A catalogue of Japanese Trichoptera
Family Brachycentridae Ulmer

Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last updated: 31 Dec. 2023


Four genera belonging to this family have been recorded from Japan.

Genus Brachycentrus Curtis

Three species belonging to this genus is known from Japan.

Brachycentrus americanus (Banks 1899)

Oligoplectrum americanum Banks 1899, 210, male, USA.
Oligoplectrodes potanini Martynov 1910, 356-359, male, Russia, Mongolia. Synonymized by Schmid (1983)
Lepneva 1966, 381-387, larva, case.
Yamanouchi 1980, 8, Hokkaido (Tokachi).
Brachycentrus americanus: Ross 1944, 263-264, male, female, larva.
Fischer 1970, 100.
Wiggins 1977, 59, larva, case, USA.
Schmid 1983, 22-23, male, female.
Flint, 1984, 17-18, male, larva, USA, Canada.
Tanida 1985, 192, larva, case.
Tanida et al. 1991, 16-17, 88, adult, larva, pupa, case, drifting behavior of larva, upstream migration of adults.
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Kuranishi and Kuhara, 1994, 1227, Hokkaido (Kushiro).
Wiggins 1996, 213, larva, case, USA.
Arefina and Levanidova 1997, 91, male, female.
Kuhara 1997a, 61, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 18-19, Hokkaido (Oshima).
Wiggins and Parker 1997, 833-834, variation of tibial spur formula, USA, Russia.
Ito et al. 1998, 11-12, 14, Hokkaido (Nemuro).
Ito et al. 1999, 25-27, Hokkaido (Kushiro).
Nozaki et al. 1999, 353.
Ohkawa 1999, 41-44, Hokkaido (Tokachi).
Ito et al. 2000b, 30-31, Hokkaido (Ishikari).
Kuhara et al. 2000, 251, Hokkaido (Iburi).
Kuhara 2001, 18, Hokkaido (Shiribeshi).
Ohkawa and Ito 2001, 264-265, larva, food, Hokkaido (Ishikari).
Kuranishi and Satake 2004, 37, Hokkaido (Okhotsk).
Nozaki 2005a, 498-499, 501, larva, case, male.
Nozaki 2005b, 333-335, male, female, larva, Hokkaido (Nemuro, Kushiro, Okhotsk, Tokachi, Akan, Kamikawa, Ishikari).
Ito and Kosugi 2007, 54, Hokkaido (Kushiro).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Muramatsu 2008, 7, Hokkaido (Okhotsk).
Ito et al. 2009, 8, Hokkaido (Okhotsk).
Ito et al. 2010, 62, 75, Hokkaido (Hiyama, Oshima).
Tsuji et al., 2010, 6, Hokkaido (Okhotsk).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Kuhara 2011, 52, 73 Hokkaido (Hidaka).
Ito 2013, 59, Hokkaido (Nemuro, Okhotsk, Kushiro).
Ijima and Tsuchiya 2014, 30-31, Hokkaido (Okhotsk, Kushiro, Tokachi).
Nozaki 2016, 361, 436, male, female.
Tanida 2016, 102.
Nozaki 2018a, 591, 593-595, male, larva.
Ito 2020, 27-37, larva, life history, Hokkaido (Ishikari).
Muramatsu 2022, 29, male, female.
Ito and Kuhara 2023, 66, Hokkaido (Kamikawa).
Type locality. USA.
Distribution. Japan (Hokkaido), Sakhalin, Kurile, Russia (Siberia, Primorye, ), North America, Mongolia.
Remarks. This species belong the subgenus Oligoplectrodes.

Brachycentrus japonicus (Iwata 1927)

