Author: Naotoshi KUHARA
E-mail: naotoshi.kuhara[AT]nifty.com
Last updated: 16 Aug. 2023
Subfamily Chimarrinae Rambur
Chimarra has been solely recorded in this subfamily in Japan.
Genus Chimarra Stephens
In Japan only one named species was recorded, but an unnamed species
accompanied with an alphabetic code was described for larva, which
was not associated with adult. Tanida (2003, 2005) illustrated
unidentified larval specimens of this genus. Kuhara (2017) pointed
out that the larva tentalively described by Furuya (1974) as Chimarra
sp. CB belongs the genus Wormaldia.
Type locality. Honshu (Nagano), Japan. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu,
Oki-shoto, Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Korea. Remarks. Kuhara (2017) considered that the larva tentatively
described as Chimarra sp. CA by Kumode (1952) and Furuya
(1974) is C. tsudai, but that by Tanida (1985) is a
different species.
Subfamily Philopotaminae Stephens
Three genera, Dolophilodes, Kisaura and Wormaldia
have been recorded from Japan.
Genus Dolophilodes Ulmer
Nine named species were recorded in Japan (Kuhara 2005a). Several
illustrations of male genitalia (Yamanouchi 1987, Morita 1996,
1998b) were also provided for unidentified species of "Dolophilodes",
but their generic placement should be reexamined because Kisaura
was elevated from a subgenus of Dolophilodes (s.l.)
to generic status (Malicky 1993, see also Blahnik 2005). Kuhara
(2017) described larvae of 8 out of 9 species recorded in Japan, but
failed to clarify the identities D. sp. DA tentatively
described based on larval specimens by Kumode (1952) and Furuya
(1974) and that of D. sp. DB by Furuya (1974). Kuhara
(2017), however, associated each larvae illustrated and tentatively
named as D. sp. DA, DB and DD by Tanida (1985, 2005) to
named species and transferred Dolophilodes sp. DC by him to
the genus Wormaldia.
Dolophilodes angustata Kuhara 2005
Dolophilodes sp. DA: Tanida 1985, 172-173, larva.
Reidentified by Kuhara (2017).
Type locality. Honshu (Kanagawa), Japan. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu,
Ryukyu (Yakushima)), Russia (southern and northern Kurils).
Type locality. Honshu (Niigata), Japan. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu,
Rishiri-to).
Genus Kisaura Ross
Eleven species of this genus have been recorded in Japan. Yamanouchi
(1987) illustrated the male genitalia and wing of an unidentified
species as "Dolophilodes sp. 2", which was treated as "Dolophilodes
(Kisaura)" by Ohkawa (1999). Morita (1997) illustrated the
male genitalia of an unidentified species of this genus. Inazu and
Nishida (2014) also showed pictures of the male genitalia of two
unidentified species.
Kisaura ashizuriensis Kuhara 2019
Kisaura ashizuriensis Kuhara 2019, 231-233, Shikoku
(Kochi), male. Type locality. Shikoku (Kochi), Japan. Distribution. Japan (Shikoku).
Type locality: Honshu (Nagano), Japan. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu,
Tsushima). Remarks: Records of this species before 1999, when some
species closely related to K. kisoensis were described from
Japan (Kuhara 1999), possibly include misidentification of other
species. The record of this species by Tanida (1991) was
misidentification of other species (Dolophilodes sp. (af. kisoensis);
Tanida et al. 1995).
Kisaura minakawai Arefina 2005
Kisaura minakawai Arefina 2005 (in Arefina and
Armitage 2005), 17, male.
Katsuma 2006, 41, Honshu (Ibaraki).
Torii and Hattori 2006, 35, Honshu (Shizuoka).
Ito et al. 2010, 58, 74, Hokkaido (Shiribeshi, Hiyama,
Oshima).
Kawase and Morita 2010, 39, Honshu (Shiga).
Ito and Kubo 2011, 6-7, Hokkaido (Tokachi).
Kuhara, 2011, 51, 70, Hokkaido (Sorachi, Ishikari).
Ito 2013, 53, Hokkaido (Tokachi, Kushiro, Nemuro).
