 177-178, 181, Honshu (Kyoto).
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Nozaki and Gyotoku 1990, 13, 16, Kyushu (Fukuoka).
Tanida 1990, 55, Honshu (Hyogo).
Gyotoku and Nozaki 1991, 75, Kyushu (Fukuoka).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Hatta and Nozaki 1991, 200-201, 208, adult, Honshu (Gifu, Aichi).
Tanida 1991a, 33-34, Honshu (Nara).
Tanida et al. 1991, 50, adult, pupa, larva.
Moriya 1992, 41, Honshu (Kanagawa).
Hori et al. 1993, 388, Honshu (Aichi).
Kuhara et al. 1993, 17, Hokkaido (Ishikari).
Morita 1993, 4, Honshu (Mie).
Nishimoto 1993, 22, Honshu (Aichi).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Kuhara and Ito 1994, 19, Hokkaido (Ishikari).
Kuranishi and Kuhara 1994, 1224, Hokkaido (Kushiro).
Morita 1994, 87, Honshu (Mie).
Moriya 1994, 72-73, Honshu (Kanagawa, Yamanashi).
Tanida et al. 1995, 2, Honshu (Nara).
Tanida and Takemon 1993, 242, 245, Honshu (Kyoto).
Kuranishi and Ohtaka 1996, 14, Honshu (Aomori).
Saito 1996, 15, Hokkaido (Kamikawa).
Saito et al. 1996, 21, Hokkaido (Kushiro).
Ito et al. 1997, 23, 27, Hokkaido (Hiyama).
Kuhara 1997a, 59, Hokkaido (Shiribeshi).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 13, 15, 18-19, Hokkaido (Oshima).
Nozaki 1997a, 35, Honshu (Kanagawa).
Nozaki 1997b, 29, Honshu (Tokyo).
Tani and Nakamura 1997, 60, Honshu (Hiroshima).
Ito et al. 1998, 10, 15, Hokkaido (Nemuro).
Kadowaki 1998, 4-5, Honshu (Shimane).
Kagaya et al. 1998, 25, 48-49, 149, Honshu (Tokyo, Yamanashi).
Morita 1998, 71-72, Honshu (Mie).
Nishimoto and Nishimoto 1998, 149, Honshu (Aichi).
Ito et al. 1999, 23, 25-27, Hokkaido (Kushiro).
Nakano and Tanida 1999, 276, Honshu (Nara).
Ohkawa 1999, 44, Hokkaido (Tokachi).
Ito et al. 2000a, 23-24, Hokkaido (Ishikari).
Kuhara et al. 2000, 246, Hokkaido (Iburi).
Nishimoto 2000, 26, Honshu (Aichi).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Kuranishi 2002, 566, adult, larva, habitat.
Nakamura et al. 2002, 67, Honshu (Hiroshima).
Nozaki and Nakamura 2002, 170, Honshu (Hiroshima).
Uchida 2002, 78, Honshu (Tochigi).
Nagayasu et al. 2003, 106-107, Hokkaido (Ishikari).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Tanida 2003, 381, male, female, larva.
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Ito et al. 2004, 11-12, Hokkaido (Kushiro).
Kawase et al. 2004, 111, Honshu (Ibaraki).
Kuranishi and Satale 2004, 35, Hokkaido (Okhotsk).
Morita and Kawase 2004, 16, Honshu (Ishikawa).
Nozaki 2004, 1231, Honshu (Kanagawa).
Yoshitaka 2004, 28, Honshu (Okayama).
Ishiwata et al. 2005, 110, 242-244, Honshu (Kanagawa).
Katsuma 2005a, 5, Honshu (Ibaraki).
Katsuma 2005b, 13, Honshu (Gumma).
Nagayasu et al. 2005, 53, Hokkaido (Ishikari).
Nozaki 2005, 108, adult, larva.
Nozaki and Nakamura 2005, 222, Honshu (Shimane).
Satake and Kuranishi 2005, 377, (Kouzu Is., Mikura Is. Hachijo Is.)
Tanida 2005, 483-490, male, female, larva.
Kameyama and Nakamura 2006, 189, Honshu (Tottori).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Kuhara et al. 2006, 45-46, Hokkaido (Ishikari).
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Ito and Kosugi 2007, 53, Hokkaido (Kushiro).
Ito et al. 2007, 148, Hokkaido (Kushiro).
Katsuma 2007, 18, Honshu (Ibaraki).
Kimura et al. 2007, 7, Honshu (Nagano).
Kuhara et al. 2007, app. 70, Hokkaido (Ishikari).