Brachycentriella japonica Iwata 1927a, 259, larva, case, Honshu (Nagano).
Iwata 1927b, 215, 218, Honshu (Nagano).
Fischer 1970, 96.
Brachycentrus japonicus: Flint 1984, 25, new combination.
Nozaki et al. 1999, 353.
Nozaki, 2005b, 329-331, male, female, larva, Honshu (Yamagata, Fukushima, Tochigi, Nagano).
Nozaki 2005c, 109, adult, larva.
Nozaki 2016, 78, 361-362, 436, 450, male, female, Honshu (Tochigi).
Tanida 2016, 102.
Nozaki 2018a, 591, 593-595, male, larva.
Egi 2022, 74, 83, 104, larva, Honshu (Okayama). Larval record.
Brachycentrus bilobatus Martynov 1935, 370-372, male, Russia. Synonymized by Nozaki (2005a).
Fischer 1970, 101.
Wiggins et al. 1985, 68-69, male, female, Honshu (Fukushima).
Arefina and Levanidova 1997, 91, male, female.
Nozaki et al. 1999, 353.
Nozaki, 2005a, 501, male.
Brachycentrus sp. BC: Akagi, 1962a, 43, larva, Honshu (Nagano). Associated by Wiggins et al. (1985) and Nozaki (2005a).
Nozaki 2005a, 499, larva.
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu), Russia (Primorye), North Korea, Mongolia.
Remarks. This species belongs to the subgenus Sphinctogaster. Nozaki (2005b) included Hokkaido as a distribution area of this species, but there is no reliable record.

Brachycentrus kuwayamai Wiggins, Tani and Tanida 1985

Brachycentrus subnubilus: Iwata 1927a, 258-259, pl. 10, larva (in part), Honshu (Gifu). Misidentification mentioned by Wiggins et al. (1985) and Nozaki (2005b).
Iwata 1927b, 214-215, pl. 6, larva, Honshu (Gifu).
Brachycentrus kuwayamai Wiggins et al. 1985, 70-71, male, Honshu (Fukushima).
Itou 1989, 55, Hokkaido (Kushiro).
Tashiro and Tashiro, 1989, 10, upstream migration of adults, Hokkaido (Kushiro).
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Itou and Muramatsu 1995, 40, Hokkaido (Okhotsk).
Nakajima et al. 1997, 18-19, Hokkaido (Oshima).
Ito et al. 1999, 25, Hokkaido (Kushiro).
Kuranishi and Kuhara 1999, 1227, Hokkaido (Kushiro).
Nozaki et al. 1999, 354.
Ito et al. 2000a, 28, Hokkaido (Soya).
Ito et al. 2000b, 31, Hokkaido (Ishikari).
Kuhara et al. 2000, 251, Hokkaido (Iburi).
Ito et al. 2004, 11, 15, Hokkaido (Kushiro).
Nozaki 2005a, 501, male.
Nozaki 2005b, 331-333, male, female, larva, Hokkaido (Kushiro, Akan, Ishikari), Honshu (Tochigi, Niigata, Gifu, Shiga, Kyoto).
Muramatsu 2008, 7, Hokkaido (Okhotsk).
Ito et al. 2010, 62, 75, Hokkaido (Oshima).
Ito 2010, 131, Hokkaido (Kushiro).
Kuhara 2011, 52, 73, Hokkaido (Sorachi).
Ito 2013, 59, Hokkaido (Okhotsk, Kushiro).
Kawase and Morita 2014, 6, Honshu (Shiga).
Ijima and Tsuchiya 2014, 31, Hokkaido (Okhotsk, Kushiro).
Ito and Nagasaka 2014, 11, Hokkaido (Sorachi).
Ito and Nagasaka 2015, case, Hokkaido (Iburi).
Nozaki 2016, 360-362, 436, male, female.
Tanaka 2016, 2, 4, male, Honshu (Akita).
Tanida 2016, 102.
Nojima 2017, 120, Honshu (Okayama).
Ito 2017b, 44, Hokkaido (Kamikawa).
Nozaki 2018a, 591, 593-595, male, lkarva, case.
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Tanaka 2018a, 21, Honshu (Akita).
Nakatani and Ito 2019, 163, Hokkaido (Kushiro).
Ito 2020, 27-37, larva, life history, Hokkaido (Iburi).
Muramatsu 2022, 29, male, female.
Tanaka 2022, 36, Honshu (Akita).
Ito and Kuhara, 2023, 66, Hokkaido (Soya, Kamikawa).
Brachycentrus sp. BA: Akagi 1957, 27, larva, case, Honshu (Nara). Associated by Nozaki (2005b).
Akagi 1962a, 43, larva.
Tsuda and Akagi 1962, 141, larva, case.
Tanida 1985, 192, larva, case.
Nozaki et al. 1999, 354
Nozaki 2005a, 499, larva.
Type locality. Honshu (Fukushima), Japan.
Distribution. Japan (Hokkaido, Honshu).
Remarks. This species belongs to the subgenus Brachycentrus. Wiggins et al. (1985) pointed out that the larval specimens recorded as B. subnubilus by Iwata (1927a) include at least two species, and that the record of B. subnubilus from Japan is doubtful. Nozaki (2005a) mentioned that a specimen collected from Gifu in Iwata’s specimens belongs to this species.