Nishimoto and Nishimoto 2014, 63-64, Honshu (Aichi).
Katsuma 2016, 47, Honshu (Ibaraki).
Tanida 2016, 81.
Yamamoto and Ito 2016, 39, Shikoku (Ehime).
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Ito 2017a, 4, Honshu (Fukui).
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 115, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Kuhara 2018, 536, 538, male.
Tanaka 2019, 26, 28, male, Honshu (Akita).
Kuhara 2019, 237-238, Shikoku (Tokushima, Ehime, Kochi).
Tezuka et al. 2021, 42, Honshu (Kanagawa).
Kisaura imparis Hur and Morse 2006, 217-219, male,
Honshu (Nara). Synonymized by Kuhara (2019)
Tanida 2016, 81.
Type locality. Sakhalin, Russia. Distribution. Japan (Hokkaido, Honshu, Kyushu, Ryukyu
(Yakushima)), Russia (Sakhalin).
Kisaura niitakaensis (Kobayashi 1973)
Sortosa niitakaensis Kobayashi 1973, 29-30, pl. 6, male,
Honshu (Yamagata).
Kisaura tsudai: Torii and Hattori 2006, 35, Honshu
(Shizuoka).
Morita 2008c, 92-93, Honshu (Mie).
Morita 2009a, 6, Honshu (Shiga).
Morita 2009b, 138, Honshu (Mie).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase and Morita 2010, 39, Honshu (Mie, Shiga).
Kawase et al. 2010, 108 Honshu (Shimane).
Inazu 2013c, 71, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 105, Honshu (Osaka).
Morita 2013, 62, Honshu (Mie).
Kawase and Morita 2014, 4, Honshu (Shiga).
Yamamoto and Ito 2014, 10-11, Shikoku (Ehime).
Nozaki 2016, 72, 331, 432, 450, adult, Shikoku (Kochi).
Yamamoto and Ito 2016, 39, Shikoku (Ehime).
Tanida 2016, 82.
Kuhara and Ito 2017, 15, Ryukyu (Yakushima).
Nojima 2017, 115, Honshu (Okayama).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Kuhara 2018, 536-537, male.
Kuhara 2019, 238, Shikoku (Tokushima, Ehime, Kochi).
Yamamoto et al. 2019, 70, Shikoku (Ehime).
Shimura and Imasaka 2020, 12, Kyushu (Fukuoka).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Yamamoto 2023, 32, Shikoku (Ehime).
Type locality. Honshu (Nagano), Japan. Distribution. Japan (Honshu (central to west), Shikoku,
Kyushu, Ryukyu (Yakushima)), Russia (southern Kurils). Remarks. Records of southern Kurils should be reexamined
(Kuhara 1999).
Genus Wormaldia McLachlan
Sixteen named species of this genus have been recorded in Japan.
Figures or pictures were also provided for male genitalia (Morita
1995, 1999a, Inazu and Nishida 2011) of unidentified species of the
genus. Larval records of Wormaldia subnigra in Japan by
Iwata (1927a, 1927b) were considered to be misidentification of
other species (Tsuda 1942b, 1958, Kuhara 2005b).
Type locality. Tsushima, Japan. Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu,
Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Russia (South
Primorye), Korea, China (Liaoning), Taiwan.
Wormaldiasaekiensis Kobayashi 1980, 98-99,
pl. 1, pl. 6, male, Honshu (Hiroshima). Synonymized by
Kuhara (2005b). Wormaldia yunotakiensis Kobayashi 1980, 98, pl. 1, pl.
5, male, Honshu (Niigata). Synonymized by Kuhara (2005b). Type locality. Kyushu (Fukuoka), Japan. Distribution. Japan (Honshu (central to west), Shikoku,
Kyushu).
Wormaldia sumuharana Kobayashi 1980
Wormaldia sumuharana Kobayashi 1980, 100, pl. 1, pl.
7, male, Honshu (Niigata).
Tanida 2016, 83.
Type locality. Honshu (Niigata), Japan. Distribution. Japan (central).
Type locality. Yakushima, Japan. Distribution. Japan (Kyushu, Fukue-jima, Ryukyu (Yakushima)).
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