Rhyacophilidae

A catalogue of Japanese Trichoptera
Family Rhyacophilidae Stephens

Authors: Akikazu TAIRA* and Takao NOZAKI
*: Correspondence. E-mail: akikazu-t[AT]mbf.nifty.com
Released: 24: Apr. 2020

Only two genera, Himalopsyche Banks and Rhyacophila Stephens, are known from Japan.

Genus Himalopsyche Banks

A single species, Himalopsyche japonica (Morton), has been known from Japan.

Himalopsyche japonica Morton 1900

Rhyacophila japonica Morton 1900, 3-4, pl. 1, male, Japan.

Matsumura 1931, 1136, adult.
Kuwayama 1932, 1513, adult.
Tsuda 1948, 2, 7-8, larva.
Ishihara et al. 1953, 37, Shikoku (Ehime).
Kuwada 1965, 163, pl. 82, adult, Shikoku (Ehime).

Himalopsyche japonica: Ross 1956, 110-111, 123, male.

Tsuda 1959, 130, larva.
Tsuda and Akagi 1962, 124-125, 127, larva.
Kuwada 1965, 163, pl. 82, adult, Shikoku (Ehime).
Saito 1965, 15, larva.
Schmid and Botosaneanu 1966, 134, 156, 158, 168, pl. IV, male.
Kobayashi 1982, 96, Honshu (Kanagawa).
Kobayashi 1984c, 28, 29, Honshu (Kanagawa).
Kobayashi 1985b, 68, 69, 71, 73, Honshu (Kanagawa).
Tanida 1985, 181, larva.
Nozaki 1988b, 40, Honshu (Kanagawa).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 34, 75, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 31, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 32, 74, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 185.
Tsuruishi 1999, 15-27, adult, pupa, larva, habitat, biology. Honshu (Nagano).
Kuranishi and Yamamoto 2000, 60, Shikoku (Ehime).
Tsuruishi and Yoshida 2002, 193-202, microhabitat, Honshu (Nagano).
Tsuruishi et al. 2002, 29-33, degree-days accumulation, Honshu (Nagano).
Tsuruishi 2003a, 11-18, larva, pupa, life cycle, Honshu (Nagano).
Tsuruishi 2003b, 24-33, adult, pupa, larva, Honshu (Nagano).
Tsuruishi and Yoshida 2003a, 23-27, respiratory rate, Honshu (Nagano).
Tsuruishi and Yoshida 2003b, gut content, Honshu (Nagano).
Nozaki 2004, 1232, Honshu (Kanagawa).
Tsuruishi 2004, larva, pupa, life history, Honshu (Nagano).
Hattori 2005, 414-415, male, larva.
Ishiwata et al. 2005, 110, 244, 245, Honshu (Kanagawa).
Nozaki 2005, 106, adult, larva.
Tsuruishi and Yoshida 2005, 436-439, developmental pause, Honshu (Nagano).
Tsuruishi 2006, 494-498, life cycle, Honshu (Nagano).
Tanida 2008, 247-248, pl. 92, adult, Shikoku (Ehime).
Tsuruishi 2010, 7-10, behavior, Honshu (Nagano).
Morita and Kawase 2012, 15, Honshu (Ishikawa).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Kawase and Morita 2014, 3, Honshu (Shiga).
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 62.
Nozaki 2016, 69, 307, 308, 429, 449, adult, Shikoku (Kochi).
Kawase et al. 2017, 23, Honshu (Shiga).
Hattori 2018, 474-475, male, larva.
Hjalmarsson et al. 2018, 89, 91-94, 96-99, larva, Honshu, Shikoku.

Type locality. Japan.
Distribution. Japan (Honshu, Shikoku).

Genus Rhyacophila Stephens

Fifty seven species have been known from Japan, but the specific status of a species redescribed as R. yukii by Schmid (1970) have not yet been clarified. Furthermore, 11 'species' separated only by larval characters have not yet been associated with their adults.

Rhyacophila arefini Lukyanchenko 1993

Rhyacophila arefini Lukyanchenko 1993, 5-6, male, female, pupa, larva, Kunashir, Hokkaido (Ishikari).

Kuhara and Ito 1994, 19, 23, Hokkaido (Eniwa).
Arefina 1997, 29, 31, 32, male, female.
Kuhara 1997b, 94, Hokkaido (Otaru).
Ito et al. 1998, 10, 12-14, 16, 17, Hokkaido (Nemuro).
Kuranishi 1999, 188.
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 24, Hokkaido (Soya).
Arefina 2001, 22, 41, female, Shikotan.
Kuhara and Sato 2001, 64, Rishiri-to.
Hattori 2005, 420, 427, 434, male, female, larva.
Ohkawa and Hattori 2007, 266, Hokkaido (Kamikawa).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Ito et al. 2009, 8, Hokkaido (Okhotsk).
Ito and Kubo 2011, 4, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Hidaka).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Kuranishi 2016, 62.
Hattori 2018, 480, 487, 495, male, female, larva.

Type locality. Kunashir.
Distribution. Japan (Hokkaido, Rishiri-to), Kunashir, Shikotan.

Rhyacophila articulata Morton 1900

Rhyacophila articulata Morton 1900, 5, pl. 1, male, Japan.

Tsuda 1940, 132-134, male, Honshu (Nagano, Shiga, Kyoto).
Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942a, 245, Honshu (Shiga, Nagano).
Tsuda and Kuwayama 1950, 414, adult.
Ross 1956, 82, 117, male.
Tsuda and Kuwayama 1950, 414, adult.
Ross 1956, 82, 117, male.
Miwa and Okawa 1958, 17, Honshu (Mie).
Kobayashi 1959, 343, Kyushu (Fukuoka).
Kuwada 1965, 163, pl. 82, adult, Honshu (Gifu).
Tsuda and Akagi 1974, 124-125, 127, larva.
Tanida 1980, 135, Honshu (Ishikawa).
Tanida 1982b, 18-19, Honshu (Ishikawa).
Kobayashi 1983, 32-36, Honshu (Kanagawa).
Nishiharu 1997, 59, Honshu (Fukui).
Kuranishi 1999, 185.
Hattori 2005, 421, 430, male.
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Kuranishi 2016, 62.
Hattori 2018, 481, 490, male.

Type locality. Japan.
Distribution. Japan (Honshu, Kyushu).
Remarks. Hattori (2005) pointed out that records as this species in previous reports were most probably R. lezeyi.

Rhyacophila azumaensis Kobayashi 1973

Rhyacophila azumaensis Kobayashi 1973, 27, pl. 5, male, Honshu (Yamagata).

Kuranishi 1999, 190.
Kuranishi et al. 2002, 98-99, Honshu (Aomori).
Hattori 2005, 426, 435, male, female.
Kuranishi 2016, 62.
Hattori 2018, 486, 495, male, female.

Rhyacophila asahiensis Kobayashi 1976, 51-52, 55, male, Honshu (Niigata). Synonymized by Kuranishi (2016).

Kuranishi 1999, 190.

Rhyacophila hayachinensis Kobayashi 1976, 53-54, 56, male, Honshu (Iwate). Synonymized by Kuranishi (2016)

Kuranishi 1999, 190.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Hattori (2005) pointed out that R. asahiensis and R. hayachinensis are most probably same species as this species based on examination of original illustrations by Kobayashi (1973, 1976). Kuranishi (2016) synonymized them with this species.

Rhaycophila bilobata Ulmer 1907

Rhyacophila bilobata Ulmer 1907, 84-85, male, Japan.

Tsuda 1940, 134, male, Honshu (Kyoto, Nagano).
Ross 1956, 76, 99, 121, male.
Schmid 1970, 43, 121, pl. 10, male.
Kobayashi 1971b, 9-10, pl. 2, male, female, Honshu (Kanagawa).
Tanida 1985, 186, larva.
Nozaki 1988b, 40, Honshu (Kanagawa).
Tanida 1991, 34, Honshu (Nara).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 34, 75, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 31, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 76, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 186.
Hattori 2005, 419, 425, 433, male, female, larva.
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Morita 2007a, 12, Honshu (Mie).
Kawase and Morita 2010, 35, Honshu (Shiga)
Moriya and Kubota 2012, 133, Honshu (Kanagawa)..
Kuranishi 2016, 62.
Hattori 2018, 479, 485, 493, male, female, larva.

Type locality. Japan.
Distribution. Japan (Honshu).

Rhyacophila brevicephala Iwata 1927

Rhyacophila brevicephala Iwata 1927a, 217, pls. 5-6, larva, Honshu (Tochigi, Yamanashi, Kyoto, Tottori).

Iwata 1927b, 205, pls. 1-2, larva, Honshu (Tochigi, Yamanashi, Kyoto, Tottori).
Tsuda 1942a, 246, Honshu (Kyoto).
Tsuda 1942b, 263-267, Honshu (Kyoto).
Tsuda 1948, 16, larva.
Tsuda and Kuwayama 1950, 414, male.
Tsuda 1959, 133, larva.
Tsuda and Akagi 1962, 126-128, larva.
Schmid 1970, 55, 123, pl. 22, male.
Tsuda and Akagi 1974, 126-128, larva.
Tani 1977, 198, male.
Yamanouchi 1978, 9, 10, adult, Hokkaido (Tokachi).
Tanida 1982a, 358, Honshu (Gunma).
Kobayashi 1983, 32-36, Honshu (Kanagawa).
Tanida 1985, 186, larva.
Kobayashi 1987a, 20, Honshu (Shimane).
Nozaki 1988a, 45, 46, Honshu (Kanagawa).
Nozaki 1988b, 40, Honshu (Kanagawa).
Kadowaki 1990 1990, 14, Okino-shima.
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Tanida 1990, 55, Honshu (Hyogo).
Hatta and Nozaki 1991, 202, Honshu (Aichi).
Tanida 1991, 34, Honshu (Nara).
Moriya 1992, 15, 42, Honshu (Kanagawa).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Morita 1993a, 5, Honshu (Mie).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Moriya 1994, 34, 75, Honshu (Kanagawa, Yamanashi).
Tanida et al. 1995, 2-3, Honshu (Nara).
Arefina, 1997, 29, 33, male.
Ito et al. 1997, 23, 25, 27, 29, 31, Hokkaido (Hiyama).
Kuhara 1997a, 59, Hokkaido (Shiribeshi).
Moriya 1997, 30, 65, Honshu (kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nishiharu 1997, 60, Honshu (Fukui).
Nozaki 1997a, 31, Honshu (Kanagawa).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kadowaki 1998, 6, Honshu (Shimane).
Kagaya et al. 1998, 21, 77, Honshu (Tokyo, Yamanashi).
Yodoe et al. 1998, 68, Honshu (Shimane).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 187.
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 25, Hokkaido (Ishikari).
Ito et al. 2000b, 23, 25-26, Hokkaido (Soya).
Kuhara et al. 2000, 247, Hokkaido (Iburi).
Kuranishi and Yamamoto 2000, 60, Shikoku (Ehime).
Maruyama and Takai 2000, 72, 74, 76, larva.
Arefina 2001, 23, 37, female, Kunashir, Shikotan.
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001a, 218, Honshu (Mie).
Morita 2001b, 336, Honshu (Mie).
Morita 2001d, 175, Honshu (Mie).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Kuranishi and Satake 2003, 36, Hokkaido (Okhotsk).
Ito et al. 2004, 11, Hokkaido (Kushiro).
Kawase et al. 2004, 106, Honshu (Ibaraki).
Uenishi and Yoshitaka 2004, 15, Honshu (Okayama).
Yoshitaka 2004, 27, Honshu (Okayama).
Hattori 2005, 420, 429, 436, male, female, larva.
Ishiwata et al. 2005, 110, 244-245, Honshu (kanagawa).
Katsuma 2005a, 2, Honshu (Ibaraki).
Katsuma 2005b, 11, Honshu (Ibaraki, Gumma).
Nagayasu et al. 2005, 53, Hokkaido (Ishikari).
Nozaki 2005, 106, adult, larva.
Kameyama et al. 2006, 92, Honshu (Hiroshima).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Ito et al. 2007, 147, Hokkaido (Kushiro).
Morita 2007a, 12, Honshu (Mie).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nakamura and Hajiri 2007, 197, Honshu (Hiroshima).
Nakamura and Tsujimura 2007, 76, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 92, Honshu (Hiroshima).
Nozaki and Tanida 2007, 242, Honshu (Shizuoka).
Ohkawa and Hattori 2007, 264-265, Hokkaido (Kamikawa).
Inazu 2008, 60-61, 86, 98, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 101, 103, Honshu (Osaka).
Tanaka 2009, 18, Honshu (Akita).
Aoyagi and Iwasaki 2010, 33, 35, Honshu (Osaka).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Morita. 2010, 35, Honshu (Mie).
Mano 2010, 22-23, Honshu (Aichi).
Tanaka 2010b, 16, Honshu (Akita).
Inazu and Nishida 2011, 171, male, larva, Honshu (Hyogo).
Ito ank Kubo 2011, 4, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Ishikari).
Morita 2011, 214, Honshu (Mie).
Taira and Tanida 2011, 29-32, 34, habitat, Honshu (Nara).
Katsuma 2012b, 63, Honshu (Ibaraki).
Morita 2012, 249, Honshu (Mie).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Tanaka 2012a, 21, 23, male, Honshu (Akita).
Ito 2013, 48, Hokkaido (Nemuro, Okhotsk, Kushiro, Tokachi).
Morita 2013, 60, Honshu (Mie).
Kawase and Morita. 2014, 3, Honshu (Shiga).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachii).
Kuranishi and Kikuchi 2014, 362, Honshu (Yamagata).
Taira and Tanida 2014, larva, behavior, Honshu (Nara).
Nishimoto 2014, 57, Honshu (Aichi).
Shimura 2015, 53, Goto-retto.
Hayashi et al. 2016, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 62.
Nozaki 2016, 70, 312, 313, male, Shikoku (Kochi).
Ito 2017a, 2l, Honshu (Fukui).
Ito 2017b, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 23, Honshu (Shiga).
Nojima 2017, 109, Honshu (Okayama).
Saito et al. 2017, 182-183, Honshu (Chiba).
Tanaka 2017, 2, Honshu (Akita).
Hattori 2018, 480, 489, 496, male, female, larva.
Nozaki et al. 2019, 167, Honshu (Nagano).
Yamamoto et al. 2019, 69, Shikoku (Ehime).