Genus Dolichocentrus Martynov

Dolichocentrus sakura Nozaki 2017

Dolichocentrus sakura Nozaki 2017, 555-561, male, female, larva, pupa, case, habitat, Honshu (Okayama).
Nojima 2017, 120, Honshu (Okayama).
Nozaki 2018a, 592, 593, 595, 596, male, larva, case.
Type locality. Honshu (Okayama), Japan.
Distribution. Japan (Honshu). Only known from the Asahi-gawa River in Okayama Prefecture.

Genus Eobrachycentrus Wiggins

Only two species belonging to this genus are recognized for Japanes fauna (Nozaki 2011).

Eobrachycentrus niigatai (Kobayashi 1968)

Brachycentrus niigatai Kobayashi 1968, 7, pl. 3, male, Honshu (Niigata).
Eobrachycentrus niigatai: Wiggins et al. 1985, 66, male.
Nozaki et al. 1999, 354.
Nozaki 2011, 394-396, 398-400, male, female, larva, pupa, case, habitat, food, Hokkaido (Rumoi, Kamikawa, Sorachi, Ishikrai, Hidaka, Shiribeshi, Hiyama), Honshu (Ibaraki, Saitama, Tokyo, Kanagawa, Yamanashi, Gifu, Shizuoka, Aichi, Mie, Shiga, Hyogo, Okayama), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka, Miyazaki).
Katsuma 2012, 65, Honshu (Ibaraki).
Inazu 2013c, 73, Honshu (Hyogo).
Ito and Nagasaka 2014, 11, Hokkaido (Sorachi).
Kawase and Morita 2014, Honshu (Shiga).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Yamamoto and Ito 2014, 13, 18-21, Shikoku (Ehime).
Nozaki 2016, 78, 360, 362-363, 436, male, Shikoku (Kochi).
Tanida 2016, 102.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Nozaki 2018a, 590, 591, 593, 595, male, larva, case.
Tsuruda 2018, 27, 34, adult, Honshu (Tokyo).
Eobrachycentrus propinquus Wiggins, Tani and Tanida 1985, 64-65, male, larva, Honshu (Mie, Osaka). Synonymized by Nozaki (2011).
Ito et al. 1997, 22, 24, 26, 28, 33, male, Hokkaido (Hiyama).
Kagaya et al. 1998, 27, 56, 179, Honshu (Tokyo).
Morita 2000, 60-61, male, Honshu (Mie).
Uenishi and Kurumi 2000, 279, Honshu (Shiga).
Kawase and Morita 2001, 9, Honshu (Ishikawa).
Kuhara 2001, 18, Hokkaido (Shiribeshi).
Morita 2001b, 339, Honshu (Mie).
Nozaki 2001, 15-16, larva, case, Honshu (Kanagawa).
Kuranishi 2002, 568, adult, habitat, Honshu (Chiba).
Uchida 2002, 78, Honshu (Tochigi).
Nozaki 2004, 1235, Honshu (Kanagawa).
Nozaki 2005a, 109, male.
Katsuma 2006, 38, 41, Honshu (Ibaraki).
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Morita 2008a, 723, Honshu (Mie).
Morita 2009a, 6-7, Honshu (Shiga).
Morita 2009b, 140, Honshu (Mie).
Kawase and Morita 2010, 43, Honshu (Mie, Shiga).
Inazu and Nishida 2011, 196, male, female, Honshu (Hyogo).
Kuhara 2011, 52, 57, 62, Hokkaido (Sorachi, Ishikari).
Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).

Eobrachycentrus vernalis (Banks 1906)

Brachycentrus vernalis Banks 1906, 108, male, Kyushu (Fukuoka).
Fischer 1970, 121.
Micrasema oharensis Iwata 1927c, 391-392, larva, case, Honshu (Kyoto). Synonymized by Nozaki (2011).
Brachycentrus sp. BB: Akagi 1959, 43, larva, Honshu (Toyama, Nara). Mentioned by Nozaki (2011).
Akagi 1962a, 43, larva.
Tsuda and Akagi 1962, 141, larva.
Kim 1974, 68-70, male, pupa, Honshu (Nara). Mentioned by Wiggins et al. (1985).
Micrasema oharense: Fischer 1970, 144.
Eobrachycentrus vernalis: Wiggins et al. 1985, 61-64, male, female, pupa, variant of male genitalia, habitat, life cycle, Honshu (Kyoto, Nara), Kyushu (Fukuoka).
Tanida 1985, 192, larva, case.
Tanida 1990, 56, Honshu (Hyogo).
Gyotoku and Nozaki 1992, 134, Kyushu (Fukuoka).
Moriya 1997, 67, Honshu (Kanagawa).
Kagaya et al. 1998, 27, 56, 180, Honshu (Tokyo, Yamanashi).
Morita 1998, 46, Honshu (Mie).
Nozaki et al. 1999, 355.
Morita 2001b, 339, Honshu (Mie).
Nozaki 2005a, 501, male.
Morita 2007, 14, Honshu (Mie)
Morita 2008, 6, Honshu (Mie).
Inazu 2008, 78, 93, 102, Honshu (Hyogo).
Morita 2009a, 6-7, Honshu (Shiga).
Morita 2009b, 140, Honshu (Mie).
Kawase and Morita 2010, Honshu (Mie).
Inazu and Nishida 2010, 196, male, female, Honshu (Hyogo).
Nozaki 2011, 391-393, 396, 398-400, male, female, larva, pupa, case, habitat, food, Honshu (Saitama, Tokyo, Ishikwa, Fukui, Yamanashi, Nagano, Shizuoka, Shiga, Kyoto, Mie, Hyogo, Nara), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka).
Inazu 2012, 8, Honshu (Hyogo).
Inazu 2013c, 73-74, Honshu (Hyogo).
Yamamoto and Ito 2014, 13-14, 18-21, Shikoku (Ehime).
Nozaki 2016, 78, 362-363, 436, 450, male, Shikoku (Kochi).
Tanida 2016, 102-103.
Nozaki 2018a, 590, 591, 593, 596, male, larva, case.
Yamamoto et al. 2021, 74, Shikoku (Ehime).
Nojima 2022, 38, Honshu (Okayama).
Eobrachycentrus oharensis: Wiggins et al. 1985, 67, Honshu (Kyoto).
Eobrachycentrus sp.: Nozaki 2005a, 499, larva, case.
Type locality. Kyushu (Fukuoka), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. Eobrachycentrus sp. described by Nozaki (2005a) is this species, since illustrations of the larva and case were reprinted from E. vernalis (Tanida 1985).

Genus Micrasema McLachlan

Six species belonging to this genus is known from Japan, but some taxonomic problems remain (Nozaki et al. 1999).

Micrasema akagiae Nozaki and Tanida 2007

Micrasema sp. MB: Akagi, 1959, 42-43, larva, male, Honshu (Aichi, Kyoto, Nara).
Tsuda and Akagi 1962, 141, larva.
Kagaya et al. 1998, 28, 57, 183, Honshu (Tokyo).
Nozaki et al. 1999, 356.
Micrasema sp.: Tanida et al., 1991, 92-93, adult, larva, pupa, Honshu (Shizuoka). Mentioned by Nozaki and Tanida (2007).
Micrasema akagiae Nozaki and Tanida 2007, 249-250, male, female, Honshu (Yamagata, Tochigi, Tokyo, Nagano, Shizuoka).
Tanida and Nozaki 2010, 89, adult, Honshu (Shizuoka).
Nozaki 2016, 363-364, 436, 450, male.
Tanaka 2016, 2, 5, male, Honshu (Akita).
Tanida 2016, 103.
Nojima 120, Honshu (Okayama).
Nozaki 2018, 592, 593, 595, 596, male, larva, case.
Tanaka 2018a, 9, Honshu (Akita).
Nozaki et al. 2019, 169, Honshu (Nagano).
Kuhara et al. 2023, 19, Honshu (Shimane).
Type locality. Honshu (Shizuoka), Japan.
Distribution. Japan (Honshu).

