Philopotamidae

A catalogue of Japanese Trichoptera Family Philopotamidae Stephens
Author: Naotoshi KUHARA
E-mail: naotoshi.kuhara[AT]nifty.com
Last updated: 6 Sept. 2024

Subfamily Chimarrinae Rambur
In Japan, only genus Chimarra has been recorded in this subfamily.

Genus Chimarra Stephens

In Japan only one named species was recorded, but an unnamed species accompanied with an alphabetic code was described for larva, which was not associated with adult. Tanida (2003, 2005) illustrated unidentified larval specimens of this genus. Kuhara (2017) pointed out that the larva tentalively described by Furuya (1974) as Chimarra sp. CB belongs the genus Wormaldia.

Chimarra (Chimarra) tsudai Ross 1956
Chimarra concolor Ulmer: Tsuda 1939b, 295, Honshu (Nagano, Kyoto). Misidentification mentioned by Tanida (2005).

Tsuda 1941, 104-105, Honshu (Nagano).
Tsuda 1942a, 255, Honshu (Kyoto).
Tsuda 1943, 163-164, Honshu (Kyoto).

Chimarra sp. CA: Kumode 1952, 12-13, larva, Honshu (Nara).

Furuya 1974, 8-9, larva.

Chimarra tsudai Ross 1956, 51, 70-72, male, Honshu (Nagano).

Fischer 1971, 219.
Kobayashi 1971, 23-24, pl. 13, male, Honshu (Kanagawa).
Tani 1976, 391, Tsushima.
Kobayashi 1980, 88, Pl. 1-2, male, Honshu (Yamagata, Saitama, Tokyo, Kanagawa).
Kobayashi 1985a, 10, Tsushima.
Kobayashi 1987, 15, Oki-shoto.
Nozaki 1987, 39, Honshu (Kanagawa).
Hatta and Nozaki 1991, 199, Honshu (Gifu).
Tanida 1991, 33, Honshu (Nara).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Nishimoto and Nishimoto 1993, 85, Honshu (Aichi).
Moriya 1994, 68, Honshu (Kanagawa).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Moriya 1997, 60, Honshu (Kanagawa).
Nishiharu 1997, 37-38, Honshu (Fukui).
Nozaki 1997a, 33, Honshu (Kanagawa).
Nishimoto and Nishimoto 1998, 149, Honshu (Aichi).
Kagaya et al. 1998, 43, 124, Honshu (Tokyo).
Morita 1999b, 82, Honshu (Mie).
Kuhara 2001, 14, Hokkaido (Shiribeshi).
Morita 2001a, 175, Honshu (Mie).
Morita 2001b, 219, Honshu (Aichi).
Morita 2001c, 337, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Kawase et al. 2004, 108-109, Honshu (Ibaraki).
Nozaki 2004, 1229, Honshu (Kanagawa).
Ishiwata et al. 2005, 242, Honshu (Kanagawa).
Katsuma 2005a, 3, Honshu (Ibaraki).
Katsuma 2005b, 12, Honshu (Gumma).
Tanida 2005, 459, 464, male.
Mey and Nozaki 2006, 24, Honshu (Gifu).
Torii and Hattori 2006, 34-35, Honshu (Shizuoka).
Katsuma 2006, 41, Honshu (Ibaraki).
Nakamura and Tsujimura 2007, 76-77, Honshu (Hiroshima).
Nozaki and Tanida 2007, 247, Honshu (Shizuoka).
Inazu 2008, 69, 89, 100, Honshu (Hyogo).
Katuma 2008, 40, Honshu (Ibaraki).
Morita 2008a, 722, Honshu (Mie).
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009b, 138, Honshu (Mie).
Ito et al. 2010, 58, 74, Hokkaido (Oshima).
Kawase and Morita 2010, 38, Honshu (Shiga).
Kawase et al. 2010, 108 Honshu (Shimane).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka).
Nozaki 2010, 216, Honshu (Kanagawa).
Inazu and Nishida 2011, 182, male, larva, Honshu (Hyogo).
Katsuma 2011b, 49-52, Honshu (Ibaraki).
Morita 2011, 215, Honshu (Mie).
Katsuma 2012, 64, Honshu (Ibaraki).
Morita 2012, 249, Honshu (Mie).
Moriya and Kubota, 2012, 131, Honshu (Kanagawa).
Inazu 2013b, 56, Honshu (Hyogo).
Inazu 2013c, 70, Honshu (Hyogo).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 9, Shikoku (Ehime).
Nishimoto and Nishimoto 2014, 63, Honshu (Aichi).
Katsuma 2015, 91, Honshu (Fukushima).
Horiguchi 2016, 410, Honshu (Tokyo).
Nozaki 2016, 72, 329, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 79.
Yamamoto and Ito 2016, 38, Shikoku (Ehime).
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Imasaka et al. 2017, 61, Kyushu (Fukuoka).
Kuhara 2017, 52, larva, Honshu (Ibaraki, Chiba, Shizuoka, Shimane) (larval record).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 114, Honshu (Okayama).
Kuhara 2018, 529-530, male, female, larva.
Tsuruda 2018, 26, 30, Honshu (Tokyo).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Taira 2020, 8, Honshu (Nara).
Shimura and Imasaka 2021, 3, Koshikijima-retto.
Higuchi 2024, 21, Honshu (Ishikawa).

Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki-shoto, Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Korea.
Remarks. Kuhara (2017) considered that the larva tentatively described as Chimarra sp. CA by Kumode (1952) and Furuya (1974) is C. tsudai, but that by Tanida (1985) is a different species.

Subfamily Philopotaminae Stephens
Three genera, Dolophilodes, Kisaura and Wormaldia have been recorded from Japan.

Genus Dolophilodes Ulmer
Nine named species were recorded in Japan (Kuhara 2005a). Several illustrations of male genitalia (Yamanouchi 1987, Morita 1996, 1998b) were also provided for unidentified species of "Dolophilodes", but their generic placement should be reexamined because Kisaura was elevated from a subgenus of Dolophilodes (s.l.) to generic status (Malicky 1993, see also Blahnik 2005). Kuhara (2017) described larvae of 8 out of 9 species recorded in Japan, but failed to clarify the identities D. sp. DA tentatively described based on larval specimens by Kumode (1952) and Furuya (1974) and that of D. sp. DB by Furuya (1974). Kuhara (2017), however, associated each larvae illustrated and tentatively named as D. sp. DA, DB and DD by Tanida (1985, 2005) to named species and transferred Dolophilodes sp. DC by him to the genus Wormaldia.

Dolophilodes angustata Kuhara 2005

Dolophilodes sp. DA: Tanida 1985, 172-173, larva. Reidentified by Kuhara (2017).

Tanida 2005, 461-462, larva.

Dolophilodes angustata Kuhara 2005a, 95-97, male, female, Honshu (Saitama, Tokyo, Kanagawa, Niigata, Yamanashi, Nagano, Gifu, Shizuoka, Aichi, Mie, Osaka, Hyogo, Nara, Wakayama, Tottori, Shimane, Okayama, Hiroshima), Shikoku (Kagawa, Ehime, Kochi), Kyushu (Kumamoto, Oita, Miyazaki) (angustatus).

Torii and Hattori 2006, 35, Honshu (Shizuoka) (angustatus).
Morita 2008b, 49, Honshu (Mie)
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Kawase and Morita 2010, 38, Honshu (Mie, Shiga).
Morita 2011, 215, Honshu (Mie).
Moriya and Kubota, 2012, 131, Honshu (Kanagawa).
Morita 2013, 61, Honshu (Mie).
Nishimoto and Nishimoto 2014, 63, Honshu (Aichi).
Morita and Kawase 2016, 22, Honshu (Ishikawa).
Nozaki 2016, 72, 329-330, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 79.
Kuhara 2017, 55-57, larva, Honshu (Niigata, Shizuoka, Nara, Tottori), Shikoku (Tokushima, Ehime) (larval record).
Nojima 2017, 114, Honshu (Okayama).
Nozaki 2018, 928, Honshu (Kanagawa).
Kuhara 2018, 531-535, male, female, larva.
Yamamoto et al. 2021, 71, Shikoku (Ehime).

Type locality. Honshu (Nara), Japan.
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).

Dolophilodes auriculata Martynov 1933
Dolophilodes auriculata Martynov 1933, 142-143, male, Honshu (Shiga) (auriculatus).

Tsuda 1939b, 296, Honshu (Nagano, Kyoto) (auriculatus).
Tsuda 1941, 104-105, Honshu (Nagano) (auriculatus).
Tsuda 1942a, 255, Honshu (Shiga) (auriculatus).
Tsuda 1943, 164, Honshu (Kyoto) (auriculatus).
Fischer 1961, 3.
Tanida and Takemon 1993, 242, Honshu (Kyoto) (auriculatus).
Tanida et al. 1995, 2, Honshu (Nara) (auriculatus).
Morita 1996, 303, Honshu (Mie) (auriculatus).
Ito et al. 1997, 23, 25, 27, 29, Hokkaido (Oshima) (auriculatus).
Kuhara 1997a, 58, Hokkaido (Shiribeshi) (auriculatus).
Nozaki 1997a, 33, Honshu (Kanagawa).
Ito et al. 1998, 14, Hokkaido (Nemuro) (auriculatus).
Kagaya et al. 1998, 43, 126, Honshu (Tokyo, Yamanashi) (auriculatus).
Ito et al. 2000, 20, Hokkaido (Ishikari) (auriculatus).
Kuhara 2001, 14, Hokkaido (Shiribeshi) (auriculatus).
Morita 2001b, 219, Honshu (Aichi).
Uchida 2002, 77, Honshu (Tochigi) (auriculatus).
Morita 2003, 24, Honshu (Mie) (auriculatus).
Kawase et al. 2004, 109, Honshu (Ibaraki) (auriculatus).
Katsuma 2005a, 4, Honshu (Ibaraki) (auriculatus).
Kuhara 2005a, 98-100, male, Hokkaido (Nemuro, Abashiri, Tokachi, Rumoi, Sorachi, Ishikari, Shiribeshi, Hiyama, Oshima), Honshu (Iwate, Akita, Ibaraki, Saitama, Tokyo, Kanagawa, Yamanashi, Nagano, Gifu, Shizuoka, Mie, Kyoto, Osaka, Hyogo, Nara, Wakayama, Tottori, Shimane, Okayama, Hiroshima), Shikoku (Kagawa, Ehime, Kochi), Kyushu (Fukuoka, Saga, Nagasaki, Kumamoto, Oita), Tsushima (auriculatus).
Tanida 2005, 460, 465, male (auriculatus).
Katsuma 2006, 41, Honshu (Ibaraki) (auriculatus).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Morita 2006, 199, Honshu (Mie).
Morita 2008a, 722, Honshu (Mie) (auriculatus).
Morita 2008c, 92-93, Honshu (Mie).
Tanida 2008, 249, Honshu (Osaka) (auriculatus).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka) (auriculatus).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Katsuma 2010, 85, Honshu (Ibaraki).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 38, Honshu (Mie, Shiga).
Kawase et al. 2010, 108 Honshu (Shimane).
Nakamura 2010, 239 Honshu (Hiroshima) (auriculatus).
Inazu and Nishida 2011, 182, male, Honshu (Hyogo).
Ito and Kubo 2011, 6, Hokkaido (Tokachi).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari).
Katsuma 2011b, 49-52, Honshu (Ibaraki).
Katsuma 2012, 64, Honshu (Ibaraki).
Inazu 2013c, 70, Honshu (Hyogo).
Morita 2013, 61, Honshu (Mie).
Ito and Nagasaka 2014, 9, Hokkaido (Sorachi).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Morita and Kawase 2016, 21, Honshu (Ishikawa).
Tanida 2016, 79.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 53, larva, Hokkaido (Sorachi, Ishikari, Hidaka), Honshu (Akita, Ibaraki, Shizuoka), Kyushu (Kumamoto, Kagoshima) (larval record).
Nojima 2017, 114, Honshu (Okayama).
Kuhara 2018, 531, 533-535, male, female, larva.
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yamamoto et al. 2021, 71, Shikoku (Ehime).
Ito and Kuhara 2023, 55, Hokkaido (Soya, Kamikawa).

Sortosa auriculata: Ross 1956, 37, 59, male.

Fischer 1971, 172.
Kobayashi 1980, 92, pl. 1, pl. 3, male, Honshu (Niigata) (misspelled as "Sortose").

Sortosa auriculata: Kobayashi 1987, 14, Honshu (Shimane).
Type locality. Honshu (Shiga), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima).

Dolophilodes babai (Kobayashi 1980)
Sortosa babai Kobayashi 1980, 94, pl. 1, Pl. 4, male, Honshu (Niigata).

Kobayashi 1985b, 68, Honshu (Kanagawa).

Dolophilodes babai: Kuhara 2005a, 101-102, male, Honshu (Niigata).

Tanida 2005, 460.
Tanida 2016, 79.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (central).

Dolophilodes commata (Kobayashi 1980)
Sortosa commata Kobayashi 1980, 95-96, Pl. 1, Pl. 5, male, Honshu (Hiroshima).

Kobayashi 1987, 14-15, Oki-shoto.
Nishimoto and Nishimoto 1993, 85, Honshu (Aichi).

Sortosa kaishoensis Kobayashi 1985a, 8-9, male, Tsushima. Synonymized by Kuhara (2005a).
Dolophilodes commata: Kagaya et al. 1998, 43, 127, Honshu (Tokyo) (commatus).

Nakano and Tanida 1999, 276, Honshu (Nara) (commatus).
Morita 2001a, 175, Honshu (Mie) (commatus).
Katsuma 2005a, 4, Honshu (Ibaraki) (commatus).
Katsuma 2005b, 12, Honshu (Tochigi, Gumma) (commatus).
Kuhara 2005a, 104-105, new synonym, male, female, Honshu (Miyagi, Tochigi, Tokyo, Aichi, Mie, Kyoto, Osaka, Hyogo, Nara, Wakayama), Shikoku (Ehime), Kyushu (Fukuoka, Miyazaki), Tsushima, Ryukyu (Yakushima) (commatus).
Tanida 2005, 460 (commatus).
Katsuma 2006, 41, Honshu (Ibaraki) (commatus).
Inazu 2008, 69-70, 89, 100, Honshu (Hyogo).
Morita 2009b, 138, Honshu (Mie).
Inazu and Nishida 2011, 182, male, female, Honshu (Hyogo).
Morita 2011, 215, Honshu (Mie).
Inazu 2012, 6, Honshu (Hyogo).
Morita 2012, 249, Honshu (Mie).
Inazu 2013c, 70, Honshu (Hyogo).
Kawase and Morita 2014, 4, Honshu (Shiga).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Morita and Kawase 2016, 22, Honshu (Ishikawa).
Nozaki 2016, 72, 330, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 79.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Kuhara 2017, 53-56, larva, Honshu (Mie, Yamaguchi), Tsushima (larval record).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 114, Honshu (Okayama).
Nozaki 2018, 928, Honshu (Kanagawa).
Kuhara 2018, 531, 533-535, male, female, larva.
Yamamoto 2023, 32, Shikoku (Ehime).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Honshu (Hiroshima), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu, Tsushima, Oki-shoto, Ryukyu (Yakushima)).

Dolophilodes dilatata Kuhara 2005
Dolophilodes dilatata Kuhara 2005a, 97-98, male, female, Hokkaido (Ishikari, Iburi, Shiribeshi), Honshu (Aomori, Iwate, Yamagata, Fukushima, Saitama, Yamanashi, Nagano, Gifu, Shizuoka, Mie, Shiga, Hyogo, Tottori, Okayama) (dilatatus).

Katsuma 2005b, 13, Honshu (Tochigi) (dilatatus).
Morita 2008b, 49, Honshu (Mie).
Katsuma 2008, 37, 40, Honshu (Ibaraki).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi).
Inazu and Nishida 2011, 183, male, female, Honshu (Hyogo).
Kuhara, 2011, 51, 70, Hokkaido (Ishikari).
Katsuma 2012, 64, Honshu (Ibaraki).
Inazu 2012, 6, Honshu (Hyogo).
Inazu 2013c, 70, Honshu (Hyogo).
Tanida 2016, 79.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 55-56, larva, Hokkaido (Iburi), Honshu (Akita, Fukushima, Yamanashi, Nagano, Shizuoka) (larval record).
Nojima 2017, 114, Honshu (Okayama).
Kuhara 2018, 531-534, male, female, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Hokkaido, Honshu).

Dolophilodes iroensis (Kobayashi 1980)
Sortosa iroensis Kobayashi 1980, 93-94, pl. 1, Pl. 4, male, Honshu (Tokyo, Kanagawa).
Dolophilodes sp. DD: Tanida 1985, 172-173, larva. Reidentified by Kuhara (2017).

Tanida 2005, 461-462, larva.

Dolophilodes kunashirensis Ivanov 1996 (in Arefina et al. 1996), 4-5, male. Synonimized by Kuhara (2005a).

Ito et al. 1998, 14, Hokkaido (Nemuro).
Ito et al. 2000, 20, Hokkaido (Ishikari).
Kuhara 2001, 14, Hokkaido (Shiribeshi).
Morita 2003, 24, Honshu (Mie).
Kawase et al. 2004, 109, Honshu (Ibaraki).

Dolophilodes iroensis: Kuhara 2005a, 102-104, new synonym, male, female, Hokkaido (Nemuro, Abashiri, Tokachi, Kamikawa, Ishikari, Hidaka, Iburi, Shiribeshi, Hiyama, Oshima), Honshu (Aomori, Iwate, Ibaraki, Saitama, Tokyo, Yamanashi, Nagano, Gifu, Shizuoka, Mie, Shiga, Nara, Tottori, Okayama), Shikoku (Ehime, Kochi), Kyushu (Fukuoka, Nagasaki, Kumamoto, Oita), Yakushima.

Tanida 2005, 460.
Katsuma 2005b, 12, Honshu (Tochigi).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Katsuma 2006, 41, Honshu (Ibaraki).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Morita 2008b, 49, Honshu (Mie).
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Ito et al. 2010, 58, 74, Hokkaido (Hiyama, Oshima).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 38, Honshu (Mie, Shiga).
Kawase et al. 2010, 108 Honshu (Shimane).
Inazu and Nishida 2011, 183, male, Honshu (Hyogo).
Katsuma 2011a, 67, Honshu (Ibaraki).
Katsuma 2011b, 49-52, Honshu (Ibaraki).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari, Hidaka, Iburi).
Inazu 2012, 6, Honshu (Hyogo).
Ito 2013, 53, Hokkaido (Tokachi).
Inazu 2013c, 70, Honshu (Hyogo).
Morita 2013, 61, Honshu (Mie).
Kawase and Morita 2014, 4, Honshu (Shiga).
Ijima and Tsuchiya 2014, 28, Hokkaido (Ohotsuku).
Ito and Nagasaka 2014, 9, Hokkaido (Sorachi).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Yamamoto and Ito 2016, 38, Shikoku (Ehime).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Tanida 2016, 80.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Ito 2017b, 41, Hokkaido (Kamikawa).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 55-57, larva, Hokkaido (Abashiri, Oshima), Honshu (Shizuoka, Kyoto), Kyushu (Fukuoka) (larval record).
Nojima 2017, 114, Honshu (Okayama).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Kuhara 2018, 531, 533-535, male, female, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Shimura and Imasaka 2020, 12, 16, Kyushu (Fukuoka, Oita).
Yamamoto et al. 2021, 71, Shikoku (Ehime).
Aoya and Tanaka, 2023, 26, Honshu (Akita).
Ito and Kuhara 2023, 55, Hokkaido (Soya, Ohotsuku, Kamikawa).
Tanaka 2023a, 42, Honshu (Akita).
Yamamoto 2023, 32, Shikoku (Ehime).
Tanaka 2024, 27, Honshu (Akita).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Ryukyu (Yakushima)), Russia (southern and northern Kurils).

Dolophilodes japonica (Banks 1906)
Philopotamus japonica Banks 1906, 111, 113, male, Kyushu (Fukuoka) (japonicus).

Ulmer 1908, 354, Kyushu (Fukuoka)

Dolophilodes exscisa Martynov 1933, 140-142, male, Honshu (Tochigi) (exscisus). Synonimized by Ross (1956).

Tsuda 1939a, 150, Honshu (Tochigi) (exscisus).
Tsuda 1939b, 295-296, Honshu (Tochigi, Nagano, Kyoto) (exscisus).
Tsuda 1941, 104-105, Honshu (Nagano) (exscisus).
Tsuda 1942a, 255, Honshu (Nagano, Shiga, Kyoto) (exscisus).
Tsuda 1943, 163-164, Honshu (Kyoto) (exscisus).
Fischer 1961, 3.

Dolophilodes japonca: Tsuda 1939b, 297 (japonicus).

Fischer 1961, 4.
Kobayashi 1971, 21-22, pl. 12, male, female, Honshu (Kanagawa).
Tani 1977, 193, thorax of male.
Tani 1978, 4, Honshu (Niigata).
Tanida 1982, 18-19 Honshu (Ishikawa) (japonicus).
Nozaki 1988a, 46, Honshu (Kanagawa) (japonicus).
Nozaki 1988b, 38, Honshu (Kanagawa) (japonicus).
Morita 1989, 38, Honshu (Mie) (japonicus).
Hatta and Nozaki 1991, 199, Honshu (Aichi) (japonicus).
Kugo and Nozaki 1991, 7, Honshu (Hyogo) (japonicus).
Tanida 1991, 33, Honshu (Nara) (japonicus).
Gyotoku and Nozaki 1991, 73, Kyushu (Fukuoka) (japonicus).
Tanida and Takemon 1993, 242, Honshu (Kyoto) (japonicus).
Morita 1993, 3, Honshu (Mie) (japonicus).
Moriya 1994, 68, Honshu (Kanagawa) (japonicus).
Tanida et al. 1995, 2, Honshu (Nara) (japonicus).
Ito et al. 1997, 23, 25, 27, 29, Hokkaido (Oshima).
Kuhara 1997a, 58, Hokkaido (Shiribeshi) (japonicus).
Moriya 1997, 60, Honshu (Kanagawa, Yamanashi) (japonicus).
Nozaki 1997a, 33-34, Honshu (Kanagawa) (japonicus).
Nozaki 1997b, 28 Honshu (Tokyo) (japonicus).
Ito et al. 1998, 14, Hokkaido (Nemuro) (japonicus).
Kagaya et al. 1998, 44, 128, Honshu (Tokyo, Yamanashi) (japonicus).
Morita 1998b, 46-47, male, Honshu (Mie) (japonicus).
Ito et al. 1999, 27, Hokkaido (Kushiro) (japonicus).
Ito et al. 2000, 20, Hokkaido (Ishikari) (japonicus).
Kuhara 2001, 14, Hokkaido (Shiribeshi) (japonicus).
Morita 2001a, 175, Honshu (Mie) (japonicus).
Morita 2001c, 337, Honshu (Mie) (japonicus).
Kuranishi and Satake 2003, 34, Hokkaido (Abashiri) (japonicus).
Morita 2003, 24, Honshu (Mie) (japonicus).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Nozaki 2004, 1229, Honshu (Kanagawa) (japonicus).
Ishiwata et al. 2005, 243, Honshu (Kanagawa) (japonicus).
Katsuma 2005a, 4, Honshu (Ibaraki) (japonicus).
Katsuma 2005b, 12, Honshu (Tochigi) (japonicus).
Kuhara 2005a, 92-94, male, female, Hokkaido (Nemuro, Kushiro, Tokachi, Ishikari, Hidaka, Iburi, Shiribeshi), Honshu (Aomori, Miyagi, Akita, Yamagata, Tochigi, Saitama, Tokyo, Kanagawa, Niigata, Toyama, Ishikawa, Fukui, Yamanashi, Nagano, Gifu, Shizuoka, Mie, Shiga, Hyogo, Kyoto, Nara, Wakayama, Tottori, Shimane, Okayama, Hiroshima), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka, Saga, Kumamoto, Oita, Miyazaki, Kagoshima), Sado-ga-shima (japonicus).
Tanida 2005, 460, 465, male (japonicus).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Torii and Hattori 2006, 35, Honshu (Shizuoka) (japonicus).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Nozaki and Nakamura 2007, 93, Honshu (Tottori) (japonicus).
Inazu 2008, 70-71, 89, 100, Honshu (Hyogo).
Morita 2008a, 722, Honshu (Mie) (japonicus).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka) (japonicus).
Morita 2009b, 138, Honshu (Mie).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Morita 2010, 38, Honshu (Shiga).
Kobayashi 2010, 140, Honshu (Tochigi).
Tanaka 2010, 24, Honshu (Akita) (japonicus).
Inazu and Nishida 2011, 183, male, female, Honshu (Hyogo).
Ito and Kubo 2011, 6, Hokkaido (Tokachi).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari, Hidaka, Iburi).
Katsuma 2012, 64, Honshu (Ibaraki).
Inazu 2012, 6, Honshu (Hyogo).
Morita 2012, 249, Honshu (Mie).
Ito 2013, 53, Hokkaido (Kushiro).
Inazu 2013c, 70, Honshu (Hyogo).
Ijima and Tsuchiya 2014, 28, Hokkaido (Ohotsuku).
Ito and Nagasaka 2014, 9, Hokkaido (Sorachi).
Horiguchi 2016, 410, Honshu (Tokyo).
Nozaki 2016, 72, 330, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 80.
Ito 2017a, 4, Honshu (Fukui).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 55-56, larva, Hokkaido (Ishikari), Honshu (Akita, Yamagata, Gumma, Kyoto, Wakayama, Okayama, Hiroshima) (larval record).
Nojima 2017, 114-115, Honshu (Okayama).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Kuhara 2018, 531-534, male, female, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yamamoto et al. 2021, 71, Shikoku (Ehime).
Tanaka 2022, 35, Honshu (Akita).
Tanaka 2023a, 42, Honshu (Akita).
Ito and Kuhara 2023, 56, Hokkaido (Soya, Kamikawa).
Yamamoto 2023, 32, Shikoku (Ehime).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).
Yamamoto 2024, 36, Shikoku (Ehime).

Sortosa japonica: Ross 1956, 59.

Fischer 1971, 174-175.
Kobayashi 1980, 92-93, pl. 1, pl. 3, male, Hokkaido (Tokachi), Honshu (Tokyo, Kanagawa, Niigata) (misspelled as "Sortose"). Kobayashi 1982, 96, Honshu (Kanagawa).
Kadowaki 1998, 2, Honshu (Shimane).

Dolophilodes sp. DB: Tanida 1985, 172-173, larva. Reidentified by Kuhara (2017).

Tanida 2005, 461-462, larva.

Dolophilodes nomugiensis: Tanaka 2008, 39 Honshu (Akita). Misidentification mentioned by Tanaka (2019).
Type locality. Kyushu (Fukuoka), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Sado-ga-shima), Russia (Sakhalin, southern Kurils).

Dolophilodes nomugiensis (Kobayashi 1980)
Sortosa nomugiensis: Kobayashi 1980, 94-95, pl. 1, Pl. 4, male, Honshu (Kanagawa, Gifu).

Kobayashi 1987, 14, Oki-shoto.

Wormaldia triangulata: Kobayashi 1984a, 1-2, male, Honshu (Niigata). Synonymized by Kuhara (2005a).
Wormaldia sp. WA: Tanida 1985, 172-173, larva. Reidentified by Kuhara (2017).
Dolophilodes nomugiensis: Kuhara 1997b, 94, Hokkaido (Shiribeshi).

Ito et al. 1998, 14, 16-17, Hokkaido (Nemuro, Abashiri).
Kagaya et al. 1998, 44, 130, Honshu (Yamanashi).
Ohkawa 1999, 40-43, Hokkaido (Tokachi).
Ito et al. 2000, 20, Hokkaido (Ishikari).
Kuhara and Sato 2001, 63, Rishiri-to.
Kawase et al. 2004, 109, Honshu (Ibaraki).
Katsuma 2005a, 4, Honshu (Ibaraki).
Kuhara 2005a, 100-101, new synonym, male, female, Hokkaido (Nemuro, Abashiri, Kushiro, Tokachi, Kamikawa, Sorachi, Ishikari, Hidaka, Iburi, Shiribeshi, Hiyama, Oshima), Honshu (Aomori, Iwate, Fukushima, Ibaraki, Saitama, Niigata, Ishikawa, Yamanashi, Nagano, Shizuoka, Nara, Tottori, Shimane, Okayama, Hiroshima), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka, Nagasaki, Saga, Oita), Rishiri-to.
Tanida 2005, 460, 465, male.
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Katsuma 2006, 41, Honshu (Ibaraki).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Nakatani et al. 2007, 24, Hokkaido (Abashiri).
Inazu 2008, 71, 89, 100, Honshu (Hyogo).
Morita 2008a, 722, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Ito et al. 2009, Hokkaido (Abashiri).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Morita 2010, 38, Honshu (Mie).
Inazu and Nishida 2011, 183, male, female, Honshu (Hyogo).
Katsuma 2011a, 67, Honshu (Ibaraki).
Katsuma 2011b, 49-52, Honshu (Ibaraki).
Ito and Kubo 2011, 6, Hokkaido (Tokachi).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari, Hidaka, Iburi).
Inazu 2012, 6, Honshu (Hyogo).
Ito 2013, 53, Hokkaido (Tokachi).
Inazu 2013c, 70-71, Honshu (Hyogo).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Ito and Nagasaka 2014, 10, Hokkaido (Sorachi).
Nakatani et al. 2015, 59, Hokkaido (Kushiro).
Ito and Kuhara 2016, 16, Hokkaido (Kushiro).
Nozaki 2016, 72, 330, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 80.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 53, 55-56, larva, Hokkaido (Tokachi, Sorachi, Ishikari, Iburi, Oshima), Honshu (Kanagawa), Rebun-to (larval record).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 531, 533-535, male, female, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Nakatani 2019, 49, Hokkaido (Kushiro).
Tanaka 2022, 35, Honshu (Akita).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Tanaka 2023a, 42, Honshu (Akita).
Ito and Kuhara 2023, 56, Hokkaido (Soya, Ohotsuku, Kamikawa, Rumoi), Rebun-to.
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Honshu (Gifu), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rishiri-to, Rebun-to, Oki-shoto), Russia (Kamchatka, Sakhalin, Kurils).

Dolophilodes shinboensis (Kobayashi 1980)
Sortosa shinboensis Kobayashi 1980, 93, pl. 1, pl. 3, male, Honshu (Niigata).
Dolophilodes shinboensis: Morita 1994, 119, Honshu (Mie).

Tanida et al. 1995, 2, Honshu (Nara).
Morita 1996, 303, Honshu (Mie).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Nozaki 1997a, 34, Honshu (Kanagawa).
Kuhara 1997a, 58, Hokkaido (Shiribeshi).
Kagaya et al. 1998, 44, 131, Honshu (Tokyo, Yamanashi).
Morita 1998a, 13, Honshu (Mie).
Ito et al. 2000, 20, Hokkaido (Ishikari).
Kuhara and Sato 2001, 63-64, Rishiri-to.
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Tanida 2005, 460.
Kuhara 2005a, 93-95, male, female, Hokkaido (Abashiri, Ishikari, Shiribeshi, Hiyama), Honshu (Aomori, Iwate, Akita, Saitama, Tokyo, Niigata, Ishikawa, Nagano, Gifu, Kyoto, Tottori), Shikoku (Tokushima, Ehime, Kochi), Fukuoka (Fukuoka, Saga, Kumamoto, Oita), Rishiri-to.
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Nozaki and Nakamura 2007, 93, Honshu (Hiroshima).
Katsuma 2008, 37, 40, Honshu (Ibaraki).
Ito et al. 2010, 58, 74, Hokkaido (Oshima).
Kawase and Morita 2010, 38, Honshu (Shiga).
Kuhara, 2011, 51, 59, 70, Hokkaido (Ishikari).
Kawase and Morita 2014, 4, Honshu (Shiga).
Nozaki et al. 2014, 60, Honshu (Mie).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Nozaki 2016, 72, 330-331, 432, 450, male, Shikoku (Kochi).
Yamamoto and Ito 2015, 33, Shikoku (Ehime).
Tanida 2016, 80.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2017, 54-56, larva, Hokkaido (Sorachi, Ishikari, Iburi), Honshu (Fukui, Nagano) (larval record).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 531-534, male, female, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Ito and Kuhara 2023, 56, Hokkaido (Soya).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rishiri-to).

Genus Kisaura Ross
Eleven species of this genus have been recorded in Japan. Yamanouchi (1987) illustrated the male genitalia and wing of an unidentified species as "Dolophilodes sp. 2", which was treated as "Dolophilodes (Kisaura)" by Ohkawa (1999). Morita (1997) illustrated the male genitalia of an unidentified species of this genus. Inazu and Nishida (2014) also showed pictures of the male genitalia of two unidentified species.

Kisaura ashizuriensis Kuhara 2019
Kisaura ashizuriensis Kuhara 2019, 231-233, Shikoku (Kochi), male.
Type locality. Shikoku (Kochi), Japan.
Distribution. Japan (Shikoku).

Kisaura borealis (Kuhara 1999)
Dolophilodes sp.: Kuhara 1997a, 58, Hokkaido (Shiribeshi) (sp. 2). Mentioned by Kuhara (2001).
Dolophilodes borealis Kuhara 1999, 181-182, male, Hokkaido (Nemuro, Sorachi, Ishikari, Shiribeshi, Hiyama, Oshima), Honshu (Yamagata, Niigata).

Ito et al. 2000, 20-21, Hokkaido (Ishikari).
Kuhara 2001, 14, Hokkaido (Shiribeshi).
Tanida 2005, 459, 464, male.

Kisaura borealis: Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).

Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Oshima).
Ito and Kubo 2011, 6, Hokkaido (Tokachi).
Kuhara 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari).
Nozaki et al. 2014, 60, Honshu (Nagano).
Tanida 2016, 81.
Ito 2017b, 41, Hokkaido (Kamikawa).
Kuhara 2018, 536-537, male.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Tanaka 2022, 35, Honshu (Akita).
Suzuki et al. 2023, Hokkaido (Sorachi, Ishikari), Honshu (Iwate, Yamagata), molecular phylogeography.

Type locality. Hokkaido (Ishikari), Japan.
Distribution. Japan (Hokkaido, Honshu (north to central)), Russia (Sakhalin, southern Kurils).

Kisaura brevis Kuhara 2019
Kisaura brevis Kuhara 2019, 235-236, Honshu (Yamanashi, Shizuoka, Shiga), Shikoku (Ehime, Kochi), male.
Type locality. Shikoku (Ehime), Japan.
Distribution. Japan (Honshu, Shikoku).

Kisaura curvispina Kuhara 2019
Kisaura curvispina Kuhara 2019, 233-235, Shikoku (Ehime, Kochi), male.
Type locality. Shikoku (Ehime), Japan.
Distribution. Japan (Shikoku).

Kisaura dichotoma Kuhara and Arefina 2004
Dolophilodes sp.: Kuhara 1997a, 58, Hokkaido (Shiribeshi) (sp. 3). Mentioned by Kuhara and Arefina (2004).

Ito et al. 1997, 23, 27, Hokkaido (Oshima) (sp. 4).
Ito et al. 2000, 21, Hokkaido (Ishikari) (sp. 2).
Kuhara 2001, 14, Hokkaido (Shiribeshi).

Kisaura dichotoma Kuhara and Arefina 2004, 81-83, male, female, Hokkaido (Abashiri, Ishikari, Shiribeshi, Hiyama, Oshima), Honshu (Aomori, Iwate, Ibaraki, Saitama, Shizuoka, Niigata, Gifu, Mie), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Nagasaki).

Katsuma 2005a, 4, Honshu (Ibaraki).
Katsuma 2006, 41, Honshu (Ibaraki).
Morita 2006, 47, Honshu (Mie).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Katuma 2008, 40, Honshu (Ibaraki).
Morita 2009a, 6, Honshu (Shiga).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 38, Honshu (Mie, Shiga).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Ito and Nagasaka 2014, 10, Hokkaido (Sorachi).
Tanida 2016, 81.
Imasaka et al. 2017, 61, Kyushu (Fukuoka).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 536, 538, male, female.
Kuhara 2019, 239, Shikoku (Ehime, Kochi).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Ito and Kuhara 2023, 56, Rebun-to.
Higuchi 2024, 21, Honshu (Ishikawa).

Type locality. Hokkaido (Oshima), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rebun-to), Russia (southern Kurils).

Kisaura hattorii (Kuhara 1999)
Dolophilodes sp.: Kuhara 1997a, 58, Hokkaido (Shiribeshi) (sp. 1). Mentioned by Kuhara (2001).
Dolophilodes hattorii Kuhara 1999, 179-180, male, Hokkaido (Sorachi, Ishikari, Shiribeshi, Hiyama), Honshu (Iwate, Ibaraki, Tokyo, Kanagawa, Niigata, Yamanashi, Nagano, Shizuoka).

Ito et al. 2000, 21, Hokkaido (Ishikari).
Kuhara 2001, 14, Hokkaido (Shiribeshi).
Kawase et al. 2004, 109, Honshu (Ibaraki).
Tanida 2005, 459, 464, male.

Kisaura hattorii: Ito et al. 2000, 21, Hokkaido (Ishikari).

Katsuma 2005a, 4, Honshu (Ibaraki).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 69, Hokkaido (Ishikari).
Katsuma 2008, 40, Honshu (Ibaraki).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari, Hidaka).
Katsuma 2011b, 49-52, Honshu (Ibaraki).
Tanida 2016, 81.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara 2018, 536-537, male.
Tanida et al. 2018, 30, Honshu (Toyama).
Tanaka 2019, 26, 28, male, Honshu (Akita).
Tanaka 2022, 35, Honshu (Akita).
Aoya and Tanaka, 2023, 26, Honshu (Akita).
Ito and Kuhara 2023, 56, Hokkaido (Kamikawa).
Tanaka 2024, 27, Honshu (Akita).
Higuchi 2024, 21, Honshu (Toyama, Ishikawa).

Type locality. Hokkaido (Sorachi), Japan.
Distribution. Japan (Hokkaido, Honshu (north to central)), Russia (southern Kurils).

Kisaura kisoensis (Tsuda 1939)
Dolophilodes kisoensis Tsuda 1939b 296-297, male, Honshu (Nagano).

Tsuda 1941, 104-105, Honshu (Nagano).
Tsuda 1942a, 255, Honshu (Nagano, Kyoto).
Tsuda 1943, 163, 166, Honshu (Kyoto).
Tanida 1982, 18, Honshu (Ishikawa).
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Ohtsuka and Tanida 1986, 4, Kyushu (Kumamoto).
Morita 1988, 5, Honshu (Mie).
Nozaki 1988a, 46, Honshu (Kanagawa)
Nozaki 1988b, 38, Honshu (Kanagawa).
Morita 1989, 38, Honshu (Mie).
Kugo and Nozaki 1990, 61, Honshu (Hyogo).
Morita 1990, 32, Honshu (Mie) (misspelled as "Dolophiloder").
Hatta and Nozaki 1991, 199, Honshu (Gifu).
Kugo and Nozaki 1991, 7, Honshu (Hyogo).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Morita 1993, 3, Honshu (Mie).
Nishimoto and Nishimoto 1993, 85, Honshu (Aichi).
Moriya 1994, 68, Honshu (Kanagawa).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Kagaya et al. 1998, 44, 129, Honshu (Tokyo).
Kuhara 1999, 177-178, male, Hokkaido (Ishikari, Hiyama, Oshima), Honshu (Ibaraki, Tokyo, Kanagawa, Ishikawa, Yamanashi, Shizuoka, Shiga, Kyoto, Osaka, Nara, Wakayama, Shimane, Hiroshima), Kyushu (Fukuoka, Kumamoto, Oita).
Morita 2001c, 337, Honshu (Mie).
Nozaki and Nakamura 2002, 168, Honshu (Hiroshima).
Nagayasu et al. 2003, 106, Hokkaido (Ishikari).
Nakamura et al. 2003, 152, Honshu (Hiroshima).
Kawase et al. 2004, 109, Honshu (Ibaraki).
Ishiwata et al. 2005, 243-244, Honshu (Kanagawa).
Tanida 2005, 459, 464, male.
Mano 2010, 22, Honshu (Aichi).
Inazu and Nishida 2011, 184, male, Honshu (Hyogo).

Sortosa kisoensis: Ross 1956, 57.

Fischer 1971, 170.
Kobayashi 1980, 89-90, pl. 1-2, male, Hokkaido (Tokachi), Honshu (Yamagata,Kanagawa).
Kobayashi 1973, 30, Honshu (Yamagata).
Kobayashi 1984b, 28, Honshu (Kanagawa).
Kobayashi 1985a, 8, Tsushima.
Kobayashi 1985b, 68, Honshu (Kanagawa).
Kobayashi 1987, 14, Honshu (Shimane).
Kadowaki 1998, 2, Honshu (Shimane), Oki-shoto.

Kisaura kisoensis: Kobayashi 1971, 22-23, pl. 13, male, female, Honshu (Kanagawa).

Katsuma 2005a, 4, Honshu (Ibaraki).
Katsuma 2006, 41, Honshu (Ibaraki).
Mey and Nozaki 2006, 24, Honshu (Gifu).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Inazu 2008, 71, 89, 100, Honshu (Hyogo).
Morita 2008a, 722, Honshu (Mie).
Morita 2008c, 92-93, Honshu (Mie).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka).
Morita 2009b, 138, Honshu (Mie).
Ito et al. 2010, 58, 74, Hokkaido (Hiyama, Oshima).
Kawase and Morita 2010, 38-39, Honshu (Shiga).
Kuhara, 2011, 51, 59, 70, Hokkaido (Sorachi, Ishikari).
Moriya and Kubota, 2012, 131, Honshu (Kanagawa).
Inazu 2013c, 71, Honshu (Hyogo).
Ito and Nagasaka 2014, 10, Hokkaido (Sorachi).
Kawase and Morita 2014, 4, Honshu (Shiga).
Nishimoto and Nishimoto 2014, 63, Honshu (Aichi).
Nozaki 2016, 72, 331, 432, 450, adult, Shikoku (Kochi).
Ito 2017a, 4, Honshu (Fukui).
Tanida 2016, 81.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 536-537, male.
Uy et al. 2019, 295-298, Hokkaido (Ishikari, Iburi), Honshu (Miyagi, Yamagata, Saitama, Niigata, Nagano, Shizuoka, Shiga, Wakayama, Okayama), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Kumamoto), male, female.
Tezuka et al. 2021, 42, Honshu (Kanagawa).
Tanaka 2023a, 42, male, Honshu (Akita).
Kuhara et al. 2023, 17, Kyushu (Kagoshima).
Higuchi 2024, 21-22, Honshu (Toyama, Ishikawa).

Type locality: Honshu (Nagano), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima).
Remarks: Records of this species before 1999, when some species closely related to K. kisoensis were described from Japan (Kuhara 1999), possibly include misidentification of other species. The record of this species by Tanida (1991) was misidentification of other species (Dolophilodes sp. (af. kisoensis); Tanida et al. 1995).

Kisaura minakawai Arefina 2005
Kisaura minakawai Arefina 2005 (in Arefina and Armitage 2005), 17, male.

Katsuma 2006, 41, Honshu (Ibaraki).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Morita 2010, 39, Honshu (Shiga).
Ito and Kubo 2011, 6-7, Hokkaido (Tokachi).
Kuhara, 2011, 51, 70, Hokkaido (Sorachi, Ishikari).
Ito 2013, 53, Hokkaido (Tokachi, Kushiro, Nemuro).
Nishimoto and Nishimoto 2014, 63-64, Honshu (Aichi).
Katsuma 2016, 47, Honshu (Ibaraki).
Tanida 2016, 81.
Yamamoto and Ito 2016, 39, Shikoku (Ehime).
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Ito 2017a, 4, Honshu (Fukui).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 115, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Kuhara 2018, 536, 538, male.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Kuhara 2019, 237-238, Shikoku (Tokushima, Ehime, Kochi).
Tezuka et al. 2021, 42, Honshu (Kanagawa).
Torii et al. 2022, 36-38, larva, molecular phylogeny, habitat, Honshu (Shizuoka, Wakayama).
Aoya and Tanaka, 2023, 26, Honshu (Akita).
Ito and Kuhara 2023, 56, Hokkaido (Soya).
Kuhara et al. 2023, 17, Kyushu (Oita).
Tanaka 2024, 27, Honshu (Akita).
Higuchi 2024, 22, Honshu (Toyama).

Kisaura imparis Hur and Morse 2006, 217-219, male, Honshu (Nara). Synonymized by Kuhara (2019)

Tanida 2016, 81.

Type locality. Sakhalin, Russia.
Distribution. Japan (Hokkaido, Honshu, Kyushu, Ryukyu (Yakushima)), Russia (Sakhalin).

Kisaura niitakaensis (Kobayashi 1973)
Sortosa niitakaensis Kobayashi 1973, 29-30, pl. 6, male, Honshu (Yamagata).

Kobayashi 1980, 90-91, pl. 1-2, male, Hokkaido (Tokachi), Honshu (Iwate, Yamagata, Kanagawa, Niigata, Nagano).

Kobayashi 1987, 14, Oki-shoto.
Kadowaki 1998, 2, Honshu (Shimane).
Tanida 2016, 81.

Type locality. Honshu (Yamagata), Japan.
Distribution. Japan (Hokkaido, Honshu).

Kisaura nozakii (Kuhara 1999)
Dolophilodes nozakii Kuhara 1999, 180-181, male, Honshu (Tokyo, Kanagawa, Niigata, Nagano, Shizuoka, Mie, Shiga, Osaka, Wakayama).

Morita 2003, 24, Honshu (Mie).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Tanida 2005, 459, 464, male.
Inazu and Nishida 2011, 184, male, Honshu (Hyogo).

Kisaura nozakii: Torii and Hattori 2006, 35, Honshu (Shizuoka).

Inazu 2008, 71, 89-90, 100, Honshu (Hyogo).
Morita 2008a, 722, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Kawase and Morita 2010, 39, Honshu (Mie, Shiga).
Inazu 2012, 6, Honshu (Hyogo).
Inazu 2013c, 71, Honshu (Hyogo).
Morita 2013, 61, Honshu (Mie).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 10, Shikoku (Ehime).
Nishimoto and Nishimoto 2014, 64, Honshu (Aichi).
Nozaki 2016, 72, 331, 432, 450, adult, Shikoku (Kochi).
Tanida 2016, 81.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 536-537, male.
Kuhara 2019, 238-239, Shikoku (Tokushima, Ehime, Kochi).

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).

Kisaura nozakii species complex
Kisaura nozakii species complex : Suzuki et al. 2023, Honshu (Kanagawa, Nagano, Shizuoka, Shiga, Wakayama, Okayama, Hiroshima), Shikoku (Ehime), Kyushu (Miyazaki), molecular phylogeography.

Higuchi 2024, 22, Honshu (Ishikawa, Toyama).

Remarks: Suzuki et al. (2023) found that “Kisaura nozakii” contains several genetically differentiated lineages, part of which were distributed sympatrically, and therefore they treated it species complex.

Kisaura tsudai (Botosaneanu 1970)
Dolophilodes tsudai Botosaneanu 1970, 294-295, 337, male, Honshu (Nagano).