Nakamura and Hajiri 2007, 198, Honshu (Shimane).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nakano et al. 2007, 39-40, Hokkaido (Nemuro).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Nozaki and Tanida 2007, 248, Honshu (Shizuoka).
Tanida 2007, 297-306, microdistribution.
Inazu 2008, 78, 93, 102, Honshu (Hyogo).
Katsuma 2008, 58, Honshu (Ibaraki).
Kimura et al. 2008, 264, Honshu (Nagano).
Morita 2008c, 723, Honshu (Mie).
Muramatsu 2008, 5, Hokkaido (Okhotsk).
Tanida 2008, 250, pl. 92, adult, Shikoku (Ehime).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka).
Kamaeyama et al. 2009, 112, Honshu (Shimane).
Tanaka 2009a, 46, adult, Honshu (Akita).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito et al. 2010, 61, Hokkaido (Shiribeshi, Hiyama, Oshima).
Katsuma 2010a, 84, Honshu (Ibaraki).
Kawase and Hayashi 2010a, 80, Honshu (Shimane).
Kawase and Morita 2010, 42, Honshu (Shiga).
Kobayashi 2010, 140, Honshu (Tochigi).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura and Kita 2010, 88, Honshu (Hiroshima).
Nozaki 2010, 216-217, Honshu (Kanagawa).
Tanaka 2010a, 24, Honshu (Akiata).
Tanaka 2010b, 17, Honshu (Akita).
Tsuji et al. 2010, 5, Hokkaido (Okhotsk).
Inazu and Nishida 2011, 193, male, larva, Honshu (Hyogo).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Katsuma 2011b, 49-50, Honshu (Ibaraki).
Katsuma 2011c, 67-68, Honshu (Ibaraki).
Katsuma 2011d, 69, Honshu (Ibaraki).
Kawase and Morita 2011b, 192, Honshu (Shimane).
Kuhara 2011, 51,55, 59, 70, Hokkaido (Sorachi, Ishikari).
Morita 2011, 98, Honshu (Mie).
Nozaki 2011, 4, Honshu (Aichi).
Tanaka and Okura 2011, 24, Honshu (Akita).
Tsukiji 2011, 140, adult.
Inazu 2012, 8, Honshu (Hyogo).
Kawase and Hayashi 2012, 256-257, Honshu (Shimane), Oki (Dogo).
Morita and Kawase 2012, 15, Honshu (Ishikawa).
Moriya and Kubota 2012, 132, Honshu (anagawa).
Hayashi et al. 2013, 13, Oki (Dogo).
Inazu 2013a, 57, Honshu (Hyogo).
Inazu 2013b, 73, Honshu (Hyogo).
Ito 2013, 57, Hokkaido (Nemuro, Okhotsk, Kushiro).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Takai et al. 2013, 84, adult.
Ijima and Tsuchiya 2014, 29-30, Hokkaido (Okhotsk, Kushiro).
Katsuma 2014, 45, Honshu (Ibaraki).
Kawase and Morita 2014, 6, Honshu (Shiga).
Kuranishi and Kikuchi 2014, 363, Honshu (Yamagata).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Nozaki 2014, 61, Honshu (Nagano).
Ito and Nagasaka, 2014, 11, Hokkaido (Okhotsk).
Shimura 2014, 43, Amami-oshima
Katsuma 2015, 91, Honshu (Fukushima).
Shimura 2015, 51, Fukue-jima.
Hirabayashi et al. 2016, 208, 210, Honshu (Nagano).
Imasaka et al. 2016, 34, Kyushu (Fukuoka).
Ito and Kuhara 2016, Hokkaido (Kushiro).
Morita and Kawase 2016, 23, Honshu (Ishikawa).
Nakamura 2016, 21, 23, Honshu (Hiroshima).
Nozaki 2016, 76, 298, 303, 345, 350-351, 435, 450, male, shikoku (Kochi).
Tanaka 2016a, 23, 25, male, Honshu (Akita).
Tanida 2016, 96-97.
Uchifune 2016, 200-201, adult, Honshu (Kanagawa).
Imasaka et al. 2017, 61, Kysuhu (Fukuoka).
Ito 2017a, 5, Honshu (Fukui).
Ito 2017b, 42, Hokkaido (Kamikawa).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kawase et al. 2017, 31-32, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Kuhara and Ito 2017, 14-15, Yaku-shima.
Matsuoka 2017, 8, Hokkaido (Tokachi).
Nojima 2017, 119, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Tanaka 2017, 2, Honshu (Akita).
Ito and Nozaki 2018, 558, Amami-o-shima, Okinawa-jima, Kume-jima.
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima). (larval record)
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka)
Tanaka 2018a, 21, Honshu (Akita).