Rhyacophila lacrimae Tsuda 1940, 125-126, male, Honshu (Kyoto). Synonymized by Tsuda (1942a).
Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Okino-shima, Goto-retto), Kuriles.
Remarks. Kuranishi (2016) synonymized Rhyacophila nakagawai Kobayashi 1969 with this species, but did not provide any evidence. Since morphology of male genitalia of R. nakagawai described or redescribed by Kobayashi (1969, 1971b) do not agree with that of this species, we list these two species separately.

Rhycophila clara Hattori and Ohkawa 2007

Rhyacophila clara: Ohkawa and Hattori, 2007, 267-270, male, female, pupa, larva, Hokkaido (Okhotsk, Kamikawa, Sorachi, Hidaka).

Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Hidaka).
Kuranishi 2016, 62-63.

Type locality. Japan (Hokkaido).
Distribution. Japan (Hokkaido).

Rhyacophila clemens Tsuda 1940

Rhyacophila clemens Tsuda 1940, 124-125, male, Honshu (Nagano, Kyoto).

Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942a, 246, Honshu (Nagano, Shiga, Kyoto).
Tsuda 1948, 11-12, larva, Honshu (Tochigi, Nagano, Kyoto).
Tsuda and Kuwayama 1950, 414, male.
Ishihara et al. 1953, 37, Shikoku (Ehime).
Ross 1956, 87, 118, male.
Kobayashi 1959, 343, Kyushu (Fukuoka).
Tsuda 1959, 237, larva.
Ishihara et al. 1953, 37, Shikoku (Ehime).
Tsuda and Akagi 1962, 125, 127-128, larva
Kuwada 1965, 163, pl. 82, male, Shikoku (Ehime).
Kobayashi 1968, 1, Honshu (Niigata).
Schmid 1970, 95, 133, male, Japan.
Kobayashi 1971b, 10-11, pl. 3, male, female, Honshu (Kanagawa).
Tsuda and Akagi 1974, 125, 127-128, larva.
Tani 1977, 198, male.
Kobayashi 1983, 32-36, Honshu (Kanagawa).
Tanida 1985, 186, larva.
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 40, Honshu (Kanagawa).
Kadowaki 1990, 14, Okino-shima.
Hatta and Nozaki 1991, 202, Honshu (Aichi, Gifu).
Tanida 1991, 34, Honshu (Nara).
Gyotoku and Nozaki 1992, 14, Kyushu (Fukuoka).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Ito et al. 1997, 23, 25, 27, 29, 31, Hokkaido (Hiyama).
Kuranishi 1997, 266, Iriomote-jima.
Nishiharu 1997, 59, Honshu (Fukui).
Kuhara 1997a, 60, Hokkaido (Shiribeshi).
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nishiharu 1997, 59, Honshu (Fukui).
Nozaki 1997a, 31, Honshu (Kanagawa).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kagaya et al. 1998, 21, 78, Honshu (Tokyo, Yamanashi).
Nishimura and Tei, 1998, 70-72, larval distribution, Honshu (Hyogo).
Sekimoto et al. 1998, 330, Honshu (Hiroshima).
Kuranishi 1999, 187.
Kuranishi and Yamamoto 2000, 60, Shikoku (Ehime).
Uenishi and Kurumi 2000, 278, Honshu (Shiga).
Kawase and Morita 2001, 8, Honshu (Ishikari).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001a, 219, Honshu (Mie).
Morita 2001b, 336, Honshu (Mie).
Aoyagi et al. 2004, 23-24, Honshu (Kyoto).
Kawase et al. 2004, 106, Honshu (Ibaraki).
Morita and Kawase 2004, 16, Honshu (Ishikawa).
Hattori 2005, 417, 423, 432, male, female, larva.
Ishiwata et al 2005, 110, 244-245, Honshu (Kanagawa).
Katsuma 2005a, 2, Honshu (Ibaraki).
Katsuma 2005b, 11, Honshu (Ibaraki).
Nozaki and Nakamura 2005, 218, Honshu (Shimane).
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Kuhara et al. 2007, ap. 67, Hokkaido (Ishikari).
Morita 2007a, 12, Honshu (Mie).
Inazu 2008, 61-62, 86, 99, Honshu (Hyogo).
Tanaka 2008, 39, Honshu (Akita).
Tanida 2008, 248, pl. 92, adult, Shikoku (Ehime).
Aoyagi and Iwasaki 2009, 101, 103, Honshu (Osaka).
Aoyagi and Iwasaki 2010, 32, 33, 35, 36, Honshu (Osaka).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Inazu and Nishida 2011, 171, male, larva, Honshu (Hyogo).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Ishikari).
Inazu 2012, 2, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Tanaka 2012a, 21, 22, male, Honshu (Akita).
Kawase and Morita 2014, 3, Honshu (Shiga).
Nozaki et al. 2014, 59, Honshu (Nagano).
Hayashi et al. 2016, 28, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 63.
Nozaki 2016, 313, 314, male.
Ito 2017a, 2, Honshu (Fukui).
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 477, 483, 492, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Okino-shima, Iriomote-jima), Taiwan.

Rhyacophila coclearis Hsu and Chen 1996

Rhyacophila coclearis Hsu and Chen 1996, 125-127, male, female, Taiwan.

Kuranishi 1999, 186, Okinawa-jima.
Hattori 2005, 422, 430, male.
Kuranishi 2016, 63.
Hattori 2018, 482, 491, male.

Type locality. Taiwan.
Distribution. Japan (Okinawa-jima), Taiwan.

Rhyacophila crassa Schmid 1970

Rhyacophila crassa Schmid 1970, 151, pl. 24, male, Honshu (Fukushima).

Kuranishi 1999, 187.
Hattori 2005, 429, 437, male.
Kuranishi 2016, 63.
Hattori 2018, 489, 497, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila curtior Schmid 1970

Rhyacophila curtior Schmid, 1970, 207-208, pls. 42, 52, male, female, Honshu (Nagano).

Kuranishi 1999, 186.
Hattori 2005, 422, 430, male.
Kuranishi 2016, 63.
Hattori 2018, 482, 490, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila diffidens Tsuda 1940

Rhyacophila diffidens Tsuda 1940, 128-129, male, Honshu (Kyoto).

Tsuda 1942a, 247, Honshu (Kyoto).
Tani 1977, 198, male.
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Kuranishi 1999, 189.
Morita 2001a, 219, Honshu (Mie).
Morita 2001b, 336, Honshu (Mie).
Hattori 2005, 428, 437, male, female.
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Morita 2007a, 12, Honshu (Mie).
Morita 2009c, 7, Honshu (Shiga).
Aoyagi and Iwasaki 2010, 32, 33, 35, 36, Honshu (Osaka).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Kuranishi 2016, 63.
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 488, 497, male, female.

Rhyacophila tsudai Ross 1956, 93, 119-120, male, Honshu (Kyoto). Synonymized by Kuranishi (2016).

Kuranishi 1999, 189.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Hattori (2005) pointed out that R. tsudai is probably the same species as this species, and Kuranishi (2016) synonymized it with this species.

Rhyacophila flinti Schmid 1970

Rhyacophila flinti Schmid 1970, 208, pl. 42, male, Kyushu (Kumamoto).

Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Kuranishi 1999, 186.
Hattori 2005, 422, 430, male, female.
Kuranishi 2016, 63.
Nozaki 2016, 309, 313, male.
Hattori 2018, 482, 490, male, female.

Type locality. Japan (Kyushu).
Distribution. Japan (Kyushu).

Rhyacophila formosana Ulmer 1927

Rhyacophila formosana Ulmer 1927, 173-174, 182, pls. 5-4, male, Taiwan.

Ross 1956, 107, 122, male.
Kobayashi 1987b, 41, male, Taiwan.
Kuranishi 1997, 266, Iriomote-jima.
Kuranishi 1999, 187.
Hattori 2005, 424, 433, male.
Kuranishi 2016, 63.
Hattori 2018, 484, 493, male.

Type locality. Taiwan.
Distribution. Japan (Iriomote-jima), Taiwan.

Rhyacophila hamata Hattori and Ohkawa 2007

Rhyacophila hamata: Ohkawa and Hattori 2007, 270-274, male, female, larva, Hokkaido (Soya, Okhotsk, Tokachi, Kamikawa, Sorachi, Hidaka, Ishikari).

Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Hidaka).
Kuranishi 2016, 63.

Type locality. Japan (Hokkaido).
Distribution. Japan (Hokkaido).

Rhyacophila hayakawai Kobayashi 1969

Rhyacophila hayakawai Kobayashi 1969, 18-19, pl. 3, male, Honshu (Kanagawa).

Kobayashi 1971b, 16-17, pl. 8, male, Honshu (Kanagawa).
Kuranishi 1999, 187.
Kuranishi 2016, 63-64.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Hattori (2005) suggested that this species resembles R. brevicephala belonging to R. brevicephala Species Group, but Kuranishi (2016) stated that this species belongs to R. anatina Species Group. Furthermore, Kuranishi (2016) mentioned that a part of description of this species reversed with that of R. nakagawai, and figures d, e, f in plate 3 (Kobayashi 1969) belongs to this species. However, Kobayashi (1971b) used those figures for redescription of R. nakagawai and different figures (c, g, h in plate 3) for this species.

Rhyacophila hokkaidensis Iwata 1927

Rhyacophila hokkaidensis Iwata 1927a, 214-215, pl. 5-6, larva, Hokkaido (Ishikari).

