A catalogue of Japanese Trichoptera
Family Hydroptilidae Stephens

Author: Tomiko ITO
E-mail: tobikera[AT]siren.ocn.ne.jp
Last updated: 3 Jan. 2024

In the family Hydroptilidae, Ptilocolepinae Martynov, treated as a subfamily of this family for a long time, is treated as family status in recent time (Holzenthal et al. 2011; Wichard 2013). Ten genera belonging to this family have been recorded from Japan. Hydroptila usuguronis Matsumura 1931 was transferred to the genus Psychomyia Latreille (Psychomyiidae) by Ito et al. (2011).


Genus Allotrichia McLachlan

One species belonging to this genus have been recorded from Japan.

Allotrichia rhynchophyllum Zhou, Yang and Morse 2016

Allotrichia rhynchophyllum Zhou, Ynag and Morse 2016, 206-208, male, northeastern China.
Ito and Shimura 2019, 32, 34-35, male, Honshu, Shikoku.
Ugandatrichia shinshiroensis Ito, Nishimoto and Nishimoto 2018, 492-500, male, female, pupa, larva, case, habitat, Honshu (Aichi, Mie), Shikoku (Kochi). Synomymized by Ito and Shimura (2019).
Ito 2018, 503, 508, 511, male, larva, case.
Type locality. Hei-long-jiang, northeastern China.
Distribution. China (Hei-long-jiang), Japan (Honshu, Shikoku).

Genus Hydroptila Dalman

Twenty-five species belonging to this genus have been recorded from Japan.

Hydroptila asymmetrica Kumanski 1990

Hydroptila asymmetrica Kumanski 1990, 50-51, male, female, North Korea (Kangvon).
Nozaki and Tanida 2007, 245, Honshu (Shizuoka).
Kawase and Morita 2010, 37, Honshu (Shiga).
Ito et al. 2010, 56, Hokkaido (Shiribeshi, Hiyama, Oshima).
Ito et al. 2011, 15-18, male, female, Hokkaido (Kushiro, Ishikari, Shiribeshi, Hiyama, Oshima), Honshu (Iwate, Miyagi, Yamagata, Niigata, Ishikawa, Shizuoka, Aichi, Mie, Shiga, Hyogo, Wakayama, Shimane, Hiroshima), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka, Nagasaki, Kumamoto, Miyazaki, Kagoshima), Tsushima, Amami-Oshima, Okinawa-jima, Ishigaki-jima.
Moriya and Kubota 2012, 134, Honshu (Kanagawa).
Ito 2013a, 50, 80, Hokkaido (Kushiro).
Ito and Nagayasu 2014, 9, Hokkaido (Sorachi).
Nishimoto and Nishimoto 2014, 62, Honshu (Aichi).
Ito 2015, 15, Amami-Oshima, Tokunoshima, Okinawa-jima.
Shimura et al. 2015, 45, Amami-Oshima.
Tanida 2016, 70.
Ito 2017a, 3, Honshu (Fukui).
Ito 2017e, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 25, Honshu (Shiga).
Kuhara and Ito 2017, 14, 16, Yakushima.
Nojima 2017, 111, Honshu (Okayama).
Saito et al. 2017, 183, Honshu (Chiba).
Ito 2018, 502, 505, male.
Morita 2018, 50, Honshu (Mie).
Nozaki 2018, 927, Honshu (Kanagawa).
Nakatani and Ito 2019, 162, Hokkaido (Kushiro).
Yamamoto et al. 2019, 69, Shikoku (Ehime).
Morita 2020a, 18, Honshu (Mie).
Morita 2020b, 43, Honshu (Mie).
Morita 2020c, 80, Honshu (Mie).
Tezuka et al. 2021, 42, Honshu (Kanagawa).
Yamamoto et al. 2021, 126, Shikoku (Ehime).
Ito and Kuhara 2023, 60, Hokkaido (Kamikawa, Rumoi).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Yakushima, Amami-Oshima, Tokunoshima, Okinawa-jima, Ishigaki-jima), Russia (South Primorye), North Korea.

Hydroptila botosaneanui Kumanski 1990

Hydroptila botosaneanui Kumanski 1990, 48-49, male, female, North Korea (Kangvon).
Ito et al. 2011, 20-22, male, female, Hokkaido (Kushiro, Ishikari), Honshu (Miyagi, Yamagata, Yamanashi, Gifu, Shizuoka, Shiga, Nara, Hyogo, Hiroshima).
Ito 2013a, 50, 80, Hokkaido (Kushiro).
Inazu 2013, 55, Honshu (Hyogo).
Tanida 2016, 70.
Ito 2017a, 3, Honshu (Fukui).
Ito 2017e, 39, Hokkaido (Kamikawa).
Kawase et al. 2017, 25, Honshu (Shiga).
Ito 2018, 502, 505, male.
Nozaki et al. 2019, 167, phenology, Honshu (Nagano).
Ito 2021, 346-347, larva, case, Hokkaido (Kushiro).
Type locality. Kangvon, North Korea.
Distribution. Japan (Hokkaido, Honshu), North Korea, Russia (South Primorye).
Remarks. Ito (2021) said that this species belonged to the H. teneoides Species Group Marshall 1979.

Hydroptila chinensis Xue and Yang 1990

Hydroptila chinensis Xue and Yang 1990, 126-127, male, female, China (Heilongjiang).
Morse et al. 2001, 97-116.
Ito et al. 2004, 12, Hokkaido (Kushiro).
Katsuma 2006, 35, Honshu (Ibaraki).
Ito et al. 2011, male, female, 5-7, Hokkaido (Nemuro, Kushiro, Soya, Tokachi).
Ito 2013a, 50, 80, Hokkaido (Kushiro, Tokachi).
Ito and Nagayasu 2014, 9, Hokkaido (Sorachi).
Tanida 2016, 70.
Ito 2017a, 3, Honshu (Fukui).
Ito 2017e, 39, Hokkaido (Kamikawa).
Ito 2018, 502, 505, male.
Ito and Kuhara 2023, 60, Hokkaido (Soya, Kamikawa, Rumoi).
Kawase and Kato 2023, 6-7, Honshu (Niigata).
Type locality. Heilongjiang, China.
Distribution. Japan (Hokkaido, Honshu (central)), Russia (Khabarovsk, South Primorye), China (Heilongjiang).

Hydroptila coreana Kumanski 1990

Hydroptila coreana Kumanski 1990, 52-54, male, female, North Korea (Kangvon).
Mey and Nozaki 2006, 24, Honshu (Shiga).
Nakamura 2010, 238, Honshu (Hiroshima).
Kawase and Morita 2010, 37, Honshu (Shiga).
Ito et al. 2010, 56, Hokkaido (Shiribeshi, Oshima).
Ito et al. 2011, 16, 17-18, male, female, Hokkaido (Kushiro, Ishikari, Hiyama, Oshima), Honshu (Iwate, Miyagi, Akita, Yamagata, Gifu, Aichi, Shiga, Kyoto, Hyogo, Nara, Shimane, Okayama, Hiroshima, Yamaguchi), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Oita, Miyazaki).
Nozaki 2011, 4, Honshu (Aichi).
Ito 2013a, 50, 80, Hokakido (Kushiro).
Nozaki 2013, 18, Honshu (Aichi).
Nishimoto and Nishimoto 2014, 62, Honshu (Aichi).
Morita 2015, 17, Honshu (Mie).
Tanida 2016, 70.
Ito 2017e, 39, Hokkaido (Kamikawa).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Nojima 2017, 111-112, Honshu (Okayama).
Ito 2018, 502, 505, male.
Katsuma 2020, 15, Kyushu (Oita).
Morita 2020c, 80, Honshu (Mie).
Yamamoto et al. 2021, 71, Shikoku (Ehime).
Ito and Kuhara 2023, 60, Hokkaido (Soya, Rumoi).
Type locality. Kangvon, North Korea.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (Khabarovsk), North Korea.

Hydroptila dampfi Ulmer 1929

Hydroptila dampfi Ulmer, 1929, 264-266, figs. 10-12, male, Germany.
Ito et al. 2011, male, female, 9, 11-12, Hokkaido (Nemuro, Kushiro, Soya, Ishikari, Iburi, Oshima), Honshu (Yamanashi, Shizuoka, Shiga).
Ito 2013a, 50, 80, Hokkaido (Nemuro, Kushiro, Tokachi).
Tanida 2016, 70.
Kawase et al. 2017, 25, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga, Kyoto).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 502, 503, 505, male, larva, case.
Ito 2021, 342-343, larva, case, Hokkaido (Kushiro, Iburi).
Ito and Kuhara 2023, 60, Hokkaido (Soya).
Hydroptila itoi Kobayashi 1977, 5, 12, male, female, Hokkaido (Iburi). Synonymized by Ito et al. (2011)
Ito and Kawamura 1980, 113-118, pupa, larva, case, habitat, life cycle, Hokkaido (Iburi).
Ito and Kawamula 1984, 315, new synonym.
Ito et al. 1993, 142.
Ito et al. 1999, 23, 25, Hokkaido (Kushiro).
Kuhara et al. 2000, 248, Hokkaido (Iburi).
Ito 2005, 442, 444, 446, male, larva, case.
Torii and Hattori 2006, 34, Honshu (Shizuoka).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Morita 2008a, 2, 4, Honshu (Mie).
Ohtaka et al. 2008, 60, Honshu (Aomori).
Kawase and Morita 2010, 37, Honshu (Shiga).
Ito et al. 2010, 56, Hokkaido (Oshima).
Oxyethira ezoensis Kobayashi 1977, 5-6, 13, male. Synonymized with H. itoi Kobayashi 1977 by Ito and Kawamula (1984).
Type locality. Germany.
Distribution. Japan (Hokkaido, Honshu), China (Heilongjiang, Yunnan, Jiangsu, Henan, Hubei, Sichuan, Guizhou), Russia (Caucasus, South Primorye), Europe.
Remarks. Ito (2021) said that this species belonged to the H. pulchricornis Species Group Marshall 1979.

Hydroptila demersa Ito and Sasaki 2023

Hydroptila demersa Ito and Sasaki, in Ito et al., 2023, 145-149, male, larva, case, habitat, Japan (Ogasawara Islands).
Type locality. Chichi-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Chichi-jima, Ani-jima).
Remarks. This species belongs to the H. ogasawaraensis Species Group Ito 2023 (Ito et al. 2023, see the remarks of H. ogasawaraensis).

Hydroptila dorsoprocessuata Botosaneanu 1993

Hydroptila dorsoprocessuata Botosaneanu 1993, 187-188, male, Russia (South Siberia).
Ito and Shimura, 2019, 30-32, male, female, Honshu (Aichi, Mie).
Type locality. Siberia, Russia.
Distribution. Japan (Honshu), Russia (Siberia, South Primorye).

Hydroptila hahajima Ito and Sasaki 2023

Hydroptila hahajima Ito and Sasaki, in Ito et al., 2023, 157-159, male, female, larva, case, habitat, Japan (Ogasawara Islands).
Type locality. Haha-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Haha-jima).
Remarks. This species belongs to the H. ogasawaraensis Species Group Ito 2023 (Ito et al. 2023, see the remarks of H. ogasawaraensis).

Hydroptila hattorii Ito 2022

Hydroptila hattorii Ito 2022, 138-139, male, Honshu (Iwate).
Type locality. Iwate, Honshu, Japan.
Distribution. Japan (Honshu).

Hydroptila introspinata Zhou and Sun 2009

Hydroptila introspinata Zhou and Sun in Zhou et al., 2009, 906-908, male, northeastern China.
Ito and Shimura 2019, 30-32, male, female, Hokkaido (Ishikari, Shiribeshi), Honshu (Iwate), Shikoku (Ehime, Kochi), Kyushu (Miyazaki).
Yamamoto et al. 2019, 70, Shikoku (Ehime).
Type locality. Hei-long-jiang, China.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), China (Hei-long-jiang), Korea.

Hydroptila ishiura Ito and Sasaki 2023

Hydroptila ishiura Ito and Sasaki, in Ito et al., 2023, 151-154, male, female, larva, case, habitat, Japan (Ogasawara Islands).
Type locality. Chichi-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Chichi-jima, Ani-jima).
Remarks. This species belongs to the H. ogasawaraensis Species Group Ito 2023 (Ito et al. 2023, see the remarks of H. ogasawaraensis).

Hydroptila kakidaensis Nozaki and Tanida 2007

Hydroptila kakidaensis Nozaki and Tanida 2007, 245-246, male, female, Honshu (Shizuoka).
Ito et al. 2011, 19-21, male, female, Honshu (Nagano, Shizuoka, Mie), Kyushu (Kumamoto, Miyazaki, Kagoshima).
Tanida 2016, 70.
Ito 2018, 502, 505, male.
Morita 2018, 50, Honshu (Mie).
Yamamoto et al. 2019, 69, Shikoku (Ehime).
Ito 2021, 344-346, larva, case, Honshu (Shizuoka), Shikoku (Ehime).
Yamamoto and Nozaki 2021, 126, 132, adult appearance months, Shikoku (Ehime).
Type locality. Shizuoka, Honshu, Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. Ito (2021) said that this species belonged to the H. teneoides Species Group Marshall 1979.

Hydroptila nagahama Ito and Sasaki 2023

Hydroptila nagahama Ito and Sasaki, in Ito et al., 2023, 157-160, male, female, larva, case, habitat, Japan (Ogasawara Islands).
Type locality. Haha-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Haha-jima).
Remarks. This species belongs to the H. ogasawaraensis Species Group Ito 2023 (Ito et al. 2023, see the remarks of H. ogasawaraensis).

Hydroptila nago Ito 2019

Hydroptila nago Ito in Ito and Shimura 2019, 27-28, male, female, Japan (Ryukyu Islands: Okinawa-jima).
Type locality. Okinawa-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands).

Hydroptila nanseiensis Ito 2011

Hydroptila nanseiensis Ito et al. 2011, 14-15, male, female, Japan (Nansei Islands: Okinawa-jima, Ishigaki-jima, Iriomote-jima).
Ito 2015, 15, Okinawa-jima, Ishigaki-jima, Iriomote-jima.
Tanida 2016, 70.
Ito 2018, 502, 505, male.
Ito 2021, 347-348, larva, case, Ryukyu (Iriomote-jima).
Type locality. Ishigaki-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands).
Remarks. Ito (2021) said that the species group of this species was unknown.

Hydroptila ogasawaraensis Ito 2011

Hydroptila ogasawaraensis Ito, in Ito et al. 2011, 6-8, male, female, Japan (Ogasawara Islands).
Tanida 2016, 70.
Ito 2018, 501, 505, male.
Ito et al. 2023, 154, male, female, larva, case, habitat, Ogasawara Islands.
Type locality. Chichi-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Chichi-jima).
Remarks. Ito et al. (2023) proposed a new species group name, H. ogasawaraensis Species Group Ito, for this species and other five new species, H. demersa, H. hahajioma, H. ishiura, H. nagahama and H. tokoyo, all endemic in the Ogasawara Islands.

Hydroptila oguranis Kobayashi 1974

Hydroptila oguranis Kobayashi 1974, 68-69, Pl. 26, male, Honshu (Kanagawa).
Ito et al. 1993, 142.
Ito et al. 1998, 10, 17, larva, case, Hokkaido (Nemuro).
Ito et al. 2000, 29, Hokkaido (Ishikari).
Kuhara et al. 2000, 248, Hokkaido (Ishikari, Iburi).
Ishiwata et al. 2005, 110, 242, Honshu (Kanagawa).
Ito 2005, 441, 446, male.
Nagayasu et al. 2005, 54, Hokkaido (Ishikari).
Nozaki 2005, 1234, Honshu (Kanagawa).
Ito and Kosugi 2007, 51, Hokkaido (Kushiro).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Tanida 2007, 245-246, Honshu (Shizuoka).
Ito et al. 2010, 56, Hokkaido (Oshima).
Ito et al. 2011, 8-11, male, female, Hokkaido (Nemuro, Kushiro, Tokachi, Ishikari, Iburi, Oshima, Oshima), Honshu (Tokyo, Kanagawa, Shizuoka, Shiga, Hyogo, Wakayama), Shikoku (Tokushima, Kochi), Kyushu (Nagasaki, Kumamoto, Miyazaki, Kagoshima), Tsushima, Amami-Oshima. Kawase and Hayashi, 2011b, 191, Honshu (Shimane).
Morita and Kubota 2012, 135, Honshu (Kanagawa).
Ito 2013a, 50, 80, Hokkaido (Nemuro, Kushiro, Tokachi).
Shimura et al. 2015, 45, Amami-Oshima.
Ito and Kuhara 2016, 15, Hokkaido (Kushiro).
Tanida 2016, 70.
Kawase et al. 2017, 25, Honshu (Shiga).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 501, 503, 505, male, larva, case.
Tsuruda 2018, 26, Honshu (Tokyo).
Yamamoto et al. 2019, 70, Shikoku (Ehime).
Taira 2020, 8, Honshu (Nara).
Katsuma 2020, 15, Kyushu (Oita).
Morita 2018, 50, Honshu (Mie).
Ito 2021, 342, 344, larva, case, Hokkaido (Nemuro, Ishikari, Iburi).
Tezuka et al. 2021, 127, Shikoku (Ehime).
Yamamoto and Nozaki 2021, 127, Shikoku (Ehime).
Ito and Kuhara 2023, 60, Hokkaido (Kamikawa).
Type locality. Kanagawa, Honshu, Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Amami-Oshima).
Remarks. Ito (2021) said that this species belonged to the H. pulchricornis Species Group Marshall 1979.

Hydroptila parapiculata Yang and Xue 1994

Hydroptila parapiculata Yang and Xue 1994, 9, male, China (Sichuan).
Kobayashi et al. 2017, 19, Honshu (Shiga, Kyoto).
Ito 2018, 502, 505, male.
Ito and Shimura 2019, 32-33, male, Honshu (Shiga, Kyoto).
Type locality. Sichuan, China.
Distribution. Japan (Honshu), China (Anhui, Fujian, Sichuan, Guizhou, Guangxi).

Hydroptila phenianica Botosaneanu 1970

Hydroptila matsuii Kobayashi 1974, 67-68, Pl. 25, male, Honshu (Nagano). Synonymized by Nozaki and Tanida (2007).
Nozaki and Gyotoku 1990, 16, Kyushu (Fukuoka).
Kadowaki 1998, 9, Honshu (Shimane).
Nishimoto and Nishimoto 1998, 150, Honshu (Aichi).
Kawase et al. 2004, 108, Honshu (Ibaraki).
Nozaki and Nakamura 2002, 167, Honshu (Hiroshima).
Nozaki 2004, 1234, Honshu (Kanagawa).
Ishiwata et al. 2005, 110, 243-244, Honshu (Kanagawa).
Ito 2005, 441, 446, male.
Nozaki and Nakamura 2005, 219, Honshu (Shimane).
Kameyama et al. 2006, 92, Honshu (Hiroshima).
Torii and Hattori 2006, 34, Honshu (Shizuoka).
Katsuma 2007, 17, Honshu (Tochigi).
Morita 2008a, 4, Honshu (Mie).
Hydroptila phenianica Botosaneanu 1970, 290-291, male, female, North Korea (Phjongjang).
Nozaki and Tanida 2007, 246, Honshu (Shizuoka).
Katsuma 2008, 57, Honshu (Ibaraki).
Nozaki 2010, 22, Honshu (Aichi).
Katsuma 2010, 84, Honshu (Ibaraki).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 238, Honshu (Hiroshima).
Kawase and Morita 2010, 37, Honshu (Mie, Shiga).
Ito et al. 2010, 56, Hokkaido (Hiyama).
Ito et al. 2011, 2-5, male, female, Hokkaido (Soya, Ishikari, Oshima), Honshu (Aomori, Iwate, Akita, Yamagata, Ibaraki, Tochigi, Tokyo, Kanagawa, Niigata, Ishikawa, Nagano, Shizuoka, Aichi, Mie, Shiga, Kyoto, Hyogo, Tottori, Shimane, Okayama, Yamaguchi), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Fukuoka, Nagasaki, Kumamoto, Oita, Miyazaki), Tsushima.
Kawase and Hayashi, 2011a, 172, Honshu (Shimane).
Kawase and Hayashi, 2011b, 191, Honshu (Shimane).
Nozaki 2011, 4, Honshu (Aichi).
Morita 2012, 249, Honshu (Mie).
Nozaki 2013, 18, Honshu (Aichi).
Ito and Nagayasu 2014, 9, Hokkaido (Sorachi).
Nishimoto and Nishimoto 2014, 62-63, male, Honshu (Aichi).
Shimura 2015, 53, Fukue-jima.
Shimura et al. 2015, 45, Amami-Oshima.
Katsuma 2016, 47, Honshu (Ibaraki).
Nozaki et al. 2016, 324, Honshu (Niigata).
Tanida 2016, 71.
Yamamoto and Ito 2016, 38, 44, Shikoku (Ehime).
Ito 2017a, 3, Honshu (Fukui).
Ito 2017e, 39-40, male, Hokkaido (Kamikawa).
Kawase et al. 2017, 25-26, Honshu (Shiga).
Nojima 2017, 112, Honshu (Okayama).
Ohira and Tsutsumi 2017, 24, Honshu (Fukushima).
Ito 2018, 501, 505, male.
Nozaki 2018, 927, Honshu (Kanagawa).
Tsuruda 2018, 26, Honshu (Tokyo).
Nozaki et al. 2019, 168, phenology, Honshu (Nagano).
Yamamoto et al. 2019, 70, Shikoku (Ehime).
Katsuma 2020, 15, Kyushu (Oita).
Kimura and Kuranishi 2020, 157, Honshu (Kyoto).
Morita 2020b, 43, Honshu (Mie).
Morita 2020c, 80, Honshu (Mie).
Ito 2021, 340-341, larva, case, Honshu (Hyogo, Hiroshima), Kyushu (Fukuoka), Tsushima.
Tezuka et al. 2021, 42, Honshu (Kanagawa).
Yamamoto and Nozaki, 2021, 126, 132, adult appearance months, Shikoku (Ehime).
Ito and Kuhara 2023, 60, Hokkaido (Soya, Rumoi).
Kawase and Kato 2023, 7, Honshu (Niigata).
Type locality. Phjongjang, North Korea.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Fukue-jima, Amami-Oshima), Russia (Khabarovsk, South Primorye), North Korea (Phjongjang).
Remaks. Ito (2021) said that this species belonged to the H. pulchricornis Species Group Marshall 1979.

Hydroptila pseudseirene Ito 2015

Hydroptila pseudseirene Ito 2015, 9-11, male, female, Ryukyu (Ishigaki-jima, Irimote-jima).
Tanida 2016, 71.
Ito 2018, 502, 505, male.
Type locality. Ishigaki-jima, Ryukyu, Japan.
Distribution. Japan (Ryukyu: Ishigaki-jima, Iriomote-jima).

Hydroptila spinosa Arefina and Armitage 2003

Hydroptila spinosa Arefina and Armitage 2003, 15-16, figs. 1-2, male, female, Russian Far East (Sakhalin).
Ito et al. 2010, 56, Hokkaido (Shiribeshi, Hiyama, Oshima).
Ito et al. 2011, male, female, 12-14, Hokkaido (Kushiro, Tokachi, Soya, Kamikawa, Ishikari, Hidaka, Shiribeshi, Hiyama, Oshima), Honshu (Aomori, Miyagi, Fukushima, Saitama, Tokyo, Kanagawa, Niigata, Nagano, Shizuoka, Mie, Shiga, Wakayama, Tottori), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Nagasaki, Miyazaki).
Moriya and Kubo 2012, 135, Honshu (Kanagawa).
Ito 2013a, 50, 80, Hokkaido (Kushiro, Tokachi).
Ito and Nagayasu 2014, 9, Hokkaido (Sorachi).
Nozaki 2016, 71, 321, 323, adult, Shikoku (Kochi).
Tanida 2016, 71.
Ito 2017a, 3, Honshu (Fukui)
Ito 2017e, 40, Hokkaido (Kamikawa).
Kawase et al. 2017, 26, Honshu (Shiga).
Ito 2018, 502, 505, male.
Morita 2018, 50, Honshu (Mie).
Nozaki 2018, 927, Honshu (Kanagawa).
Morita 2020c, 80, Honshu (Mie).
Shimura and Imasaka 2020, 13, Kyushu (Fukuoka).
Ito 2023b, 3, Honshu (Shizuoka).
Ito and Kuhara 2023, 60-61, Hokkaido (Soya, Kamikawa).
Type locality. Sakhalin, Russian Far East.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (Sakhalin).
Remarks. This species is very similar to H. introspinata Zhou and Sun, and some specimens of H. spinosa recorded in Ito et al. (2011) are incorrect.

Hydroptila spiralis Ito 2015

Hydroptila spiralis Ito 2015, 11-13, male, female, Ryukyu (Okinawa-jima, Amami-o-shima).
Tanida 2016, 71.
Ito 2018, 502, 505, male.
Type locality. Okinawa-jima, Ryukyu, Japan.
Distribution. Japan (Ryukyu: Okinawa-jima, Amami-o-shima).

Hydroptila thuna Oláh 1989

Hydroptila thuna Oláh 1989, 281-282, fig. 26, male, Vietnam.
Ito et al. 2011, 5, male, female, Yonaguni-jima, Ishigaki-jima.
Ito 2015, 13, Ishigaki-jima, Iriomote-jima.
Tanida 2016, 71.
Ito 2018, 501, 505, male.
Hydroptila phenianica: Shimura, 2010, 49, Japan (Yonaguni-jima). Misidentification mentioned by Ito et al. (2011).
Type locality. Vietnam.
Distribution. Japan (Yonaguni-jima, Ishigaki-jima, Iriomote-jima), Russia (Far East, continental part), China (Guangdong, Hainan, Henan, Hubei, Jiangsu, Anhui, Jiangxi, Fujian, Guangxi, Yunnan, Hong Kong), Taiwan, Raos, Thailand, Vietnum, Nepal, Sumatora.

Hydroptila tokoyo Ito and Sasaki 2023

Hydroptila tokoyo Ito and Sasaki, in Ito et al., 2023, 154-157, male, habitat, Japan (Ogasawara Islands).
Type locality. Chichi-jima, Ogasawara Islands, Japan.
Distribution. Japan (Ogasawara Islands: Chichi-jima).
Remarks. This species belongs to the H. ogasawaraensis Species Group Ito 2023 (Ito et al. 2023, see the remarks of H. ogasawaraensis).

Hydroptila yaeyamensis Ito 2015

Hydroptila yaeyamensis Ito 2015, 13, male, female, Ryukyu (Ishigaki-jima, Irimote-jima).
Tanida 2016, 71.
Ito 2018, 502, 505, male.
Type locality. Ishigaki-jima, Ryukyu, Japan.
Distribution. Japan (Ryukyu: Ishigaki-jima, Iriomote-jima).

Genus Microptila Ris

Three species belonging to this genus have been recorded from Japan.

Microptila genka Ito 2017

Microptila genka Ito 2017b, 107-108, figs. 2, 3, 5, male, female, Ryukyu Islands (Okinawa-jima, Ishigaki-jima).
Ito 2018, 504, 511, male.
Type locality. Okinawa-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Okinawa-jima, Iriomote-jima).

Microptila nakama Ito 2017

Microptila nakama Ito 2017b, 107-111, figs. 3-5, male, Ryukyu Islands (Iriomote-jima).
Ito 2018, 504, 511, male.
Type locality. Iriomote-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Iriomote-jima).

Microptila orienthula Kjærandsen and Ito 2009

Microptila orienthula Kjærandsen and Ito 2009, 177-180, fig. 1, male, female, Hokkaido (Kamikawa, Oshima).
Nishimoto and Nishimoto 2014, 63, Honshu (Aichi).
Tanida 2016, 71.
Ito 2017b, 105-107, figs. 1, 4, 5, male, female, Hokkaido (Kamikawa, Oshima). Honshu (Niigata, Toyama, Shizuoka, Aichi, Shiga), Shikoku (Kochi), Kyushu (Nagasaki), Yakushima.
Kuhara and Ito 2017, 14, 16, Yakushima.
Ito 2018, 502, 511, male.
Ito 2023a, 445-450, larva, case, microhabitat, Honshu (Shizuoka).
Ito 2023b, 3-4, adult appearance period, Honshu (Shizuoka).
Type locality. Oshima, Hokkaido, Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Yakushima).
Remarks. Type locality described by Tanida (2016) is incorrect.

Genus Orthotrichia Eaton

Four species belonging to this genus have been recorded from Japan. Records of Orthotrichia tetensii Kolb. 1927 in Gifu (Iwata 1927) is incorrect, because Iwata's figures and habitat description are apparently those of the genus Hydroptila.

Orthotrichia coreana Ito and Park 2016

Orthotrichia coreana Ito and Park 2016, male, female, Korea.
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 506, 511, male.
Katsuma 2020, 15, Kyushu (Oita).
Type locality. Gyeongsangbuk-do, Korea.
Distribution. Japan (Honshu, Kyushu), Korea.

Orthotrichia costalis (Curtis 1834)

Hydroptila costalis Curtis, 1834, 218, male.
Orthotrichia costalis: Malicky, 1983, 54-55, male, female.
Ito et al. 2004, 12-13, male, female, Hokkaido (Kushiro).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Ito and Kosugi 2007, 51, Hokkaido (Kushiro).
Ito et al. 2010, 57, Hokkaido (Oshima).
Ito 2013a, 50, 80, Hokkaido (Kushiro).
Ito 2013b, 40, 42-43, 45, male, female, Hokkaido (Soya, Iburi), Honshu (Miyagi, Shiga, Kyoto).
Ito and Kuhara 2016, 15, Hokkaido (Kushiro).
Tanida 2016, 71.
Kawase et al. 2017, 26, Honshu (Shiga).
Kobayashi et al. 2017, 17, 22, Honshu (Shiga).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 506, 511, male.
Ito and Kuhara 2023, 61, Hokkaido (Soya, Kamikawa, Rumoi).
Type locality. Unknown.
Distribution. Japan (Hokkaido, Honshu, Kyushu), Palearctic.

Orthotrichia iriomotensis Ito 2013
Orthotrichia iriomotensis Ito, 2013b, 43-46, male, female, Iriomote-jima.
Tanida 2016, 71.
Ito 2018, 506, 511, male.
Type locality. Iriomote-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Iriomote-jima).

Orthotrichia tragetti Mosely 1930

Orthotrichia tragetti Mosely 1930, 237, 247-249, male.
Fischer 1961, 138.
Fischer 1971, 267.
Malicky 1983, 54-55, male, female.
Ito et al. 2004, 12-14, male, female, Hokkaido (Kushiro).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Ito and Kosugi 2007, 51, Hokkaido (Kushiro).
Ohtaka et al. 2008, 60, Honshu (Aomori).
Nozaki 2010, 22, Honshu (Aichi).
Ito et al. 2010, 57, Hokkaido (Shiribeshi, Oshima).
Nozaki 2011, 4, Honshu (Aichi).
Ito 2013a, 50-51, 80, Hokkaido (Kushiro).
Ito 2013b, 39-41, 45, male, female, Hokkaido (Soya), Honshu (Miyagi, Niigata, Shiga), Kyushu (Fukuoka, Kagoshima), Amami-Oshima, Minami-daito-jima.
Nozaki 2013, 18, Honshu (Aichi).
Tanida 2016, 72.
Ito 2017a, 3, Honshu (Fukui).
Kawase et al. 2017, 26, Honshu (Shiga).
Kobayashi et al. 2017, 17, 22, Honshu (Shiga, Kyoto).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 506, 511, male.
Nozaki 2019, 43, adult, larva, Honshu (Aichi).
Yamamoto and Nozaki 2021, 127, Shikoku (Ehime).
Ito and Kuhara 2023, 61, Hokkaido (Soya).
Type locality. England.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Amami-Oshima, Minami-daito-jima), Palearctic.

Genus Oxyethira Eaton

Ten species belonging to this genus have been recorded from Japan.

Oxyethira acuta Kobayashi 1977

Oxyethira acuta Kobayashi 1977, 6-7, 13, male, Hokkaido (Iburi).
Ito and Kawamula 1984, 313-317, pupa, larva, case, life cycle, Hokkaido (Iburi).
Ito et al. 1993, 142.
Kuhara et al. 2000, 248-249, Hokkaido (Iburi).
Ito 2005, 442, 444, larva, case.
Oláh and Ito 2013, 32, male, Hokkaido (Kushiro, Ishikari, Iburi).
Tanida 2016, 72.
Ito and Oláh 2017, 3-6, 23, male, female, Hokkaido (Kushiro, Soya, Ishