A catalogue of Japanese Trichoptera
Family Goeridae Ulmer
Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last updated: 31 Dec. 2023
Subfamily Goerinae Ulmer
Two genera, Goera and Silo, were recorded from Japan, but identification of Silo is very doubtful (Nozaki et al. 2000)
Genus Goera Stephens
Eighteen species belonging to this genus are recognized for Japanese fauna.
Goera akagiae Tanida and Nozaki 2006
Goera sp. GC: Akagi 1974, 18-19, larva, case, male, Amami-Oshima. Associated with the adult stage by Nozaki and Tanida (2006).
Tanida 1985, 196, 197, larva.
Nozaki et al. 2000, 198.
Nozaki 2005a, 536, 537, larva.
Goera akagiae Nozaki and Tanida 2006, 7-9, male, female, Shikoku (Kochi), Yaku-shima, Amami-Oshima.
Shimura et al. 2014, 46-47, Amami-oshima.
Nozaki 2016, 82, 389, 391, 440, 451, male, Shikoku (Kochi).
Tanida 2016, 124.
Kuhara and Ito 2017, 14, 16, Yaku-shima.
Nozaki 2018, 637, 640, 641, male, larva.
Shimura and Imasaka 2021, 5, Kamikoshiki-jima.
Type locality. Amami-Oshima, Japan.
Distribution. Japan (Shikoku, Koshiki-jima, Yaku-shima, Amami-Oshima).
Goera curvispina Martynov 1935
Goera curvispina Martynov 1935, 367-370, male.
Tsuda and Akagi 1955, 237, Honshu (Shiga).
Akagi 1975, 3-4, larva, case, Honshu (Yamaguchi), Kyushu (Fukuoka, Oita).
Gyotoku and Nozaki 1992, 138, Honshu (Fukuoka).
Fischer 1973, 1.
Tani 1977, 205, male.
Arefina 1997, 127-128, male.
Nozaki et al. 2000, 198.
Uenishi and Yoshitaka 2004, 17, Honshu (Okayama).
Nozaki 2005a, 535-538, male, larva.
Nozaki and Tanida 2006, 10-12, male, female, Honshu (Hyogo, Shimane), Kyushu (Fukuoka), Russia.
Nakamura and Hajiri 2007, 199, Honshu (Shimane).
Inazu 2013c, 76, Honshu (Hyogo).
Nozaki 2016, 389, 391, 440, male.
Tanida 2016, 124.
Ito 2017a, Honshu (Fukui).
Kawase et al. 2017, 36, Honshu (Shiga).
Nojima 2017, 123, Honshu (Okayama).
Nozaki 2018, 637, 640, 641, male, larval leg.
Kuhara et al. 2023, 19, Kyushu (Miyazaki).
Goera gyotokui Kobayashi 1957, 276-278, male, Fukuoka. Synonymized by Nozaki and Tanida (2006).
Fischer 1973, 2.
Nozaki et al. 2000, 198.
Kobayashi 1959, 354, Fukuoka (Mis-spelled as G. gyotokuana).
Type locality. South Primorye (Ussuri R.), Russia.
Distribution. Japan (Honshu (central to western), Kyushu), Korea, Russia (Khabarovsk, Primorye).
Remarks. Tani (1976) recorded this species from Tsushima based on examination of larvae.
Goera dilatata Nozaki and Tanida 2006
Goera dilatata Nozaki and Tanida 2006, 25-27, male, female, Hokkaido (Oshima), Honshu (Iwate, Miyagi, Akita, Niigata).
Katsuma 2012, 66, Honshu (Ibaraki).
Katsuma 2015, 90-91, Honshu (Ibaraki).
Nozaki 2016, 390-391, male.
Tanida 2016, 124.
Tanaka 2017a, 26-27, male, Honshu (Akita).
Aoya 2018, 4-5, larva, case, Honshu (Akita).
Nozaki 2018, 638, 640, 641, male.
Kawase & Kato 2023, 9, Honshu (Niigata).
Tanaka 2023, 44, Honshu (Akita).
Type locality. Honshu (Akita), Japan.
Distribution. Japan (Hokkaido (southern), Honshu (northern))
Goera extrorsa Nozaki 2017
Goera extrorsa Nozaki 2017, 156-158, male, female, Honshu (Ishikawa, Fukui, Mie, Shiga).
Nojima 2022, 39, Honshu (Okayama).
Goera spicata: Morita 2009, 121; Morita and Kawase 2009, 28. Misidentification mentioned
by Nozaki (2017).
Goera spicata?: Kawase et al. 2017, 36. Mentioned by Nozaki (2017).
Goera sp.: Ito, 2017a, 6. Mentioned by Nozaki (2017).
Type locality. Honshu (Fukui), Japan.
Distribution. Japan (Honshu).
Goera japonica Banks 1906
Goera japonica Banks 1906, 108-109, male, Honshu (Shizuoka?).
Ota 1917, 489-494, parasitoid, Honshu (Kanagawa: larval record).
Matsumura 1931, 191, adult.
Matsumura 1933, 2, pl. 1, adult.
Tateishi 1934, 75-76, Kyushu (Kagoshima).
Tsuda 1941, 104, Honshu (Nagano).
Kobayashi 1942, 220, Honshu (Tokyo).
Tsuda 1942a, 323-324, Honshu (Nagano, Shizuoka, Kyoto, Osaka, Hyogo), Kyushu
(Fukuoka).
Tsuda 1942b, 263-265, Honshu (Kyoto).
Tsuda 1943, 163166-168, Honshu (Kyoto).
Tsuda and Kuwayama 1950, 428, adult.
Tsuda 1951, 66, larva, case.
Tsuda and Tatsumi 1951, 30, Honshu (Niigata).
Ishihara et al. 1953, 38, Shikoku (Ehime).
Tsuda and Akagi 1955, 235-236, larva, case.
Chihara 1956, 81-82, pupa.
Gyotoku 1957, 12-13, Kyushu (Fukuoka).
Miwa and Ohkawa 1958, 19, Honshu (Mie).
Kobayashi 1959, 353-354, Kyushu (Fukuoka).
Tsuda 1959, 150, larva, case.
Tsuda and Akagi 1962, 140, larva, case.
Kuwada 1965, 166, pl.. 83, adult, Shikoku (Ehime).
Fischer 1967, 7.
Kobayashi 1968, 6, Honshu (Niigata).
Kobayashi 1971, 35-36, male, female, Honshu (Kanagawa).
Foscher 1973, 2.
Tani 1976, 393, Tsushima.
Tani 1977, 194, 205, pl. 44, male, Honshu (Nara).
Tani 1978, 16-17, Honshu (Niigata).
Tanida 1980, 136, Honshu (Ishikawa).
Tanida and Takemon 1981, 206, Honshu (Kyoto).
Tanida 1982a, 361, Honshu (Gumma).
Tanida 1982b, 18-20, Honshu (Ishikawa).
Kobayashi 1983, 32, Honshu (Kanagawa.)
Kobayashi 1984a, 20, male, female, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kobayashi 1984b, 7-8, Sado.
Kobayashi 1984c, 28, Honshu (Kanagawa).
Kobayashi 1985a, 19, Tsushima.
Kobayashi 1985b, 68, Honshu (Kanagawa).
Tanida 1985, 196, 197, larva, case.
Togashi and Tanida, 1985, 69, Honshu (Ishikawa).
Ohtsuka and Tanida. 1986, 6, Kyushu (Kumamoto).
Kobayashi 1987, 23-24, 1987, Honshu (Shimane).
Yamanouchi 1987, 36, Hokkaido (Tokachi).
Nozaki 1987, 41-42, Honshu (Kanagawa).
Morita 1988, 5, Honshu (Mie).
Nozaki 1988a, 45-46, Honshu (Kanagawa).
Nozaki 1988b, 42, Honshu (Kanagawa).
Morita 1989, 40, Honshu (Mie).
Nozaki and Gyotoku 1990, 17, Kyushu (Fukuoka).
Kugo and Nozaki 1990, 63, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Tanida 1990, 56, Honshu (Hyogo).
Aoyagi and Ishii 1991, 113-118, parasitoid.
Hatta and Nozaki, 1991, 204, Honshu (Gifu).
Kugo and Nozaki, 1991, 8, Honshu (Hyogo).
Tanida 1991, 35, Honshu (Nara).
Tanida et al. 1991, 100-101, adult, pupa, larva, case.
Gyotoku and Nozaki 1992, 137-138, Kyushu (Fukuoka).
Moriya 1992, 44, 63, adult, case, Honshu (Kanagawa).
Hori et al. 1993, 375, 389, Honshu (Aichi).
Kobayashi 1993, 401-402, parasitoid.
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Morita 1993, 6, Honshu (Mie).
Nishimoto and Nishimoto 1993, 87, Honshu (Aichi).
Tanida and Takemon 1993, 243, Honshu (Kyoto)
Kuhara and Ito 1994, 21, Hokkaido (Ishikari).
Kuranishi and Kuhara 1994, 1229, Hokkaido (Akan).
Moriya 1994, 79, Honshu (Kanagawa, Yamanashi).
Itou and Muramatsu 1995, 41-42, Hokkaido (Okhotsk).
Kobayashi 1995, 318-321, parasitoid.
Tanida et al. 1995, 3, Honshu (Nara).
Kuranishi and Ohtaka 1996, 15, Honshu (Aomori).
Nakamura 1996, 294, Honshu (Hiroshima).
Kuhara 1997a, 61, Hokkaido (Shiribeshi).
Kuhara 1997b, 96, Hokkaido (Shiribeshi)
Nakajima et al. 1997, 14, Hokkaido (Oshima).
Ito et al. 1997, 26, 28, 30, 31, Hokkaido (Hiyama).