Tanida and Takemon 1993, 242, Honshu (Kyoto).
Tanida et al. 1995, 2, Honshu (Nara).
Morita 1996, 303, Honshu (Mie).
Kagaya et al. 1998, 44, 132, Honshu (Tokyo).
Kuhara 1999, 175-177, male, Honshu (Saitama, Tokyo, Kanagawa, Fukui, Yamanashi, Nagano, Shizuoka, Kyoto, Wakayama, Shimane, Hiroshima), Kyushu (Fukuoka, Kumamoto, Oita).

Morita 2003, 24, Honshu (Mie).
Tanida 2005, 459, 463-464, male.
Inazu and Nishida 2011, 184, male, Honshu (Hyogo).
Imasaka et al. 2016, 34, Kyushu (Fukuoka).

Kisaura tsudai: Torii and Hattori 2006, 35, Honshu (Shizuoka).

Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 39, Honshu (Mie, Shiga).
Kawase et al. 2010, 108 Honshu (Shimane).
Inazu 2013c, 71, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka).
Moriya and Kubota, 2012, 131, Honshu (Kanagawa).
Morita 2013, 62, Honshu (Mie).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 10-11, Shikoku (Ehime).
Nozaki 2016, 72, 331, 432, 450, adult, Shikoku (Kochi).
Yamamoto and Ito 2016, 39, Shikoku (Ehime).
Tanida 2016, 82.
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 115, Honshu (Okayama).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Kuhara 2018, 536-537, male.
Kuhara 2019, 238, Shikoku (Tokushima, Ehime, Kochi).
Yamamoto et al. 2019, 70, Shikoku (Ehime).
Morita 2020a, 18, Honshu (Mie).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Yamamoto 2023, 32, Shikoku (Ehime).
Higuchi 2024, 22, Honshu (Ishikawa, Toyama).
Yamamoto 2024, 36, Shikoku (Ehime).

Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu, Ryukyu (Yakushima)), Russia (southern Kurils).
Remarks. Records of southern Kurils should be reexamined (Kuhara 1999).

Genus Wormaldia McLachlan
Sixteen named species of this genus have been recorded in Japan. Figures or pictures were also provided for male genitalia (Morita 1995, 1999a, Inazu and Nishida 2011) of unidentified species of the genus. Larval records of Wormaldia subnigra in Japan by Iwata (1927a, 1927b) were considered to be misidentification of other species (Tsuda 1942b, 1958, Kuhara 2005b).

Wormaldia amamiensis Kuhara 2016
Wormaldia amamiensis Kuhara 2016, 266-267, male, Amami-oshima.

Kuhara 2018, 541-542, male.

Type locality. Amami-oshima, Japan.
Distribution. Japan (Amami-oshima).

Wormaldia apophysis Kuhara 2016
Wormaldia apophysis Kuhara 2016, 262-264, male, female Amami-oshima, Tokuno-shima.

Kuhara 2018, 540-541, male.

Type locality. Amami-oshima, Japan.
Distribution. Japan (Amami-oshima, Tokuno-shima).

Wormaldia carinata (Schmid 1991)
Doloclanes carinata Schmid 1991, 105, 107, male, Okinawa-jima.
Wormaldia carinata: Shimura et al. 2014, 44, Amami-oshima. [misspelled as “carina”]

Kuhara 2016, 259-260, male, female, Amami-oshi,a Okinawa-jima.
Tanida 2016, 82.
Kuhara 2018, 539-540, male.

Type locality. Okinawa-jima, Japan.
Distribution. Japan (Amami-oshima, Okinawa-jima).

Wormaldia fujinoensis Kobayashi 1980
Wormaldia fujinoensis Kobayashi 1980, 101-102, pl. 1, pl. 8, male, Honshu (Kanagawa).

Kuhara 2005b, 229-231, male, Honshu (Fukui, Gifu, Shizuoka, Aichi, Mie, Shiga, Osaka, Nara, Wakayama).
Tanida 2005, 460, 465, male.
Morita 2006, 199, Honshu (Mie).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Morita 2008a, 722, Honshu (Mie)
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 139, Honshu (Mie).
Kawase and Morita 2010, 39, Honshu (Mie, Shiga).
Nishimoto and Nishimoto 2014, 64, Honshu (Aichi).
Horiguchi 2016, 410, Honshu (Tokyo).
Tanida 2016, 82.
Nozaki 2018, 928, Honshu (Kanagawa).
Kuhara 2018, 539-540, male.
Taira 2020, 8, Honshu (Nara).
Morita 2020b, Honshu (Mie).
Nojima 2022, 38, Honshu (Okayama)

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Honshu (central, west)).

Wormaldia ishigakiensis Kuhara 2016
Wormaldia ishigakiensis Kuhara 2016, 267-268, male, female,Ishigaki-jima.

Kuhara 2018, 541-542, male.

Type locality. Ishigaki-jima, Japan.
Distribution. Japan (Ishigaki-jima).

Wormaldia itoae Kuhara 2016
Wormaldia itoae Kuhara 2016, 265-266, male, Yakushima.

Kuhara 2018, 539-540, male.

Type locality. Yakushima, Japan.
Distribution. Japan (Ryukyu (Yakushima)).

Wormaldia kadowakii Kobayashi 1980
Wormaldi kadowakii Kobayashi 1980, 101, pl. 1, pl. 8, male, Honshu (Shimane).

Kobayashi 1987, 15, Honshu (Shimane).
Kuhara 2005b, 232-233, male, Honshu (Shizuoka, Mie), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Nagasaki, Kumamoto).
Tanida 2005, 460.
Yamamoto and Ito 2014, 11, Shikoku (Ehime).
Tanida 2016, 82.
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 539-540, male.
Shimura and Imasaka 2020, 13, Kyushu (Fukuoka).
Yamamoto et al. 2021, 72, Shikoku (Ehime).

Type locality. Honshu (Shimane), Japan.
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).

Wormaldia kisoensis (Tsuda 1942)
Naganagapetus kisoensis (nomen nudum): Tsuda 1941, 104-105, Honshu (Nagano).
Naganagapetus kisoensis Tsuda 1942a, 253-254, male, Honshu (Nagano).
Wormaldia kisoensis: Ross 1956, 44, 66, male.

Fischer 1971, 202.
Kobayashi 1980, 102-103, male (misspelled as “Dolocanes”).
Morita 2000, 60, Honshu (Mie).

Morita 2001b, 219, Honshu (Aichi).
Morita 2003, 25, Honshu (Mie).
Kuhara 2005b, 241-243, male, Honshu (Mie).
Tanida 2005, 460.
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 139, Honshu (Mie).
Yamamoto and Ito 2014, 11, Shikoku (Ehime).
Tanida 2016, 82.
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 541-542, male.
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Higuchi 2024, 22, Honshu (Ishikawa).

Type locality: Honshu (Nagano), Japan.
Distribution. Japan (Honshu (central), Shikoku).

Wormaldia nabewarina Kobayashi 1969
Wormaldia nabewarina Kobayashi 1969, 20-21, pl. 4, male, Honshu (Kanagawa) (nabewarinus).

Kobayashi 1971, 24-25, pl. 15, male, Honshu (Kanagawa) (nabewarinus).
Kobayashi 1980, 99-100, pl. 1, pl. 7, male, Honshu (Niigata, Kanagawa) (nabewarinus).
Moriya 1994, 69, Honshu (Kanagawa) (nabewarinus).
Nozaki 1997a, 34, Honshu (Kanagawa) (nabewarinus).
Kuhara 2005b, 231-232, male, Honshu (Kanagawa, Toyama), Shikoku (Ehime, Kochi), Kyushu (Oita).
Tanida 2005, 460, 463, wings (nabewarinus).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka).
Nozaki 2016, 72, 332, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 82.
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 539-540, male.
Shimura and Imasaka 2021, 3, Koshikijima-retto.
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Tanaka 2023a, 42, male, Honshu (Akita).
Higuchi 2024, 22, Honshu (Ishikawa, Toyama).

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Honshu (central), Shikoku, Kyushu, Koshikijima-retto).