Tanaka 2018b, 8-9, Honshu (Akita).
Tanida 2018, 572, 578, 579, 580, larva, male, female.
Tsuruda 2018, 27, Honshu (Tokyo).
Nozaki et al. 2019, 169, Honshu (Nagano).
Tanaka 2019a, 27, Honshu (Akita).
Tanaka 2019b, 12, Honshu (Akita).
Yamamoto et al. 2019, 72, Shikoku (Ehime).
Morita 2020a, 18, Honshu (Mie).
Morita 2020d, 81, Honshu (Mie).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yashiro et al. 2020, 185, Kyushu (Kamikoshiki-jima, Shimokoshiki-jima).
Shimura and Imasaka 2021, 4-5, Kamikoshiki-jima, Shimokoshiki-jima.
Tezuka et al. 2021, 43, Honshu (Kanagawa).
Yamamoto and Nozaki 2021, 128, Shikoku (Ehime).
Muramatsu 2022, 23, 55, male, female, larva.
Tanaka 2022, 35-36, Honshu (Akita).
Aoya and Tanaka 2023, 27, Honshu (Akita).
Ito and Kuhara 2023, 55, Hokkaido (Soya, Kamikwa, Rumoi).
Tanaka 2023, 41-42, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).
Tanaka 2024, 26, Honshu (Akita).
Hydropsyche newae: Ulmer 1907a, 64, male, Honshu (Gifu) (nevae). Misidentification. Tsuda (1940) proposed a new name as H. ulmeri.
Hydropsyche ulmeri Tsuda 1940, 26-27, 33, male, Honshu (Nagano, Shiga, Kyoto). Junior homonym of an African species, and renamed as H. tsudai by Tani (1977).
Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942a, 279-280, Honshu (Kanagawa, Nagano, Kyoto), Kyushu (Fukuoka).
Tsuda 1942b, 263-265, 267, Honshu (Kyoto).
Tsuda 1943, 166-167, 170, Honshu (Kyoto).
Tsuda 1950, 417, male.
Tsuda and Tatsumi 1951, 28-29, 31-33, 35-41, male, larva, pupa, Honshu (Niigata)
Ishihara et al. 1953, 28, Shikoku (Ehime).
Akagi 1955, 18, larva.
Chihara 1955, 23, pupa.
Tsuda 1959, 140, larva.
Tsuda and Akagi 1962, 133, 144, larva, pupa.
Kuwada 1965, 163, male, pl. 82, Shikoku (Ehime).
Kobayashi 1971, 32-33, pl. 20, male, Honshu (Kanagawa).
Kobayashi 1973, 31, Honshu (Yamagata).
Shibata 1975, 12-75, male, larva, pupa, life history etc., Honshu (Nagano).
Kobayashi 1983, 32-33, Honshu (Kanagawa).
Kobayashi 1984c, 3, Sado.
Kobayashi 1984d, 28, 30-31, Honshu (Kanagawa).
Kobayashi 1985b, 68-73, Honshu (Kanagawa).
Nishiharu 1997a, 60, Honshu (Fukui).
Sekimoto et al. 1998, 360, Honshu (Hiroshima).
Yodoe et al. 1998, 68, Honshu (Shimane).
Nakamura et al. 2006, 101, Honshu (Hiroshima).
Hydropsyche kawamurai: Kobayashi 1964, 86, Hokkaido (Okhotsk). Misidentrification mentioned
by Ito and Kuhara (2023).
Hydropsyche nakaharai: Kobayashi 1964, 87 (a part), Hokkaido (Okhotsk, Tokachi). Misidentificatiom
mentioned by Ito and Kuhara (2023).
Hydropsyche tsudai Tani 1977, 196, pl. 43, male, Honshu (Nara). Synonymized by Tanida (1982b).
Tani 1978, 6, Honshu (Niigata).
Yamanouchi 1978, 10, Hokkaido (Tokachi).
Tanida 1980a, 136, Honshu (Ishikawa).
Tanida 1980b, 98-109, size of larval head, life history, distribution,
Honshu (Kyoto)
Tanida and Takemon 1981, 206, Honshu (Kyoto)..
Tanida 1982a, 357, Honshu (Gumma).
Hydropsyche sp. (ulmeri auct.): Tani 1976, 392, Tsushima.
Type locality. Russia.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyusyu, Sado, Kouzu Is., Mikura Is.,
Hachijo Is., Oki (Dogo), Koshiki-jima, Amami-oshima, Okinawa-jima, Kume-jima),
Russia, Korea, China.
Hydropsyche orbiculata Ulmer 1911
Hydropsyche oribiculata Ulmer 1911, 397-399, male, Taiwan.