Morita 1997a, 41, Honshu (Mie).
Morita 1997b, 96, Honshu (Mie).
Moriya 1997, 68, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 36, Honshu (Kanagawa).
Nozaki 1997b, 30, Honshu (Tokyo).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Kadowaki 1998, 8, Honshu (Shimane).
Kagaya et al. 1998, 29, 61, 199, Honshu (Tokyo, Yamanashi).
Kobayashi, 1998, 109-113, parasitoid.
Ito et al. 1998, 12, 13, 15, 16, Hokkaido (Shibetsu).
Morita 1998a, 14, Honshu (Mie).
Morita 1998b, 73, Honshu (Mie).
Sekimoto et al. 1998, 361, Honshu (Hiroshima).
Yodoe et al. 1998, 69, Honshu (Shimane).
Ito et al. 1999, 25-27, Hokkaido (Akan).
Nakano and Tanida 1999, 277, Honshu (Nara).
Ohkawa 1999, 41-44, Hokkaido (Tokachi).
Imamori 2000, 234-235, adult, larva.
Ito et al. 2000a, 27-31, Hokkaido (Soya).
Ito et al. 2000b, 35, Hokkaido (Ishikari).
Kuhara et al. 2000, 255, Hokkaido (Iburi).
Maruyama and Takai 2000, 99, 108, larva, case.
Nishimoto 2000, 26, Honshu (Aichi).
Nozaki et al. 2000, 198.
Uenishi and Ebisawa 2000, 170, Honshu (Shiga).
Kawase and Morita 2001, 9, Honshu (Ishikawa).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Morita 2001a, 221, Honshu (Aichi).
Morita 2001b, 164, Hosnhu (Mie).
Morita 2001c, 339, Honshu (Mie).
Nishimoto and Morita 2001, 77, Hosnhu (Aichi).
Kuranishi 2002, 571, larva, Honshu (Chiba).
Nakamura et al. 2002, 67, Honshu (Hiroshima).
Nozaki and Nakamura 2002, 173-174, Honshu (Hiroshima).
Karita 2003, 130-131, adult, larva, case.
Nagayasu et al. 2003, 110, Hokkaido (Ishikari).
Nakamura and Umemori 2003, 59-60, Honshu (Hiroshima).
Shinoda 2003, 145, Honshu (Yamanashi).
Ito et al. 2004, 11, 16, Hokkaido (Kushiro).
Kawase et al. 2004, 114, Honshu (Ibaraki)
Morita and Kawase 2004, 16, 18, Honshu (Ishikawa).
Nakamura et al. 2004, 154, Honshu (Hiroshima).
Nozaki 2004, 1236, Honshu (Kanagawa).
Yoshitaka 2004, 28, Honshu (Okayama).
Ishiwata et al. 2005, app. 2(3), Honshu (Kanagawa).
Kanazawa 2005, 3-7, pl. 1, case, parasitoid, Honshu (Hiroshima).
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Tochigi, Gumma).
Nagayasu et al. 2005, 56, Hokkaido (Ishikari).
Nakano 2005, 305-308, pupal aggregation, Hokkaido (Ishikari).
Nozaki 2005a, 535-538, male, larva, case.
Nozaki 2005b, 112, adult, larva, case.
Nozaki and Nakamura 2005, 224, Honshu (Shimane).
Aoyagi 2008, 1-7, case, parasitoid, Honshu (Osaka). Larval record.
Kameyama and Nakamura 2006, 190, Honshu (Tottori, Hiroshima).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Katsuma 2006, 41, Honshu (Ibaraki).
Kimura et al. 2006, 8, Honshu (Nagano).
Kuhara et al. 2006, 46, Hokkaido (Ishikari).
Mey and Nozaki 2006, 25, Honshu (Nagano, Gifu, Shiga).
Morita 2006a 201, Honshu (Mie).
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Nozaki and Tanida 2006, 3-6, male, female, Hokkaido (Nemuro, Okhotsk, Soya, Kamikawa, Hiyama), Honshu (Miyagi, Akita,
Fukushima, Tochigi, Gumma, Tokyo, Kanagawa, Niigata, Nagano, Gifu, Shizuoka,
Aichi, Mie, Hyogo, Nara, Hiroshima, Shikoku (Tokushima, Ehime, Kochi),
Sado, Tsushima, Yaku-shima.
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Gall et al. 2007, 108-111, male, female, wings of female, Hokkaido (Iburi), Honshu (Kanagawa), Sakhalin, Kuriles, Cheju (South
Korea).
Ito and Kosugi 2007, 55, Hokkaido (Kushiro).
Katsuma 2007, 18, Honshu (Ibaraki).
Kimura et al. 2007, 167, Honshu (Nagano).
Kuhara et al. 2007, ap. 72, Hokkaido (Ishikari).
Morita 2007a, 15, Honshu (Mie).
Morita 2007b, 97, Honshu (Mie).
Nakamura 2007, 78, Honshu (Hiroshima).
Nakamura and Tsujimura 2007, 78, Honshu (Hiroshima).
Nakano et al. 2007, 41-42, Hokkaido (Nemuro).
Nakano et al. 2007, 103-110, effects by case aggregation.
Nozaki and Nakamura 2007, 98, Honshu (Hiroshima).
Nozaki and Tanida 2007, 251, Honshu (Shizuoka).
Shinoda 2007, 97, Honshu (Yamanashi).
Inazu 2008, 83-84, 95, 103, Honshu (Hyogo).
Katsuma 2008, 58, Honshu (Ibaraki).
Kimura et al. 2008, 264, Honshu (Nagano).
Morita 2008a, 17, Honshu (Mie).
Morita 2008b, 724, Honshu (Mie).
Morita and Kawase 2008, 56, Honshu (Ishikawa).
Muramatsu 2008, 11-12, Hokkaido (Okhotsk).
Nakatani et al. 2008, 55, Hokkaido (Soya).
Tanida 2008, 253, pl. 94, adult, Shikoku (Ehime).
Aoyagi and Iwasaki 2009, 101, 104, 107, Honshu (Osaka).
Kameyama et al. 2009, 113, Honshu (Shimane).
Morita and Kawase 2009, 28-29, Honshu (Ishikawa).
Tanaka 2009a, 48, adult, Honshu (Akita).
Tanaka 2009b, 19, Honshu (Akita).
Aoyagi and Iwasaki 2010, 32-33, 35, 39, Honshu (Osaka).
Ito 2010, 131, Hokkaido (Kushiro).
Ito et al. 2010, 64, 77, Hokkaido (Shiribeshi, Hiyama, Oshima).
Nakamura 2010, 239, Honshu
(Hiroshima).
Nakamura and Kita 2010, 89, Honshu (Hiroshima).
Mano 2010, 21-23, Honshu (Aichi).
Katsuma 2010a, 84, Honshu (Ibaraki).
Katsuma 2010b, 86, Honshu (Ibaraki).
Kawase and Morita 2010, 45, Honshu (Mie, Shiga).
Nozaki 2010, 217, Honshu (Kanagawa).
Tanaka 2010a, 26, Honshu (Akita).
Tanaka 2010b, 17-19, male, Honshu (Akita).
Hirabayashi et al. 2011, 145, Honshu (Nagano).
Inazu and Nishida 2011, 206, male, larva, case, Honshu (Hyogo).
Ito and Kubo 2011, 10, Hokkaido (Tokachi).
Katsuma 2011, 68, Honshu (Ibaraki).
Kawase and Hayashi 2011a, 173, Honshu (Shimane).
Kawase and Hayashi 2011b, 192, Honshu (Shimane).
Kuhara 2011, 53, 57, 63, 74, Hokkaido (Sorachi, Ishikari).
Morita and Katsuma 2011, 99, Honshu (Mie).
Morita 2011, 217, Honshu (Mie).
Nakatani and Matsumoto 2011, 62, Hokkaido (Akan).
Tanaka 2011, 24, Honshu (Akita).
Tsukiji 2011, 141, adult.
Inazu 2012, 10, Honshu (Hyogo).
Katsuma 2012, Honshu (Ibaraki).
Morita 2012, 251, Honshu (Mie).
Morita and Kawase 2012, 14-16, Honshu (Ishikawa).
Moriya and Kubota 2012, 135, Honshu (Kanagawa).
Kawase and Hayashi 2012, 257, Honshu (Shimane, Oki (Dogo)).
Enju et al. 2013, 86, 90, adult, case.
Hayashi et al. 2013, 13, Oki (Dogo, Nakano-shima).
Inazu 2013a, 47, Honshu (Hyogo).
Inazu 2013b, 58, Honshu (Hyogo).
Inazu 2013c, 76, Honshu (Hyogo).
Ito 2013, 67, Hokkaido (Okhotsk, Kushiro, Nemuro).
Kawabe 2014, 115, adult, Honshu (Tokyo).
Kawase and Morita 2014, 7, Honshu (Shiga).
Ijima and Tsuchiya 2014, 33, Hokkaido (Okhotsk. Kushiro, Tokachi).
Ito and Nagasaka 2014, 12, Hokkaido (Sorachi).
Kuranishi and Kikuchi 2014, 364, Honshu (Yamagata).
Nishimoto and Nishimoto 2014, 66, Honshu (Aichi).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Shimura 2015, 54, Fukue-jima.
Hayashi & Kadowaki 2016, 33, 68-69, pupal case, Oki.
Hirabayashi et al. 2016, 208, Honshu (Nagano).
Horiguchi 2016, 414-415, Honshu (Tokyo).
Katsuma 2016, 48, Honshu (Ibaraki).
Morita and Kawase 2016, 21-23, Honshu (Ishikawa).
Nakamura 2016, 21, 24, Honshu (Hiroshima).