Wormaldia niiensis Kobayashi 1985
Wormaldia niiensis: Kobayashi 1985a, 9-10, male, Tsushima.

Nozaki and Nakamura 2005, 220, Honshu (Shimane).
Kuhara 2005b, 237-239, new synonym, male, Hokkaido (Kushiro, Hidaka, Oshima), Honshu (Akita, Yamagata, Niigata, Gifu, Kyoto, Hyogo, Nara, Wakayama, Tottori, Shimane, Hiroshima), Shikoku (Tokushima, Ehime), Kyushu (Fukuoka), Tsushima, Yakushima.
Tanida 2005, 460.
Kameyama et al. 2006, 92, Honshu (Hiroshima).
Katsuma 2006, 41, Honshu (Ibaraki).
Inazu 2008, 72-73, 90, 100, Honshu (Hyogo).
Morita 2008c, 92-93, Honshu (Mie).
Mano 2010, 22, Honshu (Aichi).
Inazu and Nishida 2011, 185, male, female, Honshu (Hyogo).
Morita 2011, 215, Honshu (Mie).
Inazu 2012, 6, Honshu (Hyogo).
Morita 2012, 250, Honshu (Mie).
Inazu 2013c, 71, Honshu (Hyogo).
Nozaki 2016, 72, 332, 432, 450, male, Shikoku (Kochi).
Tanida 2016, 82.
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 541-542, male.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Shimura and Imasaka 2021, 3, Koshikijima-retto.
Ito and Kuhara 2023, 56, Hokkaido (Kamikawa).
Higuchi 2024, 22, Honshu (Ishikawa).

Type locality. Tsushima, Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Russia (South Primorye), Korea, China (Liaoning), Taiwan.

Wormaldia okinawaensis Kuhara 2016
Wormaldia okinawaensis Kuhara 2016, 260-262, male, female, Okinoerabu-jima, Okinawa-jima.

Kuhara 2018, 540-541, male.

Type locality. Okinawa-jima, Japan.
Distribution. Japan (Okinoerabu-jima, Okinawa-jima).

Wormaldia rara (Kobayashi 1959)
Dolophilodes rara Kobayashi 1959, 345-347, male, Honshu (Fukuoka) (rarus).
Wormaldia kurokawana Kobayashi 1968, 1-3, pl. 1, male, female, Honshu (Niigata) (kurokawanus). Synonymized by Kuhara (2005b).

Kobayashi 1980, 99, pl. 1, pl. 6, male, Honshu (Niigata, Hiroshima) (kurokawanus).

Sortosa rara: Fischer 1971, 176.
Wormaldia rara: Kobayashi 1980, 100-101, pl. 7, male, Kyushu (Fukuoka) (rarus).

Gyotoku and Nozaki 1991, 73, Kyushu (Fukuoka) (rarus).
Yoshitaka 2004, 28, Honshu (Okayama) (rarus).
Kuhara 2005b, 235-237, new synonym, male, Honshu (Niigata, Nagano, Aichi, Shiga, Kyoto, Tottori, Shimane, Okayama, Hiroshima), Shikoku (Ehime), Kyushu (Fukuoka, Nagasaki, Kumamoto, Oita).

Tanida 2005, 460, 465, male.
Katsuma 2006, 41, Honshu (Ibaraki).
Nozaki and Nakamura 2007, 93, Honshu (Hiroshima).
Katsuma 2008, 40, Honshu (Ibaraki).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009c, 121, Honshu (Mie).
Katsuma 2010, 85, Honshu (Ibaraki).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 39, Honshu (Mie, Shiga).
Kawase and Hayashi 2011, 191, Honshu (Shimane).
Inazu and Nishida 2011, 184, male, Honshu (Hyogo).
Katsuma 2011a, 67, Honshu (Ibaraki).
Katsuma 2012, 64, Honshu (Ibaraki).
Inazu 2013a, 45, Honshu (Hyogo).
Inazu 2013b, 56, Honshu (Hyogo).
Inazu 2013c, 71, Honshu (Hyogo).
Morita 2013, 62, Honshu (Mie).
Yamamoto and Ito 2014, 11, Shikoku (Ehime).
Nishimoto and Nishimoto 2014, 64, Honshu (Aichi).
Tanida 2016, 82.
Yamamoto and Ito 2016, 39, Shikoku (Ehime).
Ito 2017a, 4, Honshu (Fukui).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 540-541, male.
Taira 2020, 8, Honshu (Nara).
Tanaka 2023b, 49, male, Honshu (Miyagi).
Kawase and Kato 2023, 5, Honshu (Niigata).
Higuchi 2024, 22, Honshu (Ishikawa).
Shimura and Imasaka 2024, 217, Kyushu (Fukuoka).

Wormaldia saekiensis Kobayashi 1980, 98-99, pl. 1, pl. 6, male, Honshu (Hiroshima). Synonymized by Kuhara (2005b).
Wormaldia yunotakiensis Kobayashi 1980, 98, pl. 1, pl. 5, male, Honshu (Niigata). Synonymized by Kuhara (2005b).
Type locality. Kyushu (Fukuoka), Japan.
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).

Wormaldia sumuharana Kobayashi 1980
Wormaldia sumuharana Kobayashi 1980, 100, pl. 1, pl. 7, male, Honshu (Niigata).

Tanida 2016, 83.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Honshu (central)).

Wormaldia tectum Kuhara 2016
Wormaldia tectum Kuhara 2016, 264-265, male, Okinawa-jima.

Kuhara 2018, 540-541, male.

Type locality. Okinawa-jima, Japan.
Distribution. Japan (Okinawa-jima).

Wormaldia uonumana Kobayashi 1980
Wormaldia uonumana Kobayashi 1980, 97-98, pl. 1, pl. 5, male, Honshu (Niigata).

Schmid 1991, 95-96, male, Honshu? (Higashiyama).
Kadowaki 1998, 2, Honshu (Shimane).
Kuhara 2005b, 239-241, male, Hokkaido (Ishikari, Shiribeshi), Honshu (Aomori, Yamagata, Shizuoka, Mie, Hyogo, Tottori, Shimane), Shikoku (Ehime, Kochi).
Tanida 2005, 460.
Morita 2006, 47, Honshu (Mie).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Inazu 2008, 73, 90, 100, Honshu (Hyogo).
Morita 2009a, 6, Honshu (Shiga).
Ito et al. 2010, 58, 74, Hokkaido (Hiyama, Oshima).
Kawase et al. 2010, 108 Honshu (Shimane).
Inazu and Nishida 2011, 185, male, Honshu (Hyogo).
Kuhara, 2011, 51, 70, Hokkaido (Sorachi, Ishikari).
Inazu 2013c, 71, Honshu (Hyogo).
Ito and Nagasaka 2014, 10, Hokkaido (Sorachi).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 11, Shikoku (Ehime).
Tanida 2016, 83.
Kuhara 2017, 57-58, larva, Hokkaido (Ishikari) (larval record).
Nojima 2017, 115, Honshu (Okayama).
Kuhara 2018, 539, 541-542, male, larva.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Aoya and Tanaka, 2023, 26, Honshu (Akita).
Tanaka 2023a, 42, Honshu (Akita).
Kawase and Kato 2023, 5, Honshu (Niigata).
Higuchi 2024, 22, Honshu (Ishikawa, Toyama).

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku).

Wormaldia yakuensis Kobayashi 1980
Wormaldia yakuensis Kobayashi 1980, 102, pl. 1, pl. 8, male, Yakushima.

Kuhara 2005b, 233-235, male, Kyushu (Fukuoka, Oita, Miyazaki, Kagoshima), Yakushima.
Tanida 2005, 460.
Shimura 2015, 52, male, Fukue-jima.
Kuhara 2016, 268-269, Yakushima.
Tanida 2016, 83.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Kuhara 2017, 57-58, larva, Kyushu (Miyazaki) (larval record).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Kuhara 2018, 539, 541-542, male, larva.

Type locality. Yakushima, Japan.
Distribution. Japan (Kyushu, Fukue-jima, Ryukyu (Yakushima)).

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