Malicky and Chantaramongkol 2000, 812, 851, pl. 25, male, Taiwan.
Ito and Nozaki 2018, 558-560, male, female, Ishigaki-jima, Iriomote-jima, Taiwan.
Hydrospyche yaeyamensis ms. (nomen nudum): Tanida 1984, 123-126.
Hydropsyche sp. HD: Tanida 1985, 179, larva. Mentioned by Tanida (1986a).
Hydropsyche yaeyamensis Tanida 1986a, 479-482, male, female, larva, Ishigaki-jima, Iriomote-jima. Synonymized by Ito and Nozaki (2018)
Tanida 1988, 179, larva.
Yagi et al. 1989, peculiar organ of female, 177-178, 181, Ishigaki-jima.
Tanida 2003, 381, male, female, larva.
Tanida 2005, 484-485, 489, 491, male, female, larva.
Tanida 2016, 97.
Tanida 2018, 573, 579, 581, male, female, larva.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima, Iriomote-jima), Taiwan.
Hydropsyche selysi Ulmer 1907
Hydropsyche selysi Ulmer 1907, 67-68, male, Japan (Gifu, Oiwake (Nagano?)).
Tsuda 1942a, 279, Honshu (Shiga, Kyoto).
Tsuda 1943, 168, Honshu (Kyoto).
Tsuda 1950, 417, male.
Asahina 1951, egg, 1st instar larva, pupa, pupal case, life history, Honshu (Shizuoka).
Tsuda and Tatsumi, 1951, 29, 31, 35, male, Honshu (Niigata).
Fischer 1963, 84-85.
Tani 1977, 196, male.
Tani 1978, 5, Honshu (Niigata).
Kobayashi 1982, 96, Honshu (Kanagawa).
Kobayashi 1983, 32-33, Honshu (Kanagawa).
Kobayashi 1985b, 68, Honshu (Kanagawa).
Tanida 1986b, 624-627, male, female, larva, Honshu (Kanagawa).
Kobayashi 1987, 18, Honshu (Shimane).
Yagi et al. 1989, peculiar organ of female, 178, 181, Honshu (Yamagata).
Tanida et al. 1991, 52, adult, pupa, larva.
Nishimoto and Nishimoto 1993, 86, 92, adult, Honshu (Aichi).
Moriya 1994, 73, Honshu (Kanagawa).
Moriya 1997, 63, Honshu (Kanagawa).
Nozaki 1997a, 35, Honshu (Kanagawa).
Kadowaki 1998, 5, Honshu (Shimane).
Kagaya et al. 1998, 25, 49, 150-151, Honshu (Tokyo).
Kawase et al. 2004, 111, Honshu (Ibaraki).
Nozaki 2004, 1231, Honshu (Kanagawa).
Tanida 2005, 484, 486, 489, 491, male, larva.
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Nozaki 2010, 217, Honshu (Kanagawa).
Moriya and Kubota 2012, 312, Honshu (Kanagawa).
Tanida 2016, 97.
Tanida 2018, 573, 579, 581, male, larva.
Kuhara et al. 2023, 16, Honshu (Fukushima, Nagano).
Hydropsyche ciliosa Navás 1933, 97, male, Honshu (Yamanashi). Synonymized by Tanida (1986b).
Fischer 1963, 24.
Type locality. Japan (Gifu?).
Distribution. Japan (Honshu).
Remarks. In Tanida (2005, 2018), figures of adults of this species and those of
H. dilatataare mistakenly replaced each other (Kuhara et al. 2023). A larva described as Hydropsyche sp. HE by Tanida (1985) includes this species and H. dilatata (Tanida, 1988, 1991b).
Hydropsyche setensis Iwata 1927
Hydropsyche setensis Iwata 1927a, 223, 226, pl. 5, larva, Japan (Ishikawa, Nagano, Shiga, Hyogo, Tottori, Shimane) (larval record).
Iwata 1927b 206, pl. 3, larva, Japan (Ishikawa, Nagano, Shiga, Tottori, Shimane) (larval record).
Fischer 1963, 85.
Tanida 1987, 59-60, lectotype designation, Honshu (Shiga).
Nozaki 1988a, 45 Honshu (Kanagawa).
Nozaki 1988b, 39, Honshu (Kanagawa).
Yagi et al. 1989, peculiar organ of female, 177-179, 181, Honshu (Kyoto).
Tanida 1990, 55, Honshu (Hyogo).
Gyotoku and Nozaki 1991, 75, Kyushu (Fukuoka).
Hatta and Nozaki 1991, 200, Honshu (Gifu).
Tanida 1991a, 33, Honshu (Nara).
Moriya 1992, 41, Honshu (Kanagawa).
Hori et al. 1993, 388, Honshu (Aichi).
Moriya 1994, 73, Honshu (Kanagawa).
Moriya 1997, 63, Honshu (Kanagawa, Yamanashi).
Nishiharu 1997b, 38, Honshu (Fukui)
Nozaki 1997a, 35, Honshu (Kanagawa).
Nozaki 1997b, 29, Honshu (Tokyo).
Tani and Nakamura 1997, 60-61, Honshu (Hiroshima).
Kadowaki 1998, 5, Honshu (Shimane).
Kagaya et al. 1998, 25, 49, 152, Honshu (Tokyo, Yamanashi).
Morita 1998, 72, Honshu (Mie).
Nakano and Tanida 1999, 276, Honshu (Nara).
Nishimoto 2000 26, Honshu (Aichi).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 170-171, Honshu (Hiroshima).
Nakamura and Umemori 2003, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Ishiwata et al. 2005, 110, 242-244, Honshu (Kanagawa).
Katsuma 2005a, 5, Honshu (Ibaraki).
Katsuma 2005b, 13-14, Honshu (Gumma).
Tanida 2005, 484-485, 489, 494, male, female, larva.
Kameyama and Nakamura 2006, 189, Honshu (Tottori).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Kimura et al. 2006, 7, Honshu (Nagano).
Mey and Nozaki 2006, 24, Hoshu (Gifu).
Nakamura et al. 2006, 101, Honshu (Hiroshima).
Katsuma 2007, 18, Honshu (Ibaraki).
Nakamura and Tsujimura 2007, 77, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Kimura et al. 2008, 264, Honshu (Nagano).
Kameyama et al. 2009, 112, Honshu (Shimane).
Tanaka 2009c, 18, Honshu (Akita).
Kawase and Morita 2010, 42, Honshu (Shiga).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura and Kita 2010, 88, Honshu (Hiroshima).
Inazu and Nishida 2011, 193, male, larva, Honshu (Hyogo).
Kawase and Hayashi 2011a, 173, Honshu (Shimane).
Morita and Katsuma 2011, 98, Honshu (Mie).
Kawase and Hayashi 2012, 257, Honshu (Shimane).
Moriya and Kubota 2012, 312, Honshu (Kanagawa).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nozaki et al. 2014, 61, Honshu (Nagano).
Katsuma 2015, 91, Honshu (Ibaraki).
Hirabayashi et al. 2016, 208, 210, Honshu (Nagano).
Katsuma 2016, 47-48, Honshu (Ibaraki).
Nozaki 2016, 76, 351, 435, 450, male, Honshu (Hyogo).
Tanaka 2016a, 23, 25, male, Honshu (Akita).
Tanida 2016, 97.
Kawase et al. 2017, 32, Honshu (Shiga).
Kobayashi et al. 2017, 17, 19, 20, 24, color pattern of larval head, Honshu (Shiga, Kyoto).
Nojima 2017, 119, Honshu (Okayama).
Tanaka 2018b, 9, Honshu (Akita).
Tanida 2018, 573, 579, 581, male, female, larva.
Nozaki et al. 2019, 169, Honshu (Nozaki).
Morita 2019a, 22, Honshu (Mie).
Katsuma 2020, 15, Kyushu (Oita).
Kimura and Kuranishi 2020, 157, Honshu (Kyoto).
Tanaka 2022, 36, Honshu (Akita).
Kawase & Kato 2023, 3-4, Honshu (Niigata).
Kuhara et al. 2023, 16, Honshu (Aomori).
Tanaka 2023, 42, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).
Hydropsyche nakaharai Tsuda 1949, 20-21, male, Honshu (Niigata). Synonymized by Tanida (1987).
Tsuda and Tatsumi 1951, 28-29, 31, 32, 34-35, 37-41, male, larva, pupa, Honshu (Niigata).
Akagi 1955, 18, larva.
Chihara 1955, 22-23, pupa.
Fujimoto 1956, over wintering instar, Honshu (Kyoto) (larva).
Tsuda and Kawai 1956, 73-75, Honshu (Kyoto).
Tsuda 1959, 140, larva.
Tsuda & Akagi 1962, 133, larva.
Kuwada 1965, 164, male.
Kobayashi 1973, 31, Honshu (Yamagata).
Shibata 1975, 16-20, male, larva, pupa, Honshu (Nagano).
Yamanouchi 1976, 3-4, Hokkaido (Tokachi).
Tani 1977, 196, male.
Tani 1978, 6, Honshu (Niigata).
Kobayashi 1982, 96, Honshu (Kanagawa).
Kobayashi 1983, 32, 34-35, Honshu (Kanagawa).
Kobayashi 1984b, 17, 25, male, female, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kobayashi 1984c, 3, Sado.
Kobayashi 1984d, 28-29, 31, Honshu (Kanagawa).
Tanida 1985, 179, larva.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Tanida 1986a, 482-484, male, female, larva, Honshu (Tochigi, Kanagawa, Ishikawa, Kyoto, Nara, Hyogo, Shimane), Shikoku
(Tokushima).
Tanida et al. 1995, 2, Honshu (Nara).
Hydropsyche japonica Iwata 1927a, 223, 226, larva. Synonymized by Tanida (1987).
Iwata 1927b, 207, larva.
Fischer 1963, 72.
Tanida 1987, lectotyoe designation, Honshu (Shimane).
Type locality. Japan (Shiga).
Distribution. Japan (Hokkaido?, Honshu, Shikoku, Kyushu, Sado).
Remarks. Kuhara and Kuranishi (1997) and Ito et al. (2010) suggested that the record by Kobayshi (1984) from Hokkaido is
probably based on a misidentification of H. albicephala or H. orientalis. Kuhara and Kuranishi (1997) and Ito and Kuhara (2023) pointed out that
the records from Hokkaido by Yamanouchi (1976) are also doubtful.
Hydropsyche verrucosa (Ulmer 1911)
Hydromanicus verrucosus Ulmer 1911, 397, 399-400, male, Taiwan.
Malicky and Chantaramongkol 2000, 797, 829, pl. 3, male, Taiwan.
Hydatomanicus verrucosus: Ulmer, 1951, 299.
Hydropsyche verrucosa: Geraci et al. 2010, 926.
Ito and Nozaki 2018, 560-561, male, female, Amami-o-shima, Ishigaki-jima, Iriomote-jima.
Type locality. Taiwan.
Distribution. Japan (Amami-o-shima, Ishigaki-jima, Iriomote-jima), Taiwan.
Genus Potamyia Banks
Only one species is known from Japan.
Potamyia chinensis (Ulmer 1915)
Hydrospyche chinensis Ulmer 1915, 47-48, male, China (Beijing).
Hydropsyche echigoensis Tsuda 1949, 21-22, male, Honshu (Niigata). Synonymized by Oláh et al. (2006).
Tsuda 1950, 417, male.
Tsuda and Tatsumi 1951, 28-29, 31, 33-36, male, Honshu (Niigata).
Chihara 1956, 218-220, pupa, Honshu (Kyoto).
Akagi 1956c, 215-216, larva, Honshu (Kyoto, Osaka).
Kobayashi 1959, 349, Kyushu (Fukuoka).
Tsuda and Akagi 1962, 133, larva.
Kuwada 1965, 164, male.
Tani 1977, 196, male.
Tani 1978, 6, Honshu (Niigata).
Nakamura and Fukui 2006, Honshu (Hiroshima).
Cheumatopsyche echigoensis: Tanida, 1985, 180-181, 215, larva.
Gyotoku and Nozaki 1991, 76, Kyushu (Fukuoka).
Hatta and Nozaki 1991, 201, Honshu (Gifu).
Adachi et al. 1986, 86-87, 90, larval mandibles.
Yagi et al. 1989, peculiar organ of female, 177-178, 180-181, Honshu (Kyoto).
Tani and Nakamura 1997, 60, Honshu (Hiroshima).
Morita 1998, 72, Honshu (Mie).
Sekimoto et al. 1998, 360, Honshu (Hiroshima).
Kameyama et al. 2009, 111-112, Honshu (Shimane).
Potamyia echigoensis: Tanida 1997b, 110, 115-116.
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 171, Honshu (Hiroshima).
Nozaki and Nakamura 2005, 223, Honshu (Shimane).
Tanida 2005, 483, 485, 488, larva.
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Kimura et al. 2006, 7, Honshu (Nagano).
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Inazu 2008, 78, 93, 102, Honshu (Hyogo).
Potamyia chinensis: Tian & Li 1987, 128, combination.
Katsuma 2007, 18, Honshu (Ibaraki).
Nozaki and Nakamura 2007, 95-96, Honshu (Hiroshima).
Katsuma 2008, 58, Honshu (Ibaraki).
Kimura et al. 2008, 264, Honshu (Nagano).
Katsuma 2010a, 84, Honshu (Ibaraki).
Mano 2010, 22-23, Honshu (Aichi).
Inazu and Nishida 2011, 193, male, Honshu (Hyogo).
Katsuma 2011c, 68, Honshu (Ibaraki).
Katsuma 2011d, 69, Honshu (Ibaraki).
Katsuma 2014, 45, Honshu (Ibaraki).