Nakatani 2016, 242, Hokkaido (Nemuro).
Nozaki 2016, 83, 305, 389-391, 440, 451, male, Shikoku (Kochi).
Tanida 2016, 124.
Uchifune 2016, 201-202, adult, Honshu (Kanagawa).
Yamamoto and Ito 2016, 42, 43, Shikoku (Ehime).
Ito 2017a, 6, Honshu (Fukui).
Ito 2017b, 45, Hokkaido (Kamikawa).
Kuhara and Ito 2017, 14, 16, Yaku-shima.
Kawase et al. 2017, 36, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga, Kyoto).
Matsuoka 2017, 10, Hokkaido (Tokachi).
Saito et al. 2017, 184, Hoshu (Chiba).
Nojima 2017, 123, Honshu (Okayama).
Tanaka 2017a, 26, 28, male, Honshu (Akita).
Tanaka 2017b, 3, Honshu (Akita).
Nozaki 2018, 637, 640, 641, male, larva, case.
Ohira and Tsutsumi 2018, 25, Honshu (Fukushima).
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka).
Tanaka 2018a, 22, Honshu (Akita).
Tanaka 2018b, 9, Honshu (Akita).
Aoyagi and Tada 2019, 27-31, case, parasitoid, Shikoku (Tokushima, Kagawa, Ehime, Kochi).
Morita 2019, 66, Honshu (Mie).
Nakatani and Ito 2019, 164, Hokkaido (Kushiro).
Nozaki et al. 2019, 171, Honshu (Nagano).
Yamamoto et al. 2019, 73, 77, larva, Shikoku (Ehime).
Tanaka 2019, 13, Honshu (Akita).
Katsuma 2020, 15, Kyushu (Oita).
Morimune and Nakamura 2020, 21, 23, Honshu (Okayama).
Morita 2020b, 43, Honshu (Mie).
Morita 2020c, 45, Honshu (Mie).
Tezuka et al. 2021, 43, Honshu (Kanagawa).
Watanabe and Kawashima 2021, pupal case, host of water wasp, Honshu (Kanagawa).
Yamamoto and Nozaki 2021, 129, Shikoku (Ehime).
Muramatsu 2022, 46, 63, male, female, larva
Tanaka 2022, 37, Honshu (Akita).
Aoya and Tanaka 2023, 27, Honshu (Akita).
Ito and Kuhara 2023, 67-68, Hokkaido (Soya, Kamikawa, Rumoi).
Kawase & Kato 2023, 9/15, Honshu (Niigata).
Tanaka 2023, 44, Honshu (Akita).
Yamamoto 2023, 34, Shikoku (Ehime).
Lithax karafutonis Matsumura 1911, 16, female (as male in the original description), Sakhalin. Synonymized by Gall et al. 2007).
Goera pilosa Fabricius: Iwata 1927a, 255-256, larva. Misidentification mentioned by Tsuda and Akagi (1955).
Iwata 1927b, 214, larva.
Fischer 1967, 15-16.
Goera nipponensis Navas 1933, 107-108, male, female, Honshu (Yamanashi). Synonymized by Nozaki and Tanida (2006).
Fischer 1967, 8.
Nozaki et al. 2000, 198.
Type locality. Honshu (Kawana: Shizuoka?), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku , Kyushu, Sado, Oki (Dogo, Nakanoshima),
Tsushima, Fukue-jima, Yaku-shima), Sakhalin, Kuriles, Korea (Jeju), Kazakhstan.
Remarks. Gall et al. (2007) resolved the taxonomic status of G. japonica and similar species from the eastern Palaearctic Region. Oláh et al. (2018) recently reported this species from Kazakhstan.
Goera kawamotonis Kobayashi 1987
Goera sp. GA, Tsuda and Akagi 1956, 39-40, larva, case, male, Honshu (Aichi). Associated with the adult stage by Nishimoto and Morita
(2001).
Tsuda and Akagi 1962, 140, larva.
Fischer 1973, 5.
Tanida 1985, 196, 199, larva.
Nozaki et al. 2000, 198.
Goera kawamotonis Kobayashi 1987, 24, 34, male, Honshu (Shimane).
Gyotoku and Nozaki 1992, 138, Kyushu (Fukuoka).
Arefina 1997, 126, 128, male.
Nozaki et al. 2000, 198.
Nishimoto and Morita 2001, 73, 77, adult, Honshu (Aichi).
Morita 2002, 37, Honshu (Mie).
Nozaki and Nakamura 2004, 224, Honshu (Shimane).
Yoshitaka 2004, 28, Honshu (Okayama).
Uenishi and Yoshitaka 2004, 17-18, Honshu (Okayama).
Nozaki 2005a, 535-538, male, larva, case.
Kameyama et al. 2006, 94, Honshu (Hiroshima)
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Nozaki and Tanida 2006, 11-13, male, female, Honshu (Aichi, Hyogo, Hiroshima, Shimane, Yamaguchi), Shikoku, Kyushu
(Fukuoka).
Nakamura and Tsujimura 2007, 79, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 98, Honshu (Hiroshima).
Morita 2008a, 17, Honshu (Mie).
Kameyama et al. 2009, 113, 198, Honshu (Shimane).
Mano 2010, 22-23, Honshu (Aichi).
Kawase and Hayashi 2011b, 192-193, Honshu (Shimane).
Morita and Katsuma 2011, 99, Honshu (Mie).
Nozaki 2016, 390-391, 440, male.
Tanida 2016, 124.
Nojima 2017, 123, Honshu (Okayama).
Nozaki 2018, 637, 640, 641, male, larva, case.
Kuhara et al. 2023, 19, Kyushu (Fukuoka, Oita).
Type locality. Honshu (Shimane), Japan.
Distribution. Japan (Honshu (central to western), Kyushu), Russia (Khabarovsk, Primorye),
Korea.
Remarks. Nozaki and Tanida (2006) added Shikoku as a distribution area, but no
data was provided.
Goera kyotonis Tsuda 1942
Goera kyotonis Tsuda 1942a, 324-325, male, Honshu (Kyoto).
Tsuda 1954, 26-27, Honshu (Shimane).
Tsuda and Akagi 1955, 236-237, larva, case, Honshu (Kyoto).
Kobayashi 1957, 278, pl. 44, male.
Tsuda and Akagi 1962, 140, larva, case.
Tsuda 1959, 150, larva, case.
Fischer 1967, 7.
Fischer 1973, 3.
Tani 1982, 34-35, Oki.
Morita 1996, 289, Honshu (Mie).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Nozaki et al. 2000, 198.
Nozaki 2005a, 535-538, male, larva, case.
Nozaki and Tanida 2006, 13-15, male, female, Honshu (Osaka, Hyogo, Wakayama, Tottori), Kyushu (Fukuoka).
Morita 2006b 47, Honshu (Mie).
Morita 2008b, 724, Honshu (Mie).
Kawase and Morita 2010, 45, Honshu (Shiga).
Morita 2011, 217, Honshu (Mie).
Morita 2012, 251, Honshu (Mie).
Morita 2013, 64, Honshu (Mie).
Nozaki 2016, 390-391, 440, male.
Tanida 2016, 124.
Ito 2017a, 6, Honshu (Fukui).
Imasaka et al. 2017, 61, Kyushu (Fukuoka).
Nojima 2017, 123, Honshu (Okayama).
Nozaki 2018, 637, 640, 641, male, larva, case.
Tanida et al. 2018, 31, Honshu (Toyama).
Morita 2020a, 19, Honshu (Mie).
Kuhara et al. 2023, 19, Kyushu (Oita).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu (central to western), Kyushu, Oki (based on larva)).
Goera lepidoptera Schmid 1965
Goera lepidoptera Schmid 1965, 35, male, Honshu (Nagano).
Schmid 1991, 325, male maxillary palpus.
Kagaya et al. 1998, 29, 61, 200, Honshu (Tokyo).
Nozaki et al. 2000, 198.
Nozaki 2004, 1236, Honshu (Kanagawa)
Nozaki 2005a, 538, male.
Katsuma 2006, 36, 41, Honshu (Ibaraki).
Nozaki and Tanida 2006, 15-16, male, female, Honshu (Miyagi, Yamagata, Kanagawa).
Tanaka 2010b, 17-19, male, Honshu (Akita).
Nozaki 2016, 391, 440, male.
Tanida 2016, 125.
Tanaka 2017a, 26, 28, male, Honshu (Akita).
Tanaka 2017b, 3, Honshu (Akita).
Nozaki 2018, 637, 640, 641, male.
Tsuruda 2018, 27-28, 32, Honshu (Tokyo).
Tanaka 2022, 37, Honshu (Akita).
Tanaka 2023, 44, Honshu (Akita).
Goera kyotonis Tsuda: Moriya 1992, 44, Honshu (Kanagawa). Misidentification mentioned
by Nozaki (2004).
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu central to northern))
Remarks. Larvae of this species are indistinguishable from those of G. kyotonis (Nozaki and Tanida 2006).
Goera minuta Ulmer 1927
Goera minuta Ulmer, 1927, 181-182, pls. 5-6, male, Taiwan.
Nozaki 2017, 158-159, male, Ishigaki-jima.
Nozaki 2018, 638, male.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima).
Goera nigrosoma Nozaki and Tanida 2006
Goera sp. GB: Tsuda and Akagi 1962, 140, larva.
Tanida 1985, 196, 199, larva.
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Kagaya et al. 1998, 29, 61, 202, Honshu (Tokyo).
Nozaki 2005a, 536, 537, larva.
Nozaki et al. 2000, 198.
Neophylax sp.: Nozaki 1988b, 42, Honshu (Kanagawa). Misidentification mentioned by Nozaki (1997a).