Iwata 1927b, 203-204, pls. 1-2, larva, Hokkaido (Ishikari).
Lepneva 1964, 262-265, larva, Russia (Sakhalin).
Yamanouchi 1978, 9, Hokkaido (Tokachi).
Kuhara and Ito 1994, 19, 23, Hokkaido (Ishikari).
Arefina 1997, 24, 28, male.
Ito et al. 1997, 23, 25, 27, 29, Hokkaido (Hiyama).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Ito 1999, 28-34, life history, Hokkaido (Ishikari).
Kuranishi 1999, 185-186.
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 25, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya).
Arefina 2001, 31, 43, female, Itrup, Shikotan, Kunashir.
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Kuhara and Sato 2001, 64, larva, Rishiri-to.
Kuranishi and Satake 2003, 35, Hokkaido (Okhotsk).
Hattori 2005, 417, 421, 430, male, female, larva.
Kuhara et al. 2006, 45-47, 49, Hokkaido (Ishikari).
Ito et al. 2007, 147, Hokkaido (Kushiro).
Kuhara et al. 2007, 67, Hokkaido (Ishikari).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Ohkawa and Hattori 2007, 265-266, Hokkaido (Kamikawa).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Ito et al. 2009, 8, Hokkaido (Okhotsk).
Ito et al. 2010, 53-54, 72, Hokkaido (Shiribeshi).
Tsuji et al. 2010, 5, Hokkaido (Okhotsk).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Ishikari, Hidaka).
Ito 2013, 48, Hokkaido (Nemuro, Okhotsk, Kushiro).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Kuranishi 2016, 64.
Nozaki 2016, 308-309, 313, male.
Ito 2017b, 39, Hokkaido (Kamikawa).
Hattori 2018, 477, 481, 490, male, female, larva.

Rhyacophila tenninkyoensis Kobayashi 1964, 83-85, male, Hokkaido (Kamikawa). Synonymized by Kuranishi (1999).
Rhyacophila sakhalinica Schmid 1970, 206, pl. 40, male, Sakhalin. Synonymized by Arefina (1997).
Type locality. Japan (Hokkaido).
Distribution. Japan (Hokkaido, Rishiri-to), Sakhalin, Kuriles (Itrup, Shikotan, Kunashir).

Rhyacophila impar Martynov 1914

Rhyacophila impar Martynov 1914, 77-792, male, Russia (Altai).

Lepneva 1964, 256-262, larva, Russia (Altai, Sayans, East Siberia).
Schmid 1970, 132, pls. 41, 51, male, female.
Tanida 1985, 186, larva.
Tanida 1991, 34, Honshu (Nara).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Arefina 1997, 24, 29, 31, male, female.
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Kagaya et al. 1998, 21, 79, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 187.
Arefina 2001, 32, 43, female, Russia (Khabarovsk Territory).
Hattori 2005, 419, 426, 435, male, female, larva.
Tanaka 2010b, 16, 18, 19, male, Honshu (Akita).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Tanaka 2012a, 21, 23, male, Honshu (Akita).
Nozaki et al. 2014, 59, Honshu (Nagano).
Tanaka 2014, 38, Honshu (Akita).
Tanaka 2015, 54, Honshu (Akita).
Kuranishi 2016, 64.
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 479, 486, 495, male, female, larva.

Rhyacophila sp. RG: Tsuda 1948, 14-15, larva, Honshu (Tochigi, Nagano).

Tsuda and Akagi 1962, 126-128, larva.

Rhyacophila tacita Tsuda 1940, 127-128, male, Honshu (Nagano). Synonymized by Ross (1956).

Tsuda and Akagi 1974, 126-128, larva.

Type locality. Russia (Altai).
Distribution. Japan (Honshu), Russia.
Remarks. Tsuda (1974) treated Rhyacophila sp. RG as Rhyacophila tacita, and Ross (1956) synonymized the latter with this species.

Rhyacophila ishihanaensis Kobayashi 1984

Rhyacophila ishihanaensis Kobayashi 1984a, 6-7, male, Sado-ga-shima.

Kuranishi 2016, 64.

Type locality. Japan (Sadoga-shima).
Distribution. Japan (Sadoga-shima).
Remarks. Hattori (2005) and Kuranishi (2016) suggested that this species probably the same species as Rhyacophila yamanakensis.

Rhyacophila itoi Tsuda and Kawai 1967

Rhyacophila itoi Tsuda and Kawai 1967, 111-112, male, Honshu (Tochigi, Shiga, Nara).

Tanida 1990, 55, Honshu (Hyogo).
Tanida 1991, 34, Honshu (Nara).
Morita 1998a, 46, Honshu (Mie).
Kuranishi 1999, 186.
Morita 2000, 59, Honshu (Mie).
Morita 2001a, 219, Honshu (Mie).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Hattori 2005, 419, 425, 433, male, female, larva.
Katsuma 2005a, 2, Honshu (Ibaraki).
Katsuma 2005b, 11, Honshu (Ibaraki, Gumma).
Mey and Nozaki 2006, 24, Honshu (Gifu).
Morita 2006b, 199, Honshu (Mie).
Morita 2007a, 13, Honshu (Mie).
Morita 2008b, 91, Honshu (Mie).
Morita 2009a, 136, Honshu (Mie)
Morita 2009c, 7, Honshu (Mie).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Morita 2011, 214, Honshu (Mie).
Katsuma 2012b, 63, Honshu (Ibaraki).
Morita 2013, 60, Honshu (Mie).
Kuranishi 2016, 64.
Hattori 2018, 477, 485, 493, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila kardakoffi Navás 1926

Rhyacophila kardakoffi Navás 1926, 57-58, pl. 1, male, Russia (South Ussuri).

Schmid 1970, 64, 125, pl. 48, male.
Arefina 1997, 25, 31, male, female.
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Arefina 2001, 25-26, figs. 34-37, female, Russia (Primorye Territory).
Hattori 2005, 420, 427, 434, male, female, larva.
Ito et al. 2007, 147, Hokkaido (Kushiro).
Ohkawa and Hattori 2007, 267, Hokkaido (Kamikawa).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Hidaka).
Ito 2013, 48, Hokkaido (Kushiro, Tokachi).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Kuranishi 2016, 64.
Hattori 2018, 480, 487, 494, male, female, larva.

Type locality. Russia (South Ussuri).
Distribution. Japan (Hokkaido), Russia.

Rhyacophila kawamurae Tsuda 1940

Rhyacophila kawamurae Tsuda 1940, 130-131, male, Honshu (Kyoto), North Korea (Saitoku).

Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942, 247, Honshu (Nagano, Shiga).
Tsuda and Akagi 1962, 128, larva.
Schmid 1970, 72, 127, pl. 28, male.
Kobayashi 1971b, 12-13, pl. 5, male, female, Honshu (Kanagawa).
Tsuda and Akagi 1974, 128, larva.
Tani 1977, 198, male.
Yamanouchi 1978, 9, male, Hokkaido (Tokachi).
Tanida 1985, 186, larva.
Nozaki 1988a, 45, Honshu (Kanagawa).
Nozaki 1988b, 40, Honshu (Kanagawa).
Tanida 1991, 34, Honshu (Nara).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Hori et al. 1993, 375, Honshu (Aichi).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Moriya 1994, 34, 76, Honshu (Kanagawa).
Arefina 1997, 30, 33, male.
Ito et al. 1997, 23, 25, 27, 29, Hokkaido (Hiyama).
Kuranishi 1997, 266, Amami-oshima, Okinawa-jima.
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nozaki 1997a, 31, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 80, Honshu (Tokyo, Yamanashi).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kuranishi 1999, 187.
Ito et al. 2000a, 25, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya).
Maruyama and Takai 2000, 72, 77, 78, larva, Honshu (Nara).
Uenishi and Kurumi 2000, 278, Honshu (Shiga).
Arefina 2001, 29-30, 42, female, Kunashir.
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Morita 2001a, 219, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Nozaki and Nakamura 2002, 166, Honshu (Hiroshima).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Hattori 2005, 419, 424, 432, male, female, larva.
Ishiwata et al. 2005, 110, 244-245, Honshu (Kanagawa).
Nozaki and Nakamura 2005, 218, Honshu (Shimane).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Morita 2007a, 13, Honshu (Mie).
Ohkawa and Hattori 2007, 266, Hokkaido (Kamikawa).
Inazu 2008, 61, 86, 99, Honshu (Hyogo).
Ito et al. 2010, 54, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Morita and Kawase 2010, 27, Honshu (Ishikawa).
Inazu and Nishida 2011, 171, male, larva, Honshu (Hyogo).
Ito and Kubo 2011, 52, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Ishikari).
Morita 2011, 214, Honshu (Mie).
Taira and Tanida 2011, 29-34, habitat, Honshu (Nara).
Inazu 2012, 2, 4, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Morita 2013, 60, Honshu (Mie).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Kawase and Morita 2014, 3, Honshu (Shiga).
Kuranishi and Kikuchi 2014, 362, Honshu (Yamagata).
Nishimoto 2014, 57, Honshu (Aichi).
Taira and Tanida 2014, behavior, Honshu (Nara).
Shimura 2015, 53, Goto-retto.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 64.
Nozaki 2016, 69, 310, 313, 429, 449, male, Shikoku (Kochi).
Ito 2017b, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 23, Honshu (Shiga).
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 479, 484, 492, male, female, larva.

Type locality. Japan (Honshu), North Korea.
Distribution. Japan (Honshu, Shikoku, Kyushu, Goto-retto, Amami-oshima, Okinawa-jima).
Remarks. Tsuda (1940) recorded this species from Japan and North Korea. Although Malicky (2014) synonymized this species with a continental species, Rhyacophila manuleata Martynov 1934, but Kuranishi (2016) recognized this species as a distinct species. Further study is needed to clarify the specific status of this species.

Rhyacophila kawaraboensis Kobayashi 1976

Rhyacophila kawaraboensis Kobayashi 1976, 53, 55-56, male, Honshu (Iwate).

Kuranishi 1999, 190.
Kuranishi et al. 2002, 99, Honshu (Aomori).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Hattori. 2005, 423, 432, male.
Katsuma 2006, 33, Honshu (Ibaraki).
Katsuma 2011b, 48, 49, Honshu (Ibaraki).
Tanaka 2014, 38, Honshu (Akita).
Kuranishi 2016, 64-65.
Hattori 2018, 483, 492, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila kisoensis Tsuda 1940

Rhyacophila kisoensis Tsuda 1940, 132, male, Honshu (Nagano).

Ross 1956, 95-96, 230, male.
Tsuda and Akagi 1962, 126-128, larva.
Schmid 1970, 65, 125, pl. 15, male, Japan.
Kobayashi 1971b, 15-16, pl. 7, male, female, Honshu (Kanagawa).
Kobayashi 1973, Honshu (Yamagata).
Tsuda and Akagi 1974, 126-128, larva.
Tani 1977, 198, male.
Tanida 1982a, 358, Honshu (Gunma).
Tanida 1985, 186, larva.
Hatta and Nozaki 1991, 202, Honshu (Gifu).
Tanida 1991, 34, Honshu (Nara).
Hori et al. 1993, 375, Honshu (Aichi).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Nishiharu 1997, 59, Honshu (Fukui).
Kagaya et al. 1998, 21, 81, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 188.
Kuranishi and Yamamoto 2000, 60, Shikoku (Ehime).
Maruyama and Takai 2000, 72, 76, 77, larva.
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Hattori 2005, 420, 427, 435, male, female, larva.
Nozaki 2005, 106, adult, larva.
Nozaki and Nakamura 2005, 218, Honshu (Shimane).
Inazu 2008, 62, 86, 99, Honshu (Hyogo).
Kameyama et al. 2009, 110, Honshu (Shimane, Hiroshima).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Tanaka 2010a, 23, 26, 28, male, Honshu (Akita).
Inazu and Nishida 2011, 172, male, larva, Honshu (Hyogo).
Katsuma 2011a, 37, Honshu (Ibaraki).
Morita 2011, 214, Honshu (Mie).
Inazu 2012, 4, Honshu (Hyogo).
Kawase and Hayashi 2012, 255, Honshu (Shimane).
Morita and Kawase 2012, 14, Honshu (Ishikawa).
Tanaka 2012a, 21, 23, male, Honshu (Akita).
Tanaka 2012b, 11, Honshu (Akita).
Kawase and Morita 2014, 35, male, Honshu (Shiga).
Kuranishi and Kikuchi 2014, 362, Honshu (Yamagata).
Tanaka 2014, 38, Honshu (Akita).
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 65.
Nozaki 2016, 312, 313, male.
Kawase et al. 2017, 23, Honshu (Shiga).
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 480, 487, 495, male, female, larva.
Tanaka 2018b, 8, Honshu (Akita).