Micrasema gelidum McLachlan 1876

Micrasema gelidum McLachlan 1876, 263-264, male, Finland.
Fischer 1970, 134-135.
Schmid 1983, 32-33, male, female.
Botosaneanu 1988, 181-210, male, Holarctic (Boreal and Arctic).
Kuhara et al. 1994, 20, Hokkaido (Ishikari).
Kuranishi and Kuhara 1994, 1227, Hokkaido (Kushiro).
Ito 1995, 493-501, male, female, pupa, larva, egg, case, habitat, flight period.
Kuhara 1997b, 95, Hokkaido (Shiribeshi).
Ito et al. 1998, 11-15, Hokkaido (Nemuro).
Ito et al. 1999, 27, Hokkaido (Kushiro).
Ohkawa 1999, 41-44, Hokkaido (Tokachi).
Kuhara et al. 2000, 252, Hokkaido (Iburi).
Ito et al. 2000b, 31, Hokkaido (Ishikari).
Nozaki 2005a, 498, 500-501, male, larva.
Kuhara et al. 2007, ap. 71, Hokkaido (Ishikari).
Ito et al. 2009, 8, Hokkaido (Okhotsk).
Ito et al. 2010, 62, 75, Hokkaido (Shiribeshi).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Kuhara 2011, 52, 52, 57, 62, 73, Hokkaido (Iburi).
Ito 2013, 59, Hokkaido (Okhotsk, Kushiro).
Nozaki 2016, 363-364, 436, male.
Tanida 2016, 103.
Nozaki 2018a, 592-594, 596, male, larva, case.
Nakatani and Ito 2019, 51, 163, Hokkaido (Kushiro).
Ito and Kuhara 2023, 66, Hokkaido (Kamikawa).
Type locality. Finland.
Distribution. Japan (Hokkaido), Holarctic (Boreal and Arctic).
Remarks. The case of a young instar or an early period of the final instar larva described by Ito (1995) is a case of M. hanasense (Nozaki et al. 1999).