Goera sp.: Morita 1995, 49, male, Honshu (Mie). Reidentified as this species by Morita (2007).
Nozaki 1997a, 36, Honshu (Kanagawa).
Goera nigrosoma Nozaki and Tanida 2006, 20-22, male, female, Honshu (Kanagawa, Yamanashi, Nagano, Shizuoka, Kyoto, Osaka, Wakayama,
Okayama), Shikoku (Kagawa, Ehime, Kochi).
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Morita 2007, 15, Honshu (Mie).
Inazu and Nishida 2011, 206, male, Honshu (Hyogo).
Yamamoto and Ito 2015, 15, Shikoku (Ehime).
Nozaki 2016, 83, 390-391, 440, Shikoku (Kochi).
Tanida 2016, 125.
Nozaki 2018, 637-638, 640-641, male, larva, case.
Yamamoto et al. 2021, 75, Honshu (Ehime).
Type locality. Honshu (Osaka), Japan.
Distribution. Japan (Honshu (central to western), Shikoku).
Goera ogasawaraensis Kuranishi 2005
Goera sp.: Tomokuni and Sato 1978, 117-118, larva, case, Ogasawara (Chichi-jima: larval record). Associated with the adult stage
by Satake et al. (2005).
Nozaki et al. 2000, 198.
Goera ogasawaraensis: Satake et al. 2005, 376, 378-379, male, female, Ogasawara (Chichi-jima).
Nozaki and Tanida 2006, 5-6, male, female, Ogasawara (Chichi-jima).
Yoshinari et al. 2014, 29, 32, larva, Ogasawara.
Tanida 2016, 125.
Nozaki 2018, 6381, male.
Type locality. Ogasawara (Chichi-jima), Japan.
Distribution. Japan (Ogasawara (Chichi-jima)).
Remarks. Satake and Kuranishi (2007) recorded this species from Chichi-jima and
Ani-jima based on examination of larvae.
Goera rupicola Nozaki and Shimura 2020
Goera rupicola Nozaki and Shimura 2020, 574-578, male, female, pupa, larva, case, habitat, Amami-Oshima.
Kushibiki and Nozaki 2022, 31-34 male, female, habitat, Okinawa-jima
Type locality. Amami-Oshima, Japan.
Distribution. Japan (Amami-Oshima).
Goera shikokuensis Nozaki and Tanida 2006
Goera shikokuensis Nozaki and Tanida 2006, 15-17, male, female, Shikoku (Ehime, Kochi).
Nozaki 2016, 390-391, 440, male.
Tanida 2016, 125.
Nozaki 2018, 637, 640, 641, male.
Aoyagi and Tada 2019, 29-30, case, Shikoku (Kochi).
Type locality. Shikoku (Ehime), Japan.
Distribution. Japan (Shikoku).
Goera spicata Schmid 1965
Goera spicata Schmid 1965, 35, male, Honshu (Nagano).
Schmid 1991, 325, male maxillary palpus.
Moriya 1994, 79, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 68, Honshu (Kanagawa, Yamanashi).
Nozaki 1997, 36, Honshu (Kanagawa).
Kagaya et al. 1998, 29, 61, 201, Honshu (Tokyo, Yamanashi).
Nozaki et al. 2000, 198.
Nozaki 2004, 1236, Honshu (Kanagawa).
Nozaki 2005a, 538, male.
Mey and Nozaki 2006, 25, Honshu (Nagano).
Nozaki and Tanida 2006, 24-26, male, female.
Morita 2009, 121, Honshu (Mie).
Morita and Kawase 2009, 28, Honshu (Ishikawa).
Nozaki 2016, 390-391, 440, male.
Tanida 2016, 215.
Nozaki 2017, 157, male, female, Honshu (Yamanashi).
Nozaki 2018, 638, 640, 641, male, larva, case
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu (central)).
Remarks. Larval stage of this species was associated with adult and pupal stages by Kagaya et al. (1998), but the morphology has not yet been described.
Goera tajimaensis Tanida and Nozaki 2006
Goera tajimaensis: Nozaki and Tanida 2006, 22-23, male, female, Honshu (Hyogo).
Nozaki 2016, 391, 440, male.
Tanida 2015, 125.
Nozaki 2018, 638, 640, 641, male.
Aoyagi et al. 2020, 64-65, male, female, larval exuviae, case, Kyusyu (Fukuoka, Kumamoto, Miyazaki).
Type locality. Hnshu (Hyogo), Japan.
Distribution. Japan (Honshu (central)). Known only from the type locality.
Goera tenuis Ulmer 1927
Goera tenuis Ulmer 1927, 181, male
Nozaki and Tanida 2006, 17-19, male, female, Ishigaki-jima, Iriomote-jima.
Nozaki 2016, 391, 440, male.
Tanida 2016, 125.
Nozaki 2018, 638, male.
Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima, Iriomote-jima), Taiwan.