Rhyacophila sp. RI: Tsuda 1948, 15, larva, Honshu (Tochigi). Treated by Tsuda and Akagi (1974) as this species.

Tsuda and Akagi 1962, 126-128, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Shikoku).

Rhyacophila kiyosumiensis Kuranishi 1990

Rhyacophila kiyosumiensis Kuranishi 1990, 109-112, male, female, larva, Honshu (Chiba).

Kuranishi 1992, 261-262, adult, oviposition behavior, Honshu (Chiba).
Moriya 1997, 30, 65, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 31-32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 82, Honshu (Tokyo, Yamanashi).
Kuranishi 2002, 564-565, larva, adult, Honshu (Chiba).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Katsuma 2005a, 2, Honshu (Ibaraki).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Katsuma 2011b, 48, Honshu (Ibaraki).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 65.
Saito et al. 2017, 183, Honshu (Chiba).

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila kohnoae Ross 1956

Rhyacophila kohnoae Ross 1956, 93, 119, male, Honshu (Fukushima?).

Kobayashi 1987a, 20, Honshu (Shimane).
Kadowaki 1990, 14, Okino-shima.
Kuranishi 1999, 189.
Hattori 2005, 420, 428, 436, male, female, larva.
Katsuma 2005b, 11, Honshu (Ibaraki, Gumma).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Morita 2008b, 91, 92, Honshu (Mie).
Morita 2009a, 137, Honshu (Mie).
Morita 2009c, 7, Honshu (Shiga).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Morita 2011, 214, Honshu (Mie).
Nishimoto and Nishimoto 2014, 62, Honshu (Aichi).
Tanaka 2015, 55, Honshu (Akita).
Hayashi et al. 2016, 29, Okino-shima.
Kuranishi 2016, 65.
Kawase et al. 2017, 23-24, Honshu (Shiga).
Hattori 2018, 480, 488, 496, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Okino-shima).

Rhyacophila kuramana Tsuda 1942

Rhyacophila kuramana Tsuda 1942a, 247-248, male, Honshu (Shiga, Kyoto).

Ross 1956, 92, male.
Tanida 1982b, 18-19, Honshu (Ishikawa).
Tanida 1985, 186, larva, Honshu.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Nishiharu 1997, 59, Honshu (Fukui).
Kuranishi 1999, 189.
Kuranishi and Yamamoto 2000, 60-61, Shikoku (Ehime).
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Morita and Kawase 2004, 16, Honshu (Ishikawa).
Hattori 2005, 420, 428, 436, male, female, larva.
Inazu 2008, 62, 86, 99, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 101, 103, Honshu (Osaka).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Kawase et al. 2010, 108, Honshu (Shimane).
Morita and Kawase 2010, 26, 27, Honshu (Ishikawa).
Inazu and Nishida 2011, 172, male, larva, Honshu (Hyogo).
Inazu 2012, 4, Honshu (Hyogo).
Morita and Kawase 2012, 16, Honshu (Ishikawa).
Taira et al. 2013, 125-136, larva, habitat, life cycle, Honshu (Nara).
Kawase and Morita 2014, 3, Honshu (Shiga).
Hayashi et al. 2016, 29, Okino-shima.
Kuranishi 2016, 65.
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 109, Honshu (Okayama).
Taira 2017, 83-85, Honshu (Osaka).
Hattori 2018, 480, 488, 496, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Shikoku, Kyushu).

Rhyacophila kuwayamai Schmid 1970

Rhyacophila kuwayamai Schmid 1970, 164, pl. 28, male, Honshu (Kyoto).

Tani 1976, 392, Tsushima.
Tani 1977, 198, male.
Yamanouchi 1978, 9, Hokkaido (Tokachi).
Hatta and Nozaki 1991, 202, Honshu (Aichi, Gifu).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 188.
Ito et al. 2000a, 25, Hokkaido (Soya).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya)
Kuhara et al. 2000, 247, Hokkaido (Iburi).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Hattori 2005, 419, 424, 432, male, female, larva.
Katsuma 2006, 34, Honshu (Ibaraki).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Ohkawa and Hattori 2007, 266, Hokkaido (Kamikawa).
Inazu 2008, 62-63, 86, 99, Honshu (Hyogo).
Morita 2009b, 121, Honshu (Mie).
Ito et al. 2010, 53, 72, Hokkaido (Hiyama, Oshima).
Inazu and Nishida 2011, 172, male, larva, Honshu (Hyogo).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Ishikari).
Morita 2011, 214, Honshu (Mie).
Inazu 2012, 4, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Ito 2013, 48, Hokkaido (Kushiro).
Morita 2013, 60, Honshu (Mie).
Iijima and Tsuchiya 2014, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Kawase and Morita 2014, 3, Honshu (Shiga).
Kuranishi 2016, 65.
Nakatani 2016, 240, Hokkaido (Nemuro).
Nozaki 2016, 69, 310, 313, 429, 449, male, Honshu (Gunma).
Ito 2017a, 2-3, Honshu (Fukui).
Ito 2017b, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 109, Honshu (Okayama).
Hattori 2018, 479, 484, 492-493, male, female, larva.
Nozaki et al. 2019, 167, Honshu (Nagano).

Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu, Kyushu, Tsushima).

Rhyacophila lambakanta Schmid 1970

Rhyacophila lambakanta Schmid 1970, 209, pl. 41, male, Honshu (Nagano).

Tani 1978, 7-8, male, Honshu (Niigata).
Emoto 1979, 556-560, 562-563, 565, male, female, Honshu (Yamanashi, Nagano, Gifu).
Tanida 1980,135, Honshu (Ishikawa).
Tanida 1982b, 18-19, Honshu (Ishikawa).
Kobayashi 1984b, 5, Sadoga-shima.
Nozaki 1988b, 40, Honshu (Kanagawa).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 83, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 188.
Morita and Kawase 2004, 17, Honshu (Ishikawa).
Ishiwata et al. 2005, 244-245, Honshu (Kanagawa).
Hattori 2005, 422, 430, male, female.
Katsuma 2012, 53, Honshu (Ibaraki).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Nozaki et al. 2014, 59, Honshu (Nagano).
Kuranishi 2016, 65. Honshu.
Nozaki 2016, 70, 311, 313, 429, 449, male, Honshu (Nagano).
Kawase et al. 2017, 24, Honshu (Shiga).
Hattori 2018, 482, 491, male, female.

Rhyacophila imagamiensis Kobayashi 1973, 22, pl. 3, male, Honshu (Yamagata). Synonymized by Tani (1978).
Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Sadoga-shima).

Rhyacophila lezeyi Navás 1933

Rhyacophila lezeyi Navás 1933, 94, male, Honshu? (Kofou).

Ross 1956, 82, 117, male.
Nozaki 1988b, 40, Honshu (Kanagawa).
Morita 1989, 39-40, Honshu (Mie).
Tanida 1991, 34, Honshu (Nara).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 84, Honshu (Tokyo).
Kuranishi 1999, 186.
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Morita 2001a, 219, Honshu (Mie).
Morita 2003, 23, Honshu (Mie).
Kawase et al. 2004, 107, Honshu (Ishikawa).
Morita and Kawase 2004, 17, Honshu (Ishikawa).
Hattori 2005, 417, 421, 430, male, female, larva.
Ishiwata et al. 2005, 110, 244-245, Honshu (Kanagawa).
Katsuma 2005b, 11, Honshu (Ibaraki).
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Inazu 2008, 63, 86, 99, Honshu (Hyogo).
Tanida 2008, 248, pl. 92, adult, Honshu (Kyoto).
Morita 2009a, 137, Honshu (Mie).
Morita 2009c, 7, Honshu (Shiga).
Morita and Kawase 2009, 28, Honshu (Ishikawa).
Aoyagi and Iwasaki 2010, 32, 33, 35, 36, Honshu (Osaka).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Inazu and Nishida 2011, 172, male, larva, Honshu (Hyogo).
Inazu 2012, 4, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Kawase and Hayashi 2012, 255, Honshu (Shimane).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Tanaka 2012a, 20, 21, male, Honshu (Akita).
Kawase and Morita. 2014, 3, Honshu (Shiga).
Nozaki et al. 2014, 59, Honshu (Nagano).
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 65.
Nozaki 2016, 69, 309, 313, 429, 449, male, Shikoku (Kochi).
Ito 2017a, 3, Honshu (Fukui).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 109-110, Honshu (Okayama).
Aoya 2018, 41-43, 45, behavior, Honshu (Akita).
Hattori 2018, 477, 481, 490, male, female, larva.
Aoya 2019, 23-24, Honshu (Akita).
Aoya 2020, 29-42, microhabitat, Honshu (Akita).

Rhyacophila articulata: Tsuda 1948, 5-6, larva, Honshu (Nagano). Misidentification mentioned by Kuranishi (1948).

Tsuda 1959, 130, larva. Misidentification mentioned by Kuranishi (1999).
Kobayashi 1971b, 7-9, male, female, Honshu (Kanagawa). Misidentification mentioned by Kuranishi (1999).
Tani 1977, 197, 198, male. Misidentification mentioned by Kuranishi (1999).
Tsuda and Akagi 1962. 124-125, 127, larva. Misidentification mentioned by Kuranishi (1999).
Tani 1978, 7-8, male, Honshu (Niigata). Misidentification mentioned by Kuranishi (1999).

Rhyacophila sp. RF: Tsuda 1948, 13-14, larva, Honshu (Nagano, Kyoto). Mentioned by Hattori (2005).

Tsuda and Akagi 1962, 126-128, larva.

Rhyacophila morokuboensis Kobayashi 1971, 2-3, pl. 2, male, Honshu (Kanagawa). Synonymized by Kuranishi (1999).

Kobayashi 1971b, 18-19, pl. 10, male, Honshu (Kanagawa).

Rhyacophila towadensis: Tanaka 2010b, 16, 18, 19, male, Honshu (Akita). Misidentification mentioned by Tanaka (2018a).
Type locality. Japan (Honshu?).
Distribution. Japan (Honshu, Shikoku, Kyushu).

Rhyacophila makiensis Kobayashi 1987

Rhyacophila makiensis Kobayashi 1987, 20, 33, male, Honshu (Shimane).

Kuranishi 1999, 190.
Hattori 2005, 429, 437, male.
Morita 2006a, 47, Honshu (Mie).
Inazu 2008, 63, 86, 99, Honshu (Hyogo).
Morita 2008c, 718, Honshu (Mie).
Morita 2009a, 137, Honshu (Mie).
Inazu and Nishida 2011, 173, male, Honshu (Hyogo).
Kuranishi 2016, 65.
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 489, 497, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila mayaensis Kobayashi 1973

Rhyacophila mayaensis Kobayashi 1973, 23-24, pl. 4, male, Honshu (Yamagata).

Kuranishi 1999, 189.
Uchida 2002, 77, Honshu (Tochigi).
Hattori 2005, 427, male.
Katsuma 2006, 35, Honshu (Ibaraki).
Tanaka 2012a, 21, 23, male, Honshu (Akita).
Tanaka 2014, 38, Honshu (Akita).
Kuranishi 2016, 65.
Hattori 2018, 487, male.
Tanaka 2018, 21, Honshu (Akita).