Micrasema hanasense Tsuda 1942

Micrasema hanasense (nomen nudum): Tsuda 1941, 104, Honshu (Nagano) (hanasensis).
Micrasema hanasense Tsuda 1942, 327-328, male, Honshu (Kyoto) (hanasensis).
Tsuda 1943, 163, Honshu (Kyoto) (hanasensis).
Fischer 1970, 136.
Kobayashi 1985, 19, Tsushima (hanasensis).
Nozaki 1987, 41, pl. 7, larva, Honshu (Kanagawa) (hanasensis).
Nozaki 1988, 41, Honshu (Kanagawa) (hanasensis).
Tanida 1990, 56, Honshu (Hyogo) (hanasensis).
Hatta and Nozaki 1991, 203, Honshu (Aichi) (hanasensis).
Moriya 1992, 43, Honshu (Kanagawa) (hanasensis).
Kuhara 1994, 20, Hokkaido (Ishikari) (hanasensis).
Morita 1995, 33, Honshu (Mie) (hanasensis).
Kuhara 1997a, 61, Hokkaido (Shiribeshi) (hanasensis).
Moriya 1997, 67, Honshu (Kanagawa) (hanasensis).
Nakajima et al. 1997, 18-19, Hokkaido (Oshima) (hanasensis).
Nozaki 1997, 37, Honshu (Kanagawa) (hanasensis).
Kuranishi and Kuhara 1994, 1227, Hokkaido (Kushiro) (hanasensis).
Ito et al. 1998, 11-17, Hokkaido (Nemuro, Okhotsk) (hanasensis).
Kagaya 1998, 27, 56-57, 181, Honshu (Tokyo, Yamanashi) (hanasensis).
Ito et al. 1999, 25, 27, Hokkaido (Kushiro) (hanasensis).
Ito et al. 2000, 28-30, Hokkaido (Soya) (hanasensis).
Kato and Kagaya 1999, 160-163, distribution pattern, Honshu (Tochigi) (hanasensis).
Nozaki et al. 1999, 355 (hanasensis).
Ohkawa 1999, 41-44, Hokkaido (Tokachi) (hanasensis).
Ito et al. 2000b, 31, Hokkaido (Ishikari) (hanasensis).
Kuhara et al. 2000, 252, Hokkaido (Iburi) (hanasensis).
Kawase and Morita 2001, 9, Honshu (Ishikawa) (hanasensis).
Kuhara 2001, 18, Hokkaido (Shiribeshi) (hanasensis).
Kuhara and Sato 2001, 64, Rishiri (hanasensis).
Morita 2001b, 339, Honshu (Mie) (hanasensis).
Nozaki 2001, 16, Honshu (Kanagawa) (hanasensis).
Kawase et al. 2003, 113, Honshu (Ibaraki) (hanasensis).
Morita 2003, 25, Honshu (Mie) (hanasensis).
Katsuma 2005, 7, Honshu (Ibaraki) (hanasensis).
Nozaki 2005a, 500-501, male, larva, case (hanasensis).
Katsuma 2006, 38, 41, Honshu (Ibaraki) (hanasensis).
Mey and Nozaki 2006, 24, Honshu (Nagano) (hanasensis).
Morita 2006, 200, Honshu (Mie) (hanasensis).
Torii and Hattori 2006, 38, Honshu (Shiga) (hanasensis).
Ito and Kosugi 2007, 54, Hokkaido (Kushiro) (hanasensis).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Tanida 2007, 248-249, female, Honshu (Shizuoka) (hanasensis).
Inazu 2008, 78-79, 93, 102, Honshu (Hyogo).
Morita 2008a, 723, Honshu (Mie) (hanasensis).
Morita 2008b, 92, Honshu (Mie) (hanasensis).
Aoyagi and Iwasaki 2009, 101, 104, 106, Honshu (Osaka) (hanasensis).
Morita 2009a, 6-7, Honshu (Shiga) (hanasensis).
Morita 2009b, 140, Honshu (Mie) (hanasensis).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka).
Ito et al. 2010, 62, 75-76, Hokkaido (Shiribeshi, Hiyama, Oshima).
Katsuma 2010, 85, Honshu (Ibaraki) (hanasensis).
Kawase and Hayashi 2010, 85, Honshu (Shimane) (hanasensis).
Kawase and Morita 2010, 43, Honshu (Shiga).
Shimura, 2010, 50, Yonaguni-jima.
Inazu and Nishida 2011, 196, male, female, larva, case, Honshu (Hyogo).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Katsuma 2011a, 49-51, Honshu (Ibaraki).
Katsuma 2011b, 68, Honshu (Ibaraki).
Kuhara 2011, 52, 57, 63, 73, Hokkaido (Sorachi, Ishikari, Hidaka, Iburi).
Inazu 2012, 8, Honshu (Hyogo).
Katsuma 2012, 65, Honshu (Ibaraki).
Inazu 2013b, 57, Honshu (Hyogo).
Inazu 2013c, 74, Honshu (Hyogo).
Ito 2013, 60, Hokkaido (Okhotsk, Nemuro, Tokachi).
Ijima and Tsuchiya 2014, 31, Hokkaido (Kushiro).
Ito & Nagasaka 2014, 11, Hokkaido (Honshu).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Yamamoto and Ito 2014, 14, 18-21, Shikoku (Ehime).
Katsuma 2015, 91, Honshu (Fukushima).
Morita and Kawase 2016, 22, Honshu (Ishikawa).
Nozaki 2016, 363-364, 436, male.
Tanaka 2016, 2, 5, male, Honshu (Akita).
Tanida 2016, 103.
Yamamoto and Ito 2016, 40, 43, Shikoku (Ehime).
Hayashi et al. 80, Kyushu (Miyazaki).
Ito 2017a, 5, Honshu (Fukui).
Ito 2017b, 44, Hokkaido (Kamikawa).
Katsuma 2017, 39-41, Honshu (Ibaraki).
Kuhara and Ito 2017, 14, 16, Yaku-shima.
Nojima 2017, 120, Honshu (Okayama).
Nozaki 2018a, 592-594, 596, male, larva, case.
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Tanaka 2018, 9, Honshu (Akita).
Nakatani and Ito 2019, 163, Hokkaido (Kushiro).
Nozaki et al. 2019, 169, Honshu (Nagano).
Tanaka 2019, 27, Honshu (Akita).
Yamamoto et al. 2019, 72, Shikoku (Ehime).
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka).
Yoshinari et al. 2020, 46, Honshu (Shizuoka).
Nakatani 2021, 72, Hokkaido (Kushiro).
Ito and Kuhara 2023, 66-67, Hokkaido (Soya, Kamikawa).
Micrasema sp. MA: Akagi 1957, 28, larva, case. Associated by Nozaki (1987).
Tsuda and Akagi 1962, 141, larva, case.
Tanida 1985, 193, larva, case.
Micrasema hakonense Kobayashi 1969, 19-20, pl. 4, male, Honshu (Kanagawa) (hakonensis). Synonymized by Nozaki and Tanida (2007).
Kobayashi 1971b, 36-37, pl. 23, male, Honshu (Kanagawa) (hakonensis).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rishiri, Tsushima, Yonaguni-jima), Kuriles, Korea, Taiwan.
Remarks. Tsuda (1941) used the name Micrasema hanasense (as hanasensis) ahead of the publication by Tsuda (1942). Although Tsuda (1941) assigned this species as a member of an ecological group, he did not provide any description or definition for the taxon. Therefore, the name appeared in Tsuda (1941) is not available (ICZN Art. 13.1.1).
Nozaki et al. (1999) suggested that the male genitalia illustrated by Akagi (1957) belongs to Nippoberaea gracilis (Nozaki and Kagaya 1994). Shimura (2015) recorded this species from Fukue-jima based on identification of the larval stage.