Katsuma 2016, 48, Honshu (Ibaraki).
Nozaki 2016, 77, 346, 352, 436, male, Honshu (Ibaraki).
Tanaka 2016a, 23, 26, male, Honshu (Akita).
Tanida 2016, 97.
Kawase et al. 2017, 32, Honshu (2017).
Kobayashi et al. 2017, 17, 19, 20, Honshu (Shiga, Kyoto).
Nojima 2017, 120, Honshu (Okayama).
Tanaka 2018a, 21, Honshu (Akita).
Tanaka 2018b, 9, Honshu (Akita).
Tanida 2018, 574, 583, male, wings.
Morita 2019a, 22, Honshu (Mie).
Tanaka 2019a, 27, Honshu (Akita).
Tanaka 2019b, 12, Honshu (Akita).
Morita 2020b, 43, Honshu (Mie).
Zhang and Zhou 2021, 551-553, larva, China.
Malicky 2022, 35-36, male, synonyms..
Tanaka 2023, 42, Honshu (Akita).
Type locality. China (Beijing).
Distribution. Japan (Honshu, Kyushu), Russia, Korea, China.
Subfamily Macronematinae Ulmer
Only the genus Macrostemum is known from Japan. Species of the genus Macronema recorded from Nearctic, Oriental, Ethiopian and Australasian Regions were
transferred to the genus Macrostemum by Flint and Bueno-Soria (1982).
Genus Macrostemum Kolenati
Three species have been recorded from Japan.
Macrostemum formosicolum (Matsumura 1931)
Macronema formosicolum Matsumura 1931, 1132-1133, male, Taiwan.
Macrostemum formosicolum: Malicky 1998, 771, pls. 1, 6, male, Taiwan.
Ito and Nozaki 2018, 560-562, male, variations of wing marakings, Taiwan, Ishigaki-jima.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima), Taiwan.
Remarks. A female collected from Iriomote-jima probably the same species as this
species (Ito and Nozaki 2018).
Macrostemum okinawanum (Matsumura 1931)
Macronema okinawanum Matsumura 1931, 1133, adult, Okinawa.
Fischer 1963, 192.
Macrostemum okinawanum: Tanida 1997a, 448.
Kuranishi and Kimura 2001, 17-18, adult, Amami-oshima, Tokuno-shima, Okinawa-jima.
Nozaki 2016, 77, 352, 435, 450, adult, Amami-oshima.
Tanida 2016, 94.
Ito and Nozaki 2018, 562-563, male, variations of wing markings, Amami-o-shima, Okinawa-jima.
Tanida et al. 2018, 30, Okinawa-jima.
Type locality. Japan (Okinawa).
Distribution. Japan (Amami-oshima, Tokuno-shima, Okinawa-jima).
Remarks. Ito and Nozaki (2018) pointed out that populations of Okinawa-jima and Amami-o-shima may be a subspecies of M. formosicolum.
Macrostemum radiatum (McLachlan 1872)
Macronema radiatum McLachlan 1872, 67, pl. 2, adult, eastern Siberia, Amur.
Matsumura 1904, 169-170, adult.
Banks 1906, 111, Honshu (Gifu).
Matsumura 1907, 194, adult.
Horasawa 1930, 199-200, pupa, pupal case, Honshu (Tokyo).
Matsumura 1931, 1133-1134, adult, Honshu (Tokyo).
Kuwayama 1932, 1511, adult.
Hirayama 1933, pl. 8, adult, Honshu (Tokyo).
Matsumura 1933, 3, pl. 1, adult.
Tateishi 1934, 77, Kyushu (Fukuoka).
Esaki et al. 1938, 134-135, pl. 61, adult.
Tsuda 1940, 23-24, 32, Honshu (Akita, Nagano, Shiga, Kyoto).
Tsuda 1942a, 277, Honshu (shiga, Kyoto).
Tsuda 1950, 416, adult.
Tsuda 1951, 54, larva.
Akagi 1955, 17, larva.
Chihara 1955, 21-22, pupa.
Takeuchi 1955, 74, pl. 28, adult, Honshu (Kyoto).
Tsuda and Kawai 1956, 73-75, Honshu (Kyoto).
Gyotoku 1957, 10, Kyushu (Fukuoka).
Kimmins 1957, 106, lectotype designation, Siberia.
Miwa and Okawa 1958, 18, Honshu (Mie).
Tsuda 1959, 139, larva.
Tsuda and Akagi 1962, 132, larva.
Fischer 1963, 195-196.
Kuwada 1965, 164, pl. 82, adult, Shikoku (Kochi).
Tani 1977, 197, pl. 43, adult, Honshu (Shiga).
