A catalogue of Japanese Trichoptera
Family Uenoidae Iwata


Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last update: 18 Dec. 2016


Subfamily Thremmatidae Martynov

Only the genus Neophylax is distributed in Japan.

Genus Neophylax McLachlan

Four named species belonging to this genus have been recorded from Japan. Larvae of two unnamed species separated with only tentative alphabetic designation have not been assosiated with their adult stage.

Neophylax japonicus Schmid 1964

Neophylax japonicus Schmid 1964, 832-834, male, female, Hokkaido (Ishikari), Honshu (Nagano).
Kobayashi 1984a, 7-8, female, Honshu (Kanagawa).
Tanida 1985, 199, larva, case.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Kanazawa 1987, 1-4, male, female, larva, case, parasitoid.
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 42, Honshu (Kanagawa).
Gyotoku and Nozaki 1992, 138-139, Kyushu (Fukuoka).
Konishi and Aoyagi 1994, 421-431, parasitoid.
Moriya 1994, 80, Honshu (Yamanashi).
Ito et al. 1997, 24, 27, Hokkaido (Akan).
Moriya 1997, 69, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 14, 18, 19, Hokkaido (Oshima).
Nozaki 1997a, 37, Honshu (Kanagawa).
Itou and Hirama 1998, 40, Hokkaido (Kushiro).
Ito et al. 1998, 11-15, Hokkaido (Nemuro).
Kagaya et al. 1998, 29, 62, 205, Honshu (Tokyo, Yamanashi).
Ito et al. 2000a, 35, Hokkaido (Ishikari).
Nozaki et al. 2000, 199.
Kuhara 2001, 18, Hokkaido (Shiribeshi).
Ishiwata et al. 2005, app. 2(3), Honshu (Kanagawa).
Nozaki 2005a, 532-534, male, female, larva, case.
Vineyard et al. 2005, 31-32, 100, male, female, larva, Hokkaido (Tokachi, Ishikari), Honshu (Fukushima, Kanagawa, Nagano, Hiroshima).
Kimura et al. 2006, 8, Honshu (Shiga).
Kuhara et al. 2006, 45, 47, Hokkaido (Ishikari).
Morita 2008, 719, 723, Honshu (Mie).
Inazu 2008, 82, 95, 103, Honshu (Hyogo).
Muramatsu 2008, 11, Hokkaido (Okhotsk).
Morita and Kawase 2008, 56, Honshu (Ishikawa).
Tanida 2008, 253, pl. 94, male, Honshu (Nagano).
Ito 2010, 131, Hokkaido (Kushiro).
Ito et al. 2010, 64, 77, Hokkaido (Shiribeshi, Oshima).
Tanaka 2010a, 25-28, male, Honshu (Akita).
Tanaka 2010b, 17, Honshu (Akita).
Inazu and Nishida 2011, 205, male, female, larva, case, Honshu (Hyogo).
Ito and Kubo 2011, 9, Hokkaido (Tokachi).
Kuhara 2011, 53, 64, 75, Hokkaido (Sorachi, Ishikari, Hidaka).
Inazu 2012, 10, Honshu (Hyogo).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 67, Hokkaido (Kushiro).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Iijima and Tsuchiya 2014, 33, Hokkaido (Okhotsk, Kushiro).
Ito and Nagasaka 2014, Hokkaido (Sorachi).
Nozaki et al. 2014, 33, Honshu (Nagano).
Nozaki 2016, 302, 387, 440, forewing.
Tanida 2016, 123.
Neophylax muinensis Kobayashi 1977, 2-3, 10, male, female, Hokkaido (Tokachi, Ishikari). Synonymized by Vineyard et al. (2005).
Yamanouchi 1978, 11-13, larva, case, Hokkaido (Tokachi).
Nozaki et al. 2000, 199.
Type locality. Hokkaido (Ishikari), Japan.
Distribution. Japan (Hokkaido, Honshu, Kyushu).

Neophylax koizumii (Iwata 1927)

Apatania koizumii Iwata 1927c, 390-391, 394, Honshu (Nagano).
Neophylax sp. NA: Tanida 1985, 199, larva, case. Misidentification mentioned by Mitsuhashi et al. (2000).
Konishi and Aoyagi 1994, 421-431, prasitoid.
Kagaya et al. 1998, 29, 62-63, 206, Honshu (Tokyo, Yamanashi).
Nishimoto 2000, 26-28, larva, adult, egg mass, habitat, behavior, Honshu (Aichi).
Neophylax koizumii: Mitsuhashi et al. 2000, 139-142, larva, case, Honshu (Aomori, Iwate, Fukushima, Tokyo, Kanagawa, Nagano, Aichi).
Vineyard et al. 2005, 22-23, 93, male, female, larva, Honshu (Fukushima, Tokyo).
Nozaki 2005a, 532-534, larva, case.
Nishimoto 2008, 8-11, adult, larva, life history, behavior, Honshu (Aichi).
Nishimoto 2014, 59, larva, Honshu (Aichi).
Taira et al. 2014, 173-177, larva, case, habitat, distribution, Honshu (Wakayama).
Nozaki 2016, 387, 440, forewing.
Tanida 2016, 123.
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu).

Neophylax shikoku Vineyard and Wiggins 2005

Neophylax shikoku: Vineyard et al. 2005, 32-33, 101, male, Shikoku (Tokushima).
Nozaki 2016, 82, 387, 440, 451, adult, Shikoku (Kochi).
Tanida 2016, 123.
Type locality. Shikoku (Tokushima), Japan.
Distribution. Japan (Shikoku).

Neophylax ussuriensis (Martynov 1914)

Halesus ussuriensis Martynov 1914, 276-278, female, Russia.
Martynov, 1935, 310-315, male, Russia.
Halesus uenoi Tsuda 1942, 319-320, male, Hokkaido (Tokachi, Ishikari). Synonymized by Schmid (1955).
Fischer 1967, 142.
Neophylax ussuriensis: Schmid 1955, 93-96, male, female (?).
Kobayashi 1964, 88, Hokkaido (Tokachi, Kamikawa).
Lepneva 1966, 80-86, larva, pupa, case, Russia.
Fischer 1967, 143.
Fischer 1973, 37-38.
Kobayashi 1977, 2, Hokkaido (Tokachi, Ishikari).
Kobayashi 1984b, 19, 31, male, female, Hokkaido (Shiribeshi, Hiyama, Oshima).
Tanida 1985, 199, larva, case.
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Kuhara and Ito 1994, Hokkaido (Ishikari).
Arefina 1997, 130, male, Russia.
Ito et al. 1997, 14, 18, 19, Hokkaido (Akan).
Kuhara 1997a, 61, Hokkaido (Shiribeshi).
Kuhara 1997b, 96, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 24, 28, 31, Hokkaido (Oshima).
Ito et al. 1998, 11-15, Hokkaido (Nemuro).
Ito et al. 1999, 24, 25, 27, Hokkaido (Akan).
Kushiro Insect Lovers' Society 1999, 218, Hokkaido (Nemuro).
Ohkawa et al. 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 36, Hokkaido (Ishikari).
Ito et al. 2000b, 27-30, Hokkaido (Soya).
Kuhara et al. 2000, 254, Hokkaido (Iburi).
Nozaki et al. 2000, 199.
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Kuhara and Sato 2001, 65, Hokkaido (Rishiri).
Kuranishi and Satake 2004, 39, Hokkaido (Okhotsk).
Vineyard et al. 2005, 16-18, 83, male, female, larva, Hokkaido (Nemuro, Kamikawa, Ishikari).
Nozaki 2005a, 532-534, male, female, larva, case.
Kuhara et al. 2007, ap. 72, Hokkaido (Ishikari).
Nakano et al. 2007, 41, Hokkaido (Nemuro).
Muramatsu 2008, 11, Hokkaido (Okhotsk).
Ito 2010, 131, Hokkaido (Kushiro).
Ito et al. 2010, 64, 77, Hokkaido (Shiribeshi, Hiyama, Oshima).
Tsuji et al. 2010, Hokkaido (Okhotsk).
Ito and Kubo 2011, 9, Hokkaido (Tokachi).
Kuhara 2011, 53, 75, Hokkaido (Sorachi, Ishikari).
Ito 2013, 67, Hokkaido (Nemuro, Okhotsk, Kushiro).
Iijima and Tsuchiya 2014, 33, Hokkaido (Okhotsk, Kushiro, Tokachi).
Ito and Nagasaka 2014, 12, Hokkaido (Sorachi).
Nozaki 2016, 387-388, forewing.
Tanida 2016, 123.
Neophylax sp. NB: Tani 1967, 12-13, larva, case. Associated with the adult by Tani (1980).
Type locality. Russia (Ussuri).
Distribution. Japan (Hokkaido, Honshu?), Russia (Altai, Sayans, Amur, Maritime territories, Sakhalin, Kuriles), Korea.
Remarks. Nozaki et al. (2000) suggested that illustrations of female genitalia as this species by Schmid (1955) are based on a misidentification of another species. Konishi and Aoyagi (1994) recorded this species from Honshu as a host of a parasite wasp based on identification of larval cases.

Neophylax sp. NA

Neophylax sp. NA: Akagi 1962, 41, larva, case, Honshu (Nara, Okayama).
Tsuda and Akagi 1962, 138, larva.
Mitsuhashi et al. 2000, 142, larva.
Nozaki et al. 2000, 199.
Remarks. Mitsuhashi et al. (2000) suggested that this larva is different from that of N. koizumii at species level.

Neophylax sp. NC

Neophylax sp. NC: Tanaka 1970, 28, larva, case, Honshu (Aomori).
Kim 1974, 67-68, larva, male genitalia, Honshu (Nara).
Nozaki et al. 2000, 199.
Remarks. The description of the larva and male genitalia of this species is different from other Japanese species (Nozaki et al 2000).

Subfamily Uenoinae Iwata

Only the genus Uenoa is known from Japan.

Genus Uenoa Iwata

The single species Uenoa tokunagai is known from Japan.


Uenoa tokunagai Iwata 1927

Uenoa tokunagai Iwata 1927a, 257, pl. 11, larva, case, Honshu (Shiga, Kyoto), Kyushu (Kagoshima).
Iwata 1927b, 214, larva, case, Honshu (Kyoto), Kyushu (Kagoshima).
Chihara 1956, 85-86, pupa, case.
Tsuda 1959, 153, larva, case.
Tsuda and Akagi 1962, 142, 146, pupa, case.
Fischer 1970, 286-287
Koyama 1980, 19-26, case, silk gland.
Tanida 1982, 18-19, 28, Honshu (Ishikawa).
Tanida 1985, 195, larva, case.
Wiggins et al. 1985, 774, male, female, larva, pupa, case.
Kagaya et al. 1998, 29, 63, 207, Honshu (Tokyo, Yamanashi).
Koyama 1990, 35, emergence behaivior, Honshu (Nara).
Tanida 1991, 35, Honshu (Nara).
Gyotoku and Nozaki 1992, 138, Kyushu (Fukuoka).
Morita 1993, 7, Honshu (Mie).
Moriya 1997, 69, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 37, Honshu (Kanagawa).
Nozaki 1997b, 30, 33, Honshu (Tokyo).
Maruyama and Takai 2000, 103, 109, larva, case, Honshu (Nara).
Nozaki et al. 2000, 199.
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Yamada et al. 2000, 267-272, adult, pupa, pupal aggregation, Honshu (Nara).
Morita 2003, 16, Honshu (Mie).
Aoyagi et al. 2004, 25, larva, Honshu (Kyoto).
Hosokawa et al. 2005, 181-186, case of 1st instar larva, life cycle, drift sift, food, Honshu (Nara).
Nozaki 2005a, 532-534, male, female, larva, case.
Nozaki 2005b, 112, adult, larva, case.
Mey and Nozaki 2006, 25, Honshu (Nagano).
Inazu 2008, 82-83, 95, 103, Honshu (Hyogo).
Aoyagi and Iwasaki 2009, 101, 104, 107, Honshu (Osaka).
Morita and Kawase 2009, 28, Honshu (Toyama).
Kawase & Hayashi 2010, 108, Honshu (Shimane).
Inazu and Nishida 2011, 205, male, female, larva, case, Honshu (Hyogo).
Kawase and Hayashi 2011, 192, Honshu (Shimane).
Katsuma 2012, 66, Honshu (Ibaraki).
Inazu 2012, Honshu (Hyogo).
Morita and Kawase 2012, 14, Honshu (Ishikawa).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Enju et al. 2013, 89, case.
Inazu 2013, 76, Honshu (Hyogo).
Kawase & Morita 2014, 6, Honshu (Shiga).
Nishimoto 2014, 59, larva, Honshu (Aichi).
Nozaki 2016, 81, 388, 440, 451, Honshu (Gumma).
Tanida 2016, 123.
Eothremma japonica Martynov 1933, 151-153, male, Honshu (Tochigi). Synonymized by Tsuda (1937).
Type locality. Japan.
Distribution. Japan (Honshu, Kyushu).

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