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila minoyamaensis Kobayashi 1973

Rhyacophila minoyamaensis Kobayashi 1973, 28-29, pl. 6, male, Honshu (Yamagata).

Kuranishi 1999, 189.
Hattori 2005, 428, 437, male.
Kuranishi 2016, 66.
Hattori 2018, 488, 497, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila mirabilis Levanidova and Schmid 1977

Rhyacophila mirabilis Levanidova and Schmid 1977, 501-503, male, Kunashir, Iturup.

Ito et al. 1998, 10, Hokkaido (Nemuro).
Kuranishi 1999, 189.
Arefina 2001, 22, figs. 1-4, female, Kunashir, Iutrup.
Kuhara and Sato, 2001, 64, Rishiri-to.
Kuranishi and Satake 2003, 35-36, Hokkaido (Okhotsk).
Hattori 2005, 419, 426, 434, male, female, larva.
Ito and Kosugi 2007, 40, Hokkaido (Kushiro).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Ohkawa and Hattori 2007, 267, Hokkaido (Kamikawa).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Ito et al. 2009, 8, Hokkaido (Okhotsk).
Ito and Kubo 2011, 52, 56, 57, 61, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 61, Hokkaido (Sorachi, Hidaka).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Ito 2013, 49, Hokkaido (Nemuro, Okhotsk, Tokachi).
Ito and Nagasaka 2014, 8 Hokkaido (Sorachi).
Kuranishi 2016, 66.
Nozaki, 2016, 313, 314, male.
Ito 2017b, 39, Hokkaido (Kamikawa).
Hattori 2018, 479, 486, 494, male, female, larva.

Rhyacophila sp.: Yamanouchi 1987, 39, male, Hokkaido (Tokachi). Mentioned by Kuhara and Kuranishi (1997).
Type locality. Kunashir.
Distribution. Japan (Hokkaido, Rishiri-to), Kuriles (Kunashir, Iturup).

Rhyacophila motakanta Schmid 1970

Rhyacophila motakanta Schmid 1970, 209-210, pl. 41, male, Honshu (Nagano).

Tani 1977, 198, male.
Emoto 1979, 556-560, 564-566, male, female, Honshu (Nagano, Gifu).
Ohtsuka and Tanida 1986, 5, Kyushu (Kumamoto).
Nozaki 1988b, 40, Honshu (Kanagawa).
Tanida 1990, 55, Honshu (Hyogo).
Tanida 1991, 34, Honshu (Nara).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 85, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 188.
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Hattori 2005, 422, 430, male, female.
Mey and Nozaki 2006, 24, Honshu (Nagano).
Morita 2006a, 47, Honshu (Mie).
Inazu 2008, 63, 87, 99, Honshu (Hyogo).
Tanaka 2010b, 16, Honshu (Akita).
Inazu and Nishida 2011, 173, male, Honshu (Hyogo).
Katsuma 2012, 53, Honshu (Ibaraki).
Kawase and Hayashi 2012, 255, Honshu (Shimane).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 20, 22, male, Honshu (Akita).
Tanaka 2012b, 11, Honshu (Akita).
Kawase and Morita 2014, 3, Honshu (Shiga).
Tanaka 2014, 38, Honshu (Akita).
Kuranishi 2016, 66.
Nozaki, 2016, 70, 311, 313, 429, 449, male, Shikoku (Kochi).
Ito 2017a, 3, Honshu (Fukui).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 110, Honshu (Okayama).
Tanaka 2017, 2, Honshu (Akita).
Hattori 2018, 482, 491, male, female.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Shikoku, Kyushu).

Rhyacophila nagaokaensis Kobayashi 1976

Rhyacophila nagaokaensis Kobayashi 1976, 52-53, 55, male, Honshu (Niigata).

Kuranishi 1999, 189.
Hattori 2005, 420, 428, 436, male, female, larva.
Morita 2009a, 137, Honshu (Mie).
Morita 2009c, 7, Honshu (Shiga).
Kawase and Morita 2010, 35, Honshu (Shiga, Mie).
Inazu and Nishida 2011, 173, male, Honshu (Hyogo).
Inazu 2012, 4, Honshu (Hyogo).
Morita and Kawase 2012, 14, 16, Honshu (Ishikawa).
Morita 2013, 60, Honshu (Mie).
Kawase and Morita 2014, 3, Honshu (Shiga).
Kuranishi 2016, 66.
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 480, 488, 496, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila nakagawai Kobayashi 1969

Rhyacophila nakagawai Kobayashi 1969, 17-18, pl. 3, male, Honshu (Kanagawa).

Kobayashi 1973, male, female, Honshu (Yamagata).
Tani 1977, 198, male.
Nozaki 1988a, 46, Honshu (Kanagawa).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Moriya 1994, 34, 76, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 86, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 186.
Nozaki 2004, 1233, Honshu (Kanagawa).
Hattori 2005, 419, 425, 433, male, larva.
Mey and Nozaki 2006, 24, Honshu (Gifu).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Kawase et al. 2017, 24, Honshu (Shiga).
Hattori 2018, 479, 485, 494, male, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Kuranishi (2016) synonymized this species with R. brevicephala, however, a problem between this species and R. hayakawai has not been solved (see remarks of R. hayakawai).

Rhyacophila nigrocephala Iwata 1927

Rhyacophila nigrocephala Iwata 1927a, 216, pl. 6, larva, Honshu (Tochigi, Kyoto, Hyogo, Tottori).

Iwata 1927b, 204, pl. 2, larva, Honshu (Tochigi, Kyoto, Hyogo, Tottori).
Tsuda 1948, 11, larva, Honshu (Kyoto, Nagano, Tochigi, Akita).
Tsuda and Kuwayama 1950, 414, male.
Ross 1956, 76, 105, 107, male.
Tsuda 1959, 131, larva.
Tsuda and Akagi 1962, 126, 127-128, larva.
Kobayashi 1971b, 11-12, pl. 4, male, Honshu (Kanagawa).
Tani 1977, 198, male.
Tanida 1978, 8, male, female, Honshu (Niigata).
Yamanouchi 1978, 9, Hokkaido (Tokachi).
Kobayashi 1983, 32-36, Honshu (Kanagawa).
Tanida 1985, 186, larva.
Nozaki 1988a, 45, Honshu (Kanagawa).
Kadowaki 1990, 14, Okino-shima.
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Nozaki and Gyotoku 1990, 16, Kyushu (Fukuoka).
Tanida 1990, 55, Honshu (Hyogo).
Hatta and Nozaki 1991, 202, Honshu (Gifu).
Tanida 1991, 34, Honshu (Nara).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Moriya 1992, 15, 42, Honshu (Kanagawa).
Morita 1993a, 5, Honshu (Mie).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Hori et al. 1993, 375, Honshu (Aichi).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Tanida et al. 1995, 3, Honshu (Nara).
Nishiharu 1997, 59, Honshu (Fukui).
Nishimoto and Nishimoto 1997, 150, Honshu (Aichi).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Kagaya et al. 1998, 21, 87, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 188.
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Maruyama and Takai 2000, 72-74, larva, Shikoku (Kochi).
Uenishi and Kurumi 2000, 278, Honshu (Shiga).
Morita 2001a, 219, Honshu (Mie).
Morita 2001b, 336, Honshu (Mie).
Morita 2001c, 164, Honshu (Mie).
Morita 2001d, 175, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 166, Honshu (Hiroshima).
Nakamura and Uemori 2003, 59, Honshu (Hiroshima).
Morita and Kawase 2004, 17, Honshu (Ishikawa).
Nozaki 2004, 1233, Honshu (Kanagawa).
Hattori 2005, 424, 433, male, female.
Katsuma 2005a, 3, Honshu (Ibaraki).
Katsuma 2005b, 11, Honshu (Gumma).
Nozaki and Nakamura 2005, 218, Honshu (Shimane).
Kameyama et al. 2006, 92, Honshu (Hiroshima).
Kimura et al. 2006, 7, Honshu (Nagano).
Mey and Nozaki 2006, 24, Honshu (Gifu, Shiga).
Nakamura et al. 2006, 100, Honshu (Hiroshima).
Nakamura and Fukui 2006, 186, Honshu (Hiroshima).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Morita 2007a, 13, Honshu (Mie).
Morita 2007b, 16, Honshu (Mie).
Nakamura and Hajiri 2007, 197, Honshu (Shimane).
Nakamura and Tsujimura 2007, 76, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 92, Honshu (Hiroshima).
Inazu 2008, 63, 86, 99, Honshu (Hyogo).
Kameyama et al. 2009, 110, Honshu (Shimane, Hiroshima).
Kawase and Hayashi 2010a, 80, Honshu (Shimane).
Kawase and Morita 2010, 35-36, Honshu (Shiga, Mie).
Mano 2010, 21-23, Honshu (Aichi).
Inazu and Nishida 2011, 173, male, larva, Honshu (Hyogo).
Kawase and Hayashi 2011a, 172, Honshu (Shimane).
Kawase and Hayashi 2011b, 191, Honshu (Shimane).
Morita 2011, 214, Honshu (Mie).
Inazu 2012, 4, Honshu (Hyogo).
Morita 2012, 249, Honshu (Mie).
Morita and Kawase 2012, 15, Honshu (Ishikawa).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Hayashi et al. 2013, 12-13, Okino-shima.
Kawase and Morita 2014, 3, Honshu (Shiga).
Nishimoto 2014, 57, Honshu (Aichi).
Taira and Tanida 2014, behavior, Honshu (Nara).
Hayashi et al. 2016, 28-29, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 66.
Nozaki 2016, 69, 310, 313, 429, male, Shikoku (Kochi).
Ito 2017a, 3, Honshu (Fukui).
Kawase et al. 2017, 24, Honshu (Shiga).
Kuhara and Ito 2017, 14, 16, Yakushima.
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 484, 493, male, female.
Taira 2018, 119-126, life history, microhabitat, Honshu (Nara).
Nozaki et al. 2019, 167, Honshu (Nagano).
Yamamoto et al. 2019, 69, Shikoku (Ehime).

Rhyacophila modesta Tsuda 1940, 129-130, male, Honshu (Saitama, Kyoto). Synonymized by Tsuda (1948).

Tsuda 1942a, 247, Honshu (Yamanashi, Shiga, Kyoto).
Tsuda 1942b, 263-267, Honshu (Kyoto).

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Shikoku, Kyushu, Okino-shima, Yakushima).

Rhyacophila niizakiensis Kobayashi 1976

Rhyacophila niizakiensis Kobayashi 1976, 54, 56, male, Honshu (Kanagawa).

Kuranishi 1999, 189.
Kuranishi 2016, 66.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. This species is probably the same species as Rhyacophila yosiiana Tsuda 1940 (Hattori 2005; Kuranishi 2016).

Rhyacophila nipponica Navás 1933

Rhyacophila nipponica Navás 1933, 93, male, Honshu? (Nippon moyen, Kofou).

Ross 1956, 105, 107, male.
Tani 1976, 392, Tsushima.
Tani 1977, 198, male.
Kobayashi 1984b, 5, Sado.
Kobayashi 1985, 15, Tsushima.
Kobayashi 1987a, 19-20, Honshu (Shimane).
Nozaki 1988b, 40, Honshu (Kanagawa).
Kadowaki 1990, 14, Okino-shima.
Tanida 1991, 34, Honshu (Nara).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Morita 1993b, 107, Honshu (Mie).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Moriya 1994, 34, 77, Honshu (kanagawa, Yamanashi).
Morita 1996b, 50, Honshu (Mie).
Arefina 1997, 30, 33, male.
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 88, Honshu (Tokyo, Yamanashi).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kadowaki 1998, 5, Honshu (Shimane).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 188.
Morita 1999, 27, Honshu (Mie).
Ito et al. 2000a, 25, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya).
Arefina 2001, 30-31, 42, female, Shikotan.
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Hattori 2005, 419, 424, 432, male, female, larva.
Ishiwata et al. 2005, 110, 244-245, Honshu (Kanagawa).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Kuhara et al. 2007, ap. 67, Hokkaido (Ishikari).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Tanida 2007, 242, Honshu (Shizuoka).
Ohkawa and Hattori 2007, 266, Hokkaido (Kamikawa).
Inazu 2008, 63-64, 86, 99, Honshu (Hyogo).
Morita 2008c, 719, Honshu (Mie).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Tanida 2008, 248, pl. 92, adult, Honshu (Shizuoka).
Morita and Kawase 2009, 28, Honshu (Ishikawa).
Tanaka 2009, 18, Honshu (Akita).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Oshima).
Kawase and Morita 2010, 36, Honshu (Mie).
Kobayashi 2010, 140, Honshu (Tochigi).
Inazu and Nishida 2011, 174, male, larva, Honshu (Hyogo).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 62, (Sorachi, Ishikari).
Taira and Tanida 2011, 29-34, habitat, Honshu (Nara).
Inazu 2012, 4, Honshu (Hyogo).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 21, 22, male, Honshu (Akita).
Ito 2013, 49, Hokkaido (Nemuro, Kushiro).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Taira and Tanida 2014, larva, behavior, Honshu (Nara).
Tanaka 2014, 38, Honshu (Akita).
Hayashi et al. 2016, 29, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Nozaki 2016, 311, 313, male.
Kuranishi 2016, 66.
Ito 2017a, 3, Honshu (Fukui).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 479, 484, 493, male, female, larva.
Nozaki et al. 2019, 167, Honshu (Nagano).

Type locality. Japan (Honshu?).
Distribution. Japan (Hokkaido, Honshu, Sado, Okino-shima, Tsushima), Shikotan.

Rhyacophila niwae Iwata 1927

Rhyacophila niwae Iwata 1927c, 389-390, larva, Honshu (Nagano, Gifu, Kyoto).

Tsuda 1948, 16-17, larva, Honshu (Tochigi, Nagano, Kyoto).
Tsuda 1959, 133, larva.
Tsuda and Akagi 1962, 126-128, larva.
Hattori 2005, 425, 433, male, female.
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Taira and Tanida 2011, 29-34, habitat, Honshu (Nara).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Taira and Tanida 2014, behavior, Honshu (Nara).
Kuranishi 2016, 66.
Hattori 2018, 485, 493, male, female.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Tanida (1985) synonymized this species with R. bilobata, but Hattori (2005) recognized this species as a distinct species.

Rhyacophila orthacantha Emoto 1979

Rhyacophila orthacantha Emoto 1979, 557-560, 565-568, male, female, Honshu (Aichi).

Hatta and Nozaki 1991, 202, Honshu (Aichi).
Moriya 1994, 34, 77, Honshu (Kanagawa, Yamanashi).
Nozaki 1997b, 27, 32, Honshu (Tokyo).
Kagaya et al. 1998, 21, 89, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 188.
Hattori 2005, 422, 431, male, female.
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Kawase and Morita 2010, 36, Honshu (Mie).
Kuranishi 2016, 67.
Hattori 2018, 482, 491, male, female.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila pacata Tsuda 1940

Rhyacophila pacata Tsuda 1940, 121-122, male, Honshu (Kyoto).

Tsuda 1942, 245, Honshu (Shiga).
Ross 1956, 99, 12, male.
Schmid 1970, 121, pl. 10, male.
Kuranishi 1999, 186.
Uenishi and Kurumi 2000, 278, Honshu (Shiga).
Morita 2001b, 336, Honshu (Mie).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Yoshitaka 2004, 27, Honshu (Okayama).
Hattori 2005, 425, 434, male, female.
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Morita 2007a, 13, Honshu (Mie).
Inazu 2008, 64, 86, 99, Honshu (Hyogo).
Morita 2009a, 137, Honshu (Mie).
Morita 2009c, 7, Honshu (Shiga).
Kawase and Hayshi 2010b, 84, Honshu (Shimane).
Kawase and Morita 2010, 36, Honshu (Mie).
Nakamura 2010, 238, Honshu (Hiroshima).
Inazu and Nishida 2011, 174, male, Honshu (Hyogo).
Inazu 2012, 4, Honshu (Hyogo).
Kawase and Morita. 2014, 3, Honshu (Shiga).
Kuranishi 2016, 67.
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 485, 494, male, female.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila retracta Martynov 1914

Rhyacophila retracta Martynov 1914, 75-77, male, Russia (Altai).

Ross 1956, 76, 103, 121, male.
Kobayashi 1964, 84-86, Hokkaido (Kamikawa).
Lepneva 1964, 265-271, larva.
Botosaneanu 1970, 276-277, 285, 328, female, Korea.
Schmid 1970, 90, 132, pl. 41, male.
Tani 1976, 392, Tsushima.
Tani 1977, 198, male.
Yamanouchi 1978, 9, 10, adult, Hokkaido (Tokachi).
Emoto 1979, 561-562, male, female, Hokkaido (Tokachi, Hidaka).
Kobayashi 1985, 15, Tsushima.
Tanida 1985, 181, larva.
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Saito 1996, 15, Hokkaido (Kamikawa).
Arefina, 1997, 24, 31, male, female.
Ito et al. 1997, 23, 25, 27, 29, 31, Hokkaido (Hiyama).
Kuhara 1997a, 60, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Itou and Muramatsu 1998, 39, Hokkaido (Okhotsk).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 188.
Ito et al. 2000a, 26, Hokkaido (Soya).
Arefina 2001, 33, 44, female, Russia (Khabarovska Territory).
Kuhara 2001, 16, Hokkaido (Shiribeshi).
Kuranishi and Satake 2003, 36, Hokkaido (Okhotsk).
Nagayasu et al. 2003, 106, Hokkaido (Ishikari).
Hattori 2005, 417, 421, 430, male, female, larva.
Nagayasu et al. 2005, 53, Hokkaido (Ishikari).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, ap. 67, Hokkaido (Ishikari).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Ohkawa and Hattori 2007, 266, Hokkaido (Kamikawa).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 62, Hokkaido (Sorachi, Ishikari).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Ito 2013, 49, Hokkaido (Nemuro, Okhotsk, Kushiro).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Kuranishi 2016, 67.
Nozaki, 2016, 311, 313, male.
Ito 2017b, 39, Hokkaido (Kamikawa).
Hattori 2018, 477, 481, 490, male, female, larva.

Rhyacophila uenoi Tsuda 1940, 123-124, male, Hokkaido (Kamikawa). Synonymized by Ross (1956).
Type locality. Russia (Altai).
Distribution. Japan (Hokkaido, Honshu, Tsushima), Russia, Korea.

Rhyacophila satoi Kuranishi 1997

Rhyacophila satoi Kuranishi 1997, 266-268, male, female, Amami-o-shima, Okinawa-jima.

Kuranishi 1999, 189.
Hattori 2005, 423, 432, male.
Kuranishi 2016, 67.
Hattori 2018, 483, 492, male.

Type locality. Japan (Okinawa-jima).
Distribution. Japan (Amami-o-shima, Okinawa-jima).

Rhyacophila shekigawana Kobayashi 1973

Rhyacophila shekigawana Kobayashi 1973, 25-27, pl. 5, male, Honshu (Yamagata).

Kuranishi 1999, 189.
Hattori. 2005, 428, 436, male.
Kuranishi 2016, 67.
Hattori 2018, 488, 497, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila shikotsuensis Iwata 1927

Rhyacophila shikotsuensis Iwata 1927a, 216, pl. 6, larva, Hokkaido (Ishikari), Honshu (Kyoto).

Iwata 1927b, 204, pl. 2, larva, Hokkaido (Ishikari), Honshu (Kyoto).
Tanida 1985, 186, larva.
Morita 1989, 39, Honshu (Mie).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Kuhara and Ito 1994, 19, 23, Hokkaido (Ishikari).
Ito et al. 1997, 23, 26, 28, 30, Hokkaido (Hiyama).
Kuranishi 1997, 266, Okinawa-jima.
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kagaya et al. 1998, 21, 90, Honshu (Tokyo, Yamanashi).
Ito 1999, 28-34, life history, Hokkaido (Ishikari).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 188.
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 26, Hokkaido (Soya).
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Kuhara 2001, 17, Hokkaido (Shiribeshi).
Kuranishi and Satake 2003, 36, Hokkaido (Okhotsk).
Aoyagi et al. 2004, 24, Honshu (Kyoto).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Hattori 2005, 419, 424, 432, male, female, larva.
Kuhara et al. 2006, 45-47, 49, Hokkaido (Ishikari).
Ito et al. 2007, 147, Hokkaido (Kushiro).
Kuhara et al. 2007, ap. 67, Hokkaido (Ishikari).
Ohkawa and Hattori 2007, 266, male, female, Hokkaido.
Satake and Kuranishi 2007, 280-281, Mikura-jima.
Inazu 2008, 64, 86, 99, Honshu (Hyogo).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Tanaka 2010b, 16, 18, 19, male, Honshu (Akita).
Inazu and Nishida 2011, 174, male, larva, Honshu (Hyogo).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Katsuma 2011b, 48, Honshu (Ibaraki).
Kuhara 2011, 52, 56, 57, 62, Hokkaido (Sorachi, Ishikari).
Inazu 2012, 4, Honshu (Hyogo).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 21, 22, male, Honshu (Akita).
Tanaka 2012b, 11, Honshu (Akita).
Ito 2013, 49, Hokkaido (Okhotsk, Kushiro).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Nozaki et al. 2014, 59, Honshu (Nagano).
Taira and Tanida 2014, behavior, Honshu (Nara).
Tanaka 2014, 38, Honshu (Akita).
Yamamoto and Ito 2014, 32, Shikoku (Ehime).
Hayashi et al. 2016, 28, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 67.
Nozaki, 2016, 70, 311, 313, 429, 449, male, Shikoku (Kochi).
Hayashi et al. 2017, 78, Kyushu (Miyazaki).
Ito 2017b, 39, Hokkaido (Kamikawa).
Nojima 2017, 110, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Hattori 2018, 479, 484, 492, male, female, larva.

Rhyacophila sp. RE: Tsuda 1948, 13, larva, Honshu (Tochigi, Nagano, Kyoto). Mentioned by Tanida (1985).

Tsuda and Akagi 1962, 126-128, larva.

Rhyacophila remingtoni Ross, 1956, 107, 122, male, Hokkaido (Ishikari), Honshu (Fukushima). Synonymized by Kuranishi (1997).

Schmid 1970, 72, 128, pl. 27, male.
Tani 1977, 198, male.
Nozaki 1988b, 40, Honshu (Kanagawa).
Hatta and Nozaki 1991, 202, Honshu (Aichi).
Gyotoku and Nozaki 1992, 13, Kyushu (Fukuoka).
Moriya 1994, 34, 77, Honshu (Kanagawa, Yamanashi).
Morita 1996a, 303, Honshu (Mie).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kadowaki 1998, 5, Honshu (Shimane).

Rhyacopjila similis Kobayashi 1959, 343-345, male, Kyushu (Fukuoka). Synonymized with R. remingtoni by Schmid (1970).

Kobayashi 1968, 1, Honshu (Niigata).

Type locality. Japan (Hokkaido, Honshu).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Mikura-jima, Okinawa-jima), Taiwan.

Rhyacophila shiraishiensis Kobayashi 1971

Rhyacophila shiraishiensis Kobayashi 1971a, 1-2, pl. 1, male, Honshu (Kanagawa).

Kobayashi 1971b, 19-20, pl. 11, male, Honshu (Kanagawa).
Kobayashi 1973, Honshu (Yamagata).
Kuranishi 1999, 186.
Kuranishi 2016, 67.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. This species is probably a junior synonym of R. bilobata (Hattori 2005, Kuranishi 2016).

Rhyacophila tachikawana Kobayashi 1973

Rhyacophila tachikawana Kobayashi 1973, 24-25, pl. 4, male, Honshu (Yamagata).

Kuranishi 1999, 189.
Kuranishi 2016, 67.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila towadensis Iwata 1927

Rhyacophila towadensis Iwata 1927a, 215-216, pl. 6, male, Honshu (Akita, Kyoto).

Iwata 1927b, 204, pl. 2, larva, Honshu (Akita, Kyoto).
Tanida 1985, 181, larva.
Moriya 1994, 34, 77, Honshu (Kanagawa, Yamanashi).
Kuranishi and Ohtaka 1996, 14, Honshu (Aomori).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 91, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 186.
Maruyama and Takai, 2000, 72, larva.
Kuranishi et al. 2002, 99, Honshu (Aomori).
Hattori 2005, 417, 421, 430, male, female, larva.
Katsuma 2005a, 3, Honshu (Ibaraki).
Nozaki 2005, 106, adult.
Ito et al. 2010, 53, 72, male, female, variation, Hokkaido (Hiyama, Oshima).
Inazu and Nishida 2011, 174, male, larva, Honshu (Hyogo).
Katsuma 2011b, 48, Honshu (Ibaraki).
Inazu 2012, 4, Honshu (Hyogo).
Katsuma 2012b, 48, Honshu (Ibaraki).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 20, 21, male, Honshu (Akita).
Tanaka 2014, 37, Honshu (Akita).
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 68.
Tanaka 2017, 2, Honshu (Akita).
Hattori 2018, 477, 481, 490, male, female, larva.

Rhyacophila vaoides Ross 1956, 82, 117, male, Honshu (Tochigi). Synonymized by Kuranishi (1999).

Hatta and Nozaki 1991, 202, Honshu (Gifu).

Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu).
Remarks. Tsuda (1948) synonymized this species with R. articulata. However, Kuranishi (1999) recognized that this species is a distinct species and R. vaoides is the same species as this species.

Rhyacophila transquilla Tsuda 1940

Rhyacophila transquilla Tsuda 1940, 131, male, Honshu (Kyoto).

Tsuda 1942a, 247, Honshu (Shiga).
Tsuda 1948, 14, larva, Honshu (Nagano, Tochigi).
Ross 1956, 95-96, 120, male, Japan (Nagano).
Tsuda 1959, 132, larva.
Tsuda and Akagi 1962, 126-128, larva.
Kuwada 1965, 163, pl. 82, male, Shikoku (Ehime).
Kobayashi 1971b, 14-15, pl. 6, male, female, Honshu (Kanagawa).
Tsuda and Akagi 1974, 126-128, larva.
Tani 1977, 197-198, male.
Tani 1978, 8, Honshu (Niigata).
Yamanouchi 1978, 9, Hokkaido (Tokachi).
Tanabe 1980, 65, adult, Hokkaido (Tokachi).
Tanida 1985, 186, larva.
Nozaki 1988a, 45, 46, Honshu (Kanagawa).
Nozaki 1988b, 40, Honshu (Kanagawa).
Morita 1989, 39, Honshu (Mie).
Kadowaki 1990, 14, Okino-shima.
Hatta and Nozaki 1991, 202, Honshu (Aichi, Gifu).
Tanida 1991, 34, Honshu (Nara).
Morita 1993a, 5, Honshu (Mie).
Nishimoto and Nishimoto 1993, 86, Honshu (Aichi).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 34, 77, Honshu (Kanagawa, Yamanashi).
Arefina 1997, 28, 32, male, female.
Ito et al. 1997, 23, 26, 28, 30, Hokkaido (Hiyama).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Nishiharu 1997, 59, Honshu (Fukui).
Nozaki 1997a, 32, Honshu (Kanagawa).
Tani and Nakamura 1997, 61, Honshu (Hiroshima).
Ito et al. 1998, 10, Hokkaido (Nemuro).
Kagaya et al. 1998, 21, 92, Honshu (Tokyo, Yamanashi).
Morita 1998b, 71, Honshu (Mie).
Sekimoto et al. 1998, 330, Honshu (Hiroshima).
Yodoe et al. 1998, 68, Honshu (Shimane).
Kuranishi 1999, 189.
Ito et al. 2000a, 26, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya).
Maruyama and Takai 2000, 72, 77, 78, larva, Honshu (Nara).
Arefina 2001, 28-29, 41, female, Sakhalin.
Kawase and Morita 2001a, 8, Honshu (Ishikawa).
Morita 2001a, 219, Honshu (Mie).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Hattori 2005, 420, 427, 434, male, female, larva.
Ishiwata et al. 2005, 110, 244-245, Honshu (Kanagawa).
Katsuma 2005b, 11, Honshu (Ibaraki).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Kuhara et al. 2007, ap. 67, Hokkaido (Ishikari).
Morita 2007b, 96, Honshu (Mie).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Ohkawa and Hattori 2007, 267, Hokkaido (Kamikawa).
Inazu 2008, 65, 86, 99, Honshu (Hyogo).
Muramatsu 2008, 3, Hokkaido (Okhotsk).
Tanida 2008, 248, pl. 92, adult, Shikoku (Ehime).
Aoyagi and Iwasaki 2009, 101, 103, Honshu (Osaka).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Morita 2010, 36, Honshu (Shiga, Mie).
Inazu and Nishida 2011, 175, male, larva, Honshu (Hyogo).
Ito and Kubo 2011, 5, Hokkaido (Tokachi).
Kuhara 2011, 52, 56, 57, 62, Hokkaido (Sorachi, Ishikari).
Taira and Tanida 2011, 29-34, habitat, Honshu (Nara).
Inazu 2012, 4, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Kawase and Hayashi 2012, 255, Honshu (Shimane).
Morita and Kawase 2012, 14, 15, Honshu (Ishikawa).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 21, 23, male, Honshu (Akita).
Tanaka 2012b, 11, Honshu (Akita).
Ito 2013, 49, Hokkaido (Kushiro).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Iijima and Tsuchiya 2014, 8, Hokkaido Sorachi).
Ito and Nagasaka 2014, 8, Hokkaido (Sorachi).
Kuranishi and Kikuchi 2014, 362, Honshu (Yamagata).
Nozaki et al. 2014, 59, Honshu (Nagano).
Taira and Tanida 2014, behavior, Honshu (Nara).
Tanaka 2014, 38, Honshu (Akita).
Hayashi et al. 2016, 29, Okino-shima.
Horiguchi 2016, 409, Honshu (Tokyo).
Kuranishi 2016, 68.
Nozaki 2016, 70, 312, 313, 429, 449, male, Shikoku (Kochi).
Ito 2017b, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 480, 487, 494, male, female, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Okino-shima), Russia.
Remarks. Tsuda (1940) used two names, R. transquila and R. transquilla, in the original description of this species, but he used the latter name for the first time (Tsuda 1942).

Rhyacophila tsusimaensis Kobayashi 1985

Rhyacophila tsusimaensis Kobayashi 1985a, 13-14, male, Tsushima.

Kuranishi 1999, 187.
Hattori 2005, 429, 436, male.
Kuranishi 2016, 68.
Hattori 2018, 489, 496, male.

Type locality. Japan (Tsushima).
Distribution. Japan (Tsushima).

Rhyacophila ulmeri Navás 1907

Rhyacophila ulmeri Navás 1907, 309-400, male, China.

Ross 1956, 92-93, 119, male, China.
Kobayashi 1973, Honshu (Yamagata).
Tani 1978, 8, Honshu (Niigata).
Kuranishi 1999, 189.
Hattori 2005, 420, 428, 436, male, larva.
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2015, 54-55, Honshu (Akita).
Kuranishi 2016, 68.
Hattori 2018, 480, 488, 496, male, larva.

Type locality. China.
Distribution. Japan (Honshu), China.

Rhyacophila verecunda Tsuda 1940

Rhyacophila verecunda Tsuda 1940, 120-121, male, Honshu (Kyoto).

Kuranishi 1999, 186.
Hattori. 2005, 425, 434, male.
Inazu 2008, 65, 86, 99, Honshu (Hyogo).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Inazu and Nishida 2011, 175, male, Honshu (Hyogo).
Kuranishi 2016, 68.
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 485, 494, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila yamanakensis Iwata 1927

Rhyacophila yamanakensis Iwata 1927a, 217, pl. 6, larva, Honshu (Ishikawa, Tottori).

Tsuda and Kuwayama 1950, 414, male.
Kobayashi 1959, 343, Kyushu (Fukuoka).
Tsuda 1959, 13, larva.
Tsuda and Akagi 1962, 124-125, 127, larva.
Lepneva 1964, 251-256, larva.
Schmid 1970, 91, 132, pl. 42, male.
Tani 1977, 198, male.
Tanida 1985, 181, larva.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Kobayashi 1987a, 20, Honshu (Shimane).
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 40, Honshu (Kanagawa).
Kadowaki 1990, 14, Okino-shima.
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Tanida 1990, 55, Honshu (Hyogo).
Hatta and Nozaki 1991, 202, Honshu (Gifu).
Tanida 1991, 34, Honshu (Nara).
Moriya 1992, 15, 42, Honshu (Kanagawa).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Tanida and Takemon 1993, 242, Honshu (Kyoto).
Moriya 1994, 34, 77, Honshu (Kanagawa).
Tanida et al. 1995, 3, Honshu (Nara).
Ito et al. 1997, 23, 31, Hokkaido (Hiyama).
Moriya 1997, 30, 66, Honshu (Kanagawa, Yamanashi).
Nishiharu 1997, 59, Honshu (Fukui).
Nozaki 1997a, 32, Honshu (Kanagawa).
Nozaki 1997b, 27, 32, Honshu (Tokyo).
Tani and Nakamura 1997, 62, Honshu (Hiroshima).
Kadowaki 1998, 5, Honshu (Shimane).
Kagaya et al. 1998, 21, 93, Honshi (Tokyo, Yamanashi).
Sekimoto et al. 1998, 330, Honshu (Hiroshima).
Ito et al. 1999, 23, 25-26, Hokkaido (Kushiro).
Kuranishi 1999, 187.
Ito et al. 2000a, 26, Hokkaido (Ishikari).
Ito et al. 2000b, 26, 28-29, Hokkaido (Soya).
Kuhara et al. 2000, 247, Hokkaido (Iburi).
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Maruyama and Takai, 2000, 72, 76, 77, larva.
Kuhara 2001, 17, Hokkaido (Shiribeshi).
Morita 2001a, 219, Honshu (Mie).
Morita 2001b, 336, Honshu (Mie).
Kuranishi et al. 2002, 99, Honshu (Aomori).
Nozaki and Nakamura 2002, 166, Honshu (Hiroshima).
Nakamura et al. 2003, 152, Honshu (Hiroshima).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Kawase et al. 2004, 107, Honshu (Ibaraki).
Uenishi and Yoshitaka 2004, 15, Honshu (Okayama).
Yoshitaka 2004, 27, Honshu (Okayama).
Hattori 2005, 417, 422, 430, male, female, larva.
Hirabayashi et al. 2005, Honshu (Nagano).
Ishiwata et al. 2005, 110, 244-245, Honshu (Kanagawa).
Katsuma 2005b, 11, Honshu (Gumma).
Nozaki and Nakamura 2005, 218, Honshu (Hiroshima).
Kameyama and Nakamura 2006, 92, Honshu (Hiroshima).
Kimura et al. 2006, 7, Honshu (Nagano).
Mey and Nozaki 2006, 24, Honshu (Shiga).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Nakamura et al. 2006, 100, Honshu (Hiroshima).
Katsuma 2007, 17, Honshu (Tochigi).
Nakamura and Tsujimura 2007, 76, Honshu (Hiroshima).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Nakamura 2007, 92, Honshu (Hiroshima).
Inazu 2008, 65, 86, 99, Honshu (Hyogo).
Kimura et al. 2008, 264, Honshu (Nagano).
Kameyama et al. 2009, 110, Honshu (Shimane, Hiroshima).
Ito et al. 2010, 53, 72, Hokkaido (Shiribeshi, Hiyama, Oshima).
Katsuma 2010, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 36, Honshu (Shiga, Mie).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 238, Honshu (Hiroshima).
Tanaka 2010a, 23, 28, male, Honshu (Akita).
Tanaka 2010b, 16, Honshu (Akita).
Nozaki 2010, 215, Honshu (Akita).
Inazu and Nishida 2011, 175, male, larva, Honshu (Hyogo).
Morita 2011, 214, Honshu (Mie).
Taira and Tanida 2011, 31, habitat, Honshu (Nara).
Tsukiji 2011, 140, adult.
Inazu 2012, 4, Honshu (Hyogo).
Katsuma 2012b, 63, Honshu (Ibaraki).
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Tanaka 2012a, 20, 21, male, Honshu (Akita).
Tanaka 2012b, 11, Honshu (Akita).
Ito 2013, 49, Hokkaido (Okhotsk, Kushiro).
Iijima and Tsuchiya 2014, 27, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 9, Hokkaido (Sorachi).
Kawase and Morita. 2014, 3, Honshu (Shiga).
Tanaka 2014, 37, Honshu (Akita).
Hayashi et al. 2016, 28, Okino-shima.
Hirabayashi et al. 2016, 208, Honshu (Nagano).
Horiguchi 2016, 409, (Tokyo).
Kuranishi 2016, 68.
Nozaki 2016, 69, 309, 313, 429, male, Honshu (Fukushima).
Ito 2017b, 39, Hokkaido (Kamikawa).
Kobayashi et al. 2017, 17, 18, 19, Honshu (Kyoto).
Kawase et al., 2017, 24, Honshu (Shiga).
Nojima 2017, 110, Honshu (Okayama).
Tanaka 2017, 2, Honshu (Akita).
Saito et al. 2017, 183, Honshu (Chiba).
Hattori 2018, 477, 482, 490 male, female, larva.
Morita 2019, 22, Honshu (Mie).
Nozaki et al. 2019, 167, Honshu (Nagano).