Tanida 1985, 176-177, larva.
Adachi et al. 1986, 86-89, 91, 93, larval mandibles
Kobayashi 1987, 17, Honshu (Shimane).
Kadowaki 1998, 3-4, Honshu (Shimane).
Kameyama et al. 2004, 213, Honshu (Hiroshima).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Nakamura and Hajiri 2007, 198, Honshu (Shimane).
Nakamura and Tsujimura 2007, 78, Honshu (Hiroshima).
Kameyama et al. 2009, 112, Honshu (Shimane, Hiroshima).
Nakamura and Kita 2010, 88, Honshu (Hiroshima).
Macronema lacustris Iwata 1927a, 223-224, pls. 5, 7, larva. Synonymized by Horasawa (1930).
Iwata 1927b, 206, pls. 1, 3, larva.
Fischer 1963, 189.
Sekimoto 1998, 360, Honshu (Hiroshima).
Yodoe et al. 1998, 68, Honshu (Shimane).
Macrostemum radiatum:
Ohtsuka & Tanida 1986, 5, Kyushu (Kumamoto).
Yagi et al. 1989, peculiar organ of female, 179, 181, Honshu (Kyoto).
Gyotoku and Nozaki 1991, 74, pl. 9, adult, Kyushu (Fukuoka).
Hatta and Nozaki 1991, 200, 208, larva.
Tanida et al. 1991, 48, adult, pupa, larva.
Nishimoto and Nishimoto 1993, 86, 92, adult, Honshu (Aichi).
Morita 1997, 41, Honshu (Mie).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Morita 1998, 72, Honshu (Mie).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338, Honshu (Mie).
Nishimoto and Morita 2001, 72, 76, adult, Honshu (Aichi).
Nakamura et al. 2002, 67, Honshu (Hiroshima).
Nozaki and Nakamura 2002, 171, Honshu (Hiroshima).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Tanida 2003, 381, larva.
Katsuma 2005a, 5, Honshu (Ibaraki).
Nozaki 2005, 108, adult larva.
Nozaki and Nakamura 2005, 222-223, Honshu (Shimane).
Okazaki 2005, 35-39, early instar larva, life history, Honshu (Nara).
Tanida 2005, 482, 487, adult, larva.
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 95, Honshu (Hiroshima).
Morita 2008a, 17, Honshu (Mie).
Tanida 2008, 250, pl. 92, adult, Shikoku (Kochi).
Tanaka 2009a, 45, adult, Honshu (Akita).
Tanaka 2009b, 49, adult, Honshu (Akita).
Kawase and Morita 2010, 42, Honshu (Shiga).
Mano 2010, 21-23, Honshu (Aichi).
Tanaka 2010a, 24, Honshu (Akita).
Inazu and Nishida 2011, 191, male, larva, Honshu (Hyogo).
Katsuma 2011a, 37, Honshu (Ibaraki).
Katsuma 2011d, 69, Honshu (Ibaraki).
Kawase and Hayashi 2011b, 192, Honshu (Shimane).
Morita 2011, 216, Honshu (Mie).
Morita and Katsuma 2011, Honshu (Mie).
Kawase and Hayashi 2012, 257, Honshu (Shimane).
Kawase and Minami 2013, 11, Honshu (Shiga).
Takai et al. 2013, 84, adult.
Nishimoto 2014, 58, larva.
Ohira and Tsutsumi 2014, 106-109, larva, life history, Honshu (Fukushima).
Ohira et al. 2015, 193, larva, mitochondrial DNA, Honshu (Fukushima).
Nozaki 2016, 77, 346, 352, 435, 450, adult, Shikoku (Kochi).
Tanaka 2016a, 23, 25, male, Honshu (Akita).
Tanida 2016, 94.
Kawase et al. 2017, Honshu (Shiga).
Kobayashi et al. 2017, 17, 19, 20, Hosnhu (Shiga, Kyoto).
Nojima 2017, 119, Honshu (Okayama).
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Tanida 2018, 572, adult, larva.
Morita 2019a, 22, Honshu (Mie).
Kitahara 2019, 122-124, larva, Honshu (Shizuoka: larval record).
Morita 2020c, 45, Honshu (Mie).
Morita 2020d, 81, Honshu (Mie).
Inadzu and Watanabe 2021, 53-58, retreat making behavior, Honshu (Hyogo).
Nishida, A. et al. 2022, 15-28, environmental factor, Honshu (Hyogo).
Tanaka 2023, 42, Honshu (Akita).
Type locality. Russia (Siberia).
Distribution. Japan (Honshu, Shikoku, Kyushu), Russia, Korea, China.
References
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