Micrasema quadriloba Martynov 1933

Micrasema quadriloba Martynov 1933, 147-149, male, Honshu (Tochigi).
Tsuda 1942, 327, Honshu (Nagano).
Akagi 1956, 77-78, larva, case, Honshu (Kyoto).
Chihara 1956, 79-80, pupa, Honshu (Kyoto).
Tsuda 1959, 151, larva, case.
Tsuda and Akagi 1962, 141, larva, case.
Tanida 1985, 193, larva, case.
Isobe et al. 1994, case, life cycle, 27-32, Honshu (Nara).
Kato and Kagaya 1999, 160-163, distribution pattern, Honshu.
Nozaki et al. 1999, 356.
Katano et al. 2002, 523-528, food, Honshu (Nara).
Isobe 2003, 12-15, life cycle, case.
Katano et al. 2005a, larval drift, 18-20, Honshu (Nara).
Katano et al. 2005b, 219-221, crawling activity, Honshu (Nara).
Katano et al. 2005c, 854-859, reach-scale distribution, Honshu (Nara).
Nozaki 2005a, 500-501, male, larva, case.
Katsuma 2008, 38, 40, Honshu (Ibaraki) (quadrilobum).
Aoyagi and Iwasaki 2010, 32-33, 35, 38, Honshu (Osaka) (quadrilobum).
Nishimoto and Nishimoto 2014, Honshu (Aichi) (quadrilobum).
Nozaki 2016, 364, 436, male.
Tanida 2016, 103.
Ito 2017a, 5, Honshu (Fukui).
Katano et al. 2017, drift dynamics.
Katsuma 2017, 39-41, Honshu (Ibaraki).
Nozaki 2018a, 592, 593, 595, 596, male, larva, case.
Type locality. Honshu (Tochigi), Japan.
Distribution. Japan (Honshu).

Micrasema spinosum Nozaki and Tanida 2007

Micrasema spinosum Nozaki and Tanida 2007, 249-250, male, female, Hokkaido (Hiyama, Oshima), Honshu (Akita, Shizuoka, Shiga).
Ito et al. 2010, 62, 76, Hokkaido (Shiribeshi, Hiyama).
Kawase and Morita 2010, 43, Honshu (Shiga).
Inazu and Nishida 2011, 197, male, Honshu (Hyogo).
Inazu 2012, 8, Honshu (Hyogo).
Nozaki 2016, 364, 436, male.
Tanaka 2016, 2, 5, male, Honshu (Akita).
Tanida 2016, 103.
Nojima 2017, 120, Honshu (Okayama).
Nozaki 2018a, 592-594, male, larva.
Tanaka 2018, 9, Honshu (Akita).
Nozaki et al. 2019, 170, 172, 174-175.
Yamamoto et al. 2019, 72, Shikoku (Ehime).
Ohtaka et al. 2020, 12, Honshu (Aomori).
Type locality. Honshu (Shizuoka), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku).

Micrasema uenoi Martynov 1933

Micrasema uenoi Martynov 1933, 149-150, male, Honshu (Tochigi).
Tsuda 1942, 327, Honshu (Nagano).
Fischer 1970, 150.
Nozaki 1988, 41, Honshu (Kanagawa).
Hatta and Nozaki 1991, 204, Honshu (Aichi).
Kato 1995, 131-135, habitat, Honshu (Tochigi).
Kato 1997, 213-217, habitat, food, Honshu (Tochigi).
Moriya 1997, 68, Honshu (Kanagawa).
Nozaki 1997, 37, Honshu (Kanagawa).
Kagaya et al. 1998, 27, 57, 182, Honshu (Tokyo, Yamanashi).
Kato and Kagaya 1999, 160-163, distribution pattern, Honshu (Tochigi).
Nozaki et al. 1999, 356.
Kawase and Morita 2001, 9, Honshu (Ishikawa).
Morita 2001a, 221, Honshu (Mie).
Morita 2001b, 339, Honshu (Mie).
Kawase et al. 2003, 113, Honshu (Ibaraki).
Inazu 2008, 79, 93, 102, Honshu (Hyogo).
Morita 2009a, 6-7, Honshu (Shiga).
Morita and Kawase 2009, 28, Honshu (Toyama).
Kawase and Morita 2010, 43, Honshu (Mie, Shiga).
Kawase and Hayashi 2010, 85, Honshu (Shimane).
Kawase et al. 2010, 108, Honshu (Shimane).
Inazu and Nishida 2011, 196, male, female, larva, case, Honshu (Hyogo).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 65, Honshu (Ibaraki).
Inazu 2013a, 46, Honshu (Hyogo).
Inazu 2013c, 74, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nozaki et al. 2014, 31, Honshu (Nagano).
Nozaki 2016, 78, 364-365, 436, 450, male, Honshu (Nagano).
Tanida 2016, 103.
Kawase et al. 33, Honshu (Shiga).
Nojima 2017, 120, Honshu (Okayama).
Nozaki 2018a, 592-594, 596, male, larva.
Nozaki 2018b, 930, Honshu (Kanagawa).
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Ohtaka et al. 2020, 12, Honshu (Aomori).
Yamamoto et al. 2021, 74, Shikoku (Ehime).
Aoya and Tanaka 2023, 27, Honshu (Akita).
Micrasema sp. MC: Akagi 1962b, 44, larva, case, Honshu (Aichi) . Associated with adult stage by Kagaya et al. (1998).
Tanida 1985, 193, larva.
Tada and Satake, 1994, 161-163, larval habitat, Honshu (Tochigi).
Micrasema genijiroense Kobayashi 1971a, 6-7, pl. 6, male (genjiroensis). Synonymized by Nozaki and Tanida (2007).
Kobayashi 1971b, 37-38, pl. 24, male, Honshu (Kanagawa) (genjiroensis).
Micrasema sp.: Kimura and Hirabayashi 2013, 185, Honshu (Nagano). Mentioned by Nozaki et al. (2014).
Type locality. Honshu (Tochigi), Japan.
Distribution. Japan (Honshu, Shikoku).

Micrasema sp.

Micrasema minimum: Iwata, 1927, 260, larva, case.
Micrasema sp.: Nozaki et al. 1999, 356.

Micrasema spp.

Lasiocephala basalis Kolenati: Iwata 1927a, 262-263, larva, case.
Micrasema spp.: Nozaki et al. 1999, 356.
Remarks. Larvae recorded as L. basalis by Iwata (1927a) belong to the genus Micrasema (Ito 1990).

Genus Tsudaea Nozaki 2009

Recentry, Nozaki (2009) erected this genus for Brachycentrus kitayamanus Tsuda 1942.

Tsudaea kitayamana (Tsuda 1942)

Brachycentrus kitayamanus Tsuda 1942, 326, male, Honshu (Kyoto).
Micrasema sp. MD: Akagi, 1962b, 44-45, larva, case, Honshu (Nagano).
Eobrachycentrus kitayamanus: Wiggins et al. 1985, 66.
Morita and Kawase 2004, 18, Honshu (Ishikawa).
Tsudaea kitayamana Nozaki 2009, 56-63, male, female, larva, pupa, case, food, habitat, Honshu (Ishikwa, Yamanashi, Nagano, Gifu, Mie, Hyogo).
Morita 2009c, 121, Honshu (Mie).
Inazu and Nishida 2011, 197, male, larva, case, Honshu (Hyogo).
Inazu 2013a, 46, 50, adult, larva, Honshu (Hyogo).
Nozaki et al. 2014, Honshu (Nagano).
Nozaki 2016, 78, 360, 365, 436, 450, male, Honshu (Yamanashi).
Tanida 2016, 104.
Nojima 2017, 121, Honshu (Okayama).
Nozaki 2018a, 590, 591, 593, 595, male, larva, case.
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu).
Remarks. Ohira and Tsutsumi (2018) recordes a female as possibly T. kitamana from Fukushima, northern Honshu.

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