Rhyacophila quieta Tsuda 1940, 126-127, male, Honshu (Kyoto). Synonymized by Tsuda (1948).

Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942, 246-247, Honshu (Kyoto, Nagano).
Ross 1956, 99, 121, male.
Kuwada 1965, 163, pl. 82, male, Shikoku (Ehime).
Kobayashi 1983, 32-36, Honshu (Kanagawa).
Kadowaki 1998, 5, Honshu (Shimane).

Type locality. Japan (Honshu).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Okino-shima).

Rhyacophila yoshinensis Tsuda and Kawai 1967

Rhyacophila yoshinensis Tsuda and Kawai 1967, 111-112, male, Honshu (Nara).

Tani 1977, 198, male.
Kuranishi 1999, 187.
Kuranishi and Yamamoto 2000, 61, Shikoku (Ehime).
Hattori 2005, 422, 430-431, male, female.
Morita 2006a, 47, Honshu (Mie).
Inazu and Nishida 2011, 175, male, larva, Honshu (Hyogo).
Katsuma 2012, 53, Honshu (Ibaraki).
Morita and Kawase 2012, 15, Honshu (Ishikwawa).
Tanaka 2012b, 11, Honshu (Akita).
Kawase and Morita 2014, 3-4, Honshu (Shiga).
Kuranishi 2016, 68. Honshu.
Nozaki 2016, 69, 310, 313, 429, 449, male, Shikoku (Kochi).
Nojima 2017, 110, Honshu (Okayama).
Hattori 2018, 482, 491, male, female.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu, Shikoku).

Rhyacophila yosiiana Tsuda 1940

Rhyacophila tristis: Iwata 1927a, 216, pl. 6, larva, Honshu (Shiga, Kyoto). Misidentification mentioned by Tsuda (1948).
Rhyacophila yosiiana Tsuda 1940, 119-1202, male, Honshu (Nagano, Kyoto).

Tsuda 1941, 104-105, 107, Honshu (Nagano).
Tsuda 1942, 245, Honshu (Shiga).
Tsuda and Kuwayama 1950, 414, male.
Ross 1956, 76, 92-93, male.
Tani 1977, 198, male.
Kugo and Nozaki 1990, 62, Honshu (Hyogo).
Hatta and Nozaki 1991, 202, Honshu (Aichi).
Tanida 1991, 34, Honshu (Nara).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Nishiharu 1997, 59, Honshu (Fukui).
Nozaki 1997a, 32, Honshu (Kanagawa).
Kagaya et al. 1998, 21, 94, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 190.
Kuranishi and Yamamoto 2000, 62, Shikoku (Ehime).
Uenishi and Kurumi 2000, 278, Honshu (Shiga).
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Morita 2001b, 336, Honshu (Mie).
Morita 2003, 23-24, Honshu (Mie).
Hattori 2005, 416, 417, 423, 432, male, female, larva.
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Morita 2006b, 199, Honshu (Mie).
Torii and Hattori 2006, 33, Honshu (Shizuoka).
Inazu 2008, 65, 86, 99, Honshu (Hyogo).
Morita 2008b, 91, Honshu (Mie).
Aoyagi and Iwasaki 2009, 101, 105, Honshu (Osaka).
Morita 2009a, 137, Honshu (Mie).
Morita 2009c, 7, Honshu (Mie).
Aoyagi and Iwasaki 2010, 32, 33, 35, 36, Honshu (Osaka).
Morita and Kawase 2010, 27, Honshu (Ishikawa).
Morita 2011, 214, Honshu (Mie).
Inazu 2012, 4, Honshu (Hyogo).
Morita and Kawase 2012, 15, Honshu (Ishikawa).
Tanaka 2012a, 20, 22, male, Honshu (Akita).
Kuranishi 2016, 68. Honshu.
Morita and Kawase 2016, 23, Honshu (Toyama).
Kawase et al. 2017, 24, Honshu (Shiga).
Nojima 2017, 110-111, Honshu (Okayama).
Hattori 2018, 476, 477, 483, 492, male, female, larva.

Rhyacophila sp. RD: Tsuda 1948, 12-13, larva, Honshu (Nagano, Shiga, Kyoto).

Tsuda and Akagi 1962, 126-128, larva.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).
Remarks. Tsuda (1948) pointed out that a larva recorded as a European species, R. tristris, by Iwata (1927a) was the same species as Rhyacophila sp. RD. Later, Hattori (2005) treated R. sp. RD as the larval stage of R. yosiiana.

Rhyacophila yukii Tsuda 1942

Rhyacophila yukii Tsuda 1942, 248-249, male, Honshu (Shiga).

Kagaya et al. 1998, 21, 95, Honshu (Tokyo, Yamanashi).
Kuranishi 1999, 189.
Kawase and Morita 2001, 8, Honshu (Ishikawa).
Hattori 2005, 427, 435, male, Honshu.
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Inazu 2008, 65, 86, 99, Honshu (Hyogo).
Morita 2009c, 7, Honshu (Shiga).
Kawase and Hayashi 2010b, 84, Honshu (Shimane).
Inazu and Nishida 2011, 176, male, Honshu (Hyogo).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Kuranishi 2016, 68.
Kawase et al. 2017, 25, Honshu (Shiga).
Nojima 2017, 111, Honshu (Okayama).
Hattori 2018, 487, 495, male.

Type locality. Japan (Honshu).
Distribution. Japan (Honshu).

Rhyacophila sp. (yukii sensu Schmid 1970)

Rhyacophila yukii: Schmid 1970, 126, pl. 15, male. Misidentification mentioned by Hattori (2005).
Rhyacophila sp. (yukii sensu Schmid 1970)

Hattori 2005, 427, 435, male.
Inazu and Nishida 2011, 176, male, Honshu (Hyogo).
Nojima 2017, 111, Honshu (Okayama).
Hattori 2018, 487, 495, male.

Distribution. Japan (Honshu).
Remarks. Schmid (1970) described a male as R. yukii, but Hattori (2005) pointed out that it does not belong to R. yukii described by Tsuda (1942).

Rhyacophila sp.

Rhyacophila sp.: Tanaka 2012a, 20, 22, male, Honshu (Akita).
Remarks. This species is very similar to R. yamanakensis, but distinguishable from the latter by male genitalia (Tanaka 2012a).

Rhyacophila sp.

Rhyacophila sp.: Tanaka 2012a, 20-21, 22, male, Honshu (Akata).
Remarks. This species is very similar to R. motakanta, but distinguishable from the latter by male genitalia (Tanaka 2012a).

Rhyacophila sp. RA

Rhyacophila sp. RA: Tsuda 1948, 6-7, larva, Honshu (Kyoto, Nagano, Tochigi).

Tsuda and Akagi 1962, 124-125, 127, larva.

Remarks. Tanida (1985) pointed out that this species was described probably based on an early instar larva.

Rhyacophila sp. RB

Rhyacophila sp. RB: Tsuda 1948, 9-10, larva, Honshu (Kyoto).

Tsuda and Akagi 1962, 124-125, 127, larva.
Tanida 1985, 183, 186, larva.
Hattori 2005, 416, 417, larva.
Hattori 2018, 476, 477, larva.

Rhyacophila sp. RC

Rhyacophila sp. RC: Tsuda 1948, 10, larva, Honshu (Tochigi, Kyoto, Wakayama).

Tsuda and Akagi 1962, 124-125, 127, larva.
Tanida 1985, 181, 183, larva.
Taira and Tanida 2011, 25-35, behavior, Honshu (Nara).
Taira and Tanida 2014, 23-37, behavior, Honshu (Nara).

Remarks. Two or more species belong to this 'species' (Hattori 2005).

Rhyacophila sp. RK

Rhyacophila sp. RK: Tanida 1985, 181, 182, larva, Honshu.

Maruyama and Takai 2000, 75, 76, larva, Shikoku (Kochi).

Rhyacophila sp. RL

Rhyacophila sp. RL: Tsuda 1948, 185, 186, larva, Honshu.

Tsuda and Akagi 1962, 126-128, larva.

Rhyacophila sp. RM?

Rhyacophila sp. aff. nigrocephala: Tanida 1985, 185.
Rhyacophila sp. RM?: Hattori 2005, 416, 419, larva, Honshu.

Torii and Hattori 2006, 33, habitat, Honshu (Shizuoka).
Hattori 2018, 478-479, larva.

Rhyacophila sp. (Kaltatica group-sp. 1)

Rhyacophila sp. (Kaltatica group): Hattori 2005, 418, 419, 435, larva, Hokkaido (Ishikari).

Hattori 2018, 478, 479, 495, larva, Hokkaido (Ishikari).

Remarks. This is the larval stage of an undescribed species related to Rhyacophila azumaensis (Hattori 2005).

Rhyacophila sp. (Sibirica group-sp. 1)

Rhyacophila sp. (sibirica group-sp. 1): Hattori 2005, 418, 420, 435, larva, Honshu (Shizuoka, Nara).

Hattori 2018, 480, 483, 495, larva, Honshu (Shizuoka, Nara).

Remarks. This is the larval stage of an undescribed species belonging to the Sibirica species group (Hattori, 2005). Hattori (2005) also recognized another undescribed species as Rhyacophila sp. (Sibirica group-sp. 2) from Gifu, Honshu.

Rhyacophila sp. (Betteni group?)

Rhyacophila sp. (Betteni group?): Hattori 2005, 420, 437, larva, Hokkaido (South-east).

Hattori 2018, 480, 497, larva, Hokkaido (South-east).

Remarks. This is the larval stage of an undescribed species probably belonging to the Betteni species group (Hattori, 2005).

Rhyacophila sp. (species group unknown)

Rhyacophila sp. (species group?): Hattori 2005, 420, 424, 437, larva.

Hattori 2018, 480-497, larva.

Rhyacophila sp. (species group unknown)

Rhyacophila sp.: Satake and Kuranishi 2007, 281-282, larva, habitat, Haha-jima.

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