Goera tungusensis Martynov 1909
Goera tungusensis Martynov 1909, 246-250, male, female
Nozaki and Tanida 2006, 18-20, male, female, Hokkaido (Soya, Rumoi, Sorachi, Shiribeshi).
Nozaki 2016, 390-391, 440, male.
Tanida 2016, 125.
Ito 2017b, 45, Hokkaido (Kamikawa).
Nozaki 2018, 638, 640, 641, male, larva, case.
Ito and Kuhara 2023, 68, Hokkaido (Soya).
Goera sp. 1: Ito et al. 2000, 23, 27-29, Hokkaido (Soya). Reidentified as this species by Nozaki
and Tanida (2006).
Goera sajanensis Martynov 1924, 103-105, male, female, Russia.Synonymized by Lavanidova (1982).
Lepneva 1966, 328-331, larva, Russia.
Type locality. Northern Siberia, Russia.
Distribution. Japan (Hokkaido), Russia (Siberia, continental Far East), Canada (Quebec),
USA (Alaska).
Goera uchina Tanida and Nozaki 2006
Goera sp. GD: Tanida 1997, 449.
Tanida 2003, 379, male, larva.
Goera uchina: Nozaki and Tanida 2006, 8-10, male, female, Okinawa, Ishigaki-jima.
Nozaki 2016, 389, 391, 440, male.
Tanida 2016, 125.
Nozaki 2018, 637, 640, 641, male.
Type locality. Okinawa, Japan.
Distribution. Japan (Okinawa, Ishigaki-jima).
Remarks. Larvae of this species are indistinguishable from those of G. akagiae (Nozaki and Tanida 2006).
Goera sp.
Goera sp.: Shimura and Imasaka 2020, 14-15, 18, female, Kyushu (Fukuoka).
Remarks. Shimura and Imasaka (2020) recorded a female, which is similar to that of G. kyotonis but possibly an undescribed species.
Genus Silo Curtis
A single species, Silo nigricellus Matsumura, is recorded from Hokkaido, but the generic identification is
very doubtful (Nozaki et al. 2000).
Silo? nigricellus Matsumura 1907, Hokkaido
Silo nigricellus Matsumura 1907, 191, adult.
Nozaki et al. 2000, 198.
Tanida 2016, 126.
Type locality. Hokkaido, Japan.
Distribution. Japan (Hokkaido).
Remarks. Only the color, size, and morphology of the antenna were provided in the original description by Matsumura (1907), but they are insufficient even for generic identification (Nozaki et al. 2000). The holotype of this species was not found in the collection of
Matsumura deposited in Systematic Entomology, Faculty of Agriculture, Hokkaido
University (Nozaki et al. 2000).
Subfamily Larcasinae Navas
Only the genus Larcasia belongs to this subfamily.
Genus Larcasia Navas
Two species of this genus were recorded from Japan (Nishimoto et al 1999).
Larcasia akagiae Nishimoto and Tanida 1999
Goerinae sp.: Akagi 1959, 42, larva, male, Shikoku (Ehime). Associated with the adult stage by Nishimoto et al. (1999)
Tsuda and Akagi 1962, 140, larva.
Imania sp. IA: Tanida 1985, 195, comb. Misidentification mentioned by Nishimoto et al. (1999).
Larcasia akagiae: Nishimoto et al. 1999, 426-435, male, female, pupa, larva, egg mass, case, habitat, adult behavior, life history, Honshu (Aichi, Nara).
Nishimoto 2000, 26-28, larva, adult, egg mass, habitat, adult behavior, Honshu (Aichi).
Nozaki et al. 2000, 199.
Nozaki 2005a, 536-538, male, larva, case.
Inazu 2008, 84, 95, 103, Honshu (Hyogo).
Inazu and Nishida 2011, 206, male, larva, case, Honshu (Hyogo).
Inazu 2013b, 58, Honshu (Hyogo).
Miyahara 2014, 1, 4-6, larva, Honshu (Gumma).
Nishimoto 2014, larva).
Horiguchi 2016, 415, Honshu (Tokyo).
Nozaki 2016, 83, 391-392, 440, 451, male, Honshu (Tokyo).
Tanida 2016, 126.
Nojima 2017, Honshu (Okayama).
Nozaki 2018, 638, 642, male, larva, case.
Ohira and Tsutsumi 2018, 26, Honshu (Fukushima).
Type locality. Honshu (Aichi).
Distribution. Japan (Honshu (central), Shikoku).
Larcasia minor Nishimoto 1999
Larcasia minor: Nishimoto et al. 1999, 435-438, male, female, larva, egg mass, case, habitat, adult behavior, life cycle, Honshu (Tochigi).
Nozaki et al. 2000, 199.
Nozaki 2005a, 536-538, male, larva.
Tanida 2016, 126.
Nozaki 2018, 638, 642, male, larva.
Type locality, Honshu (Tochigi).
Distribution. Japan (Honshu (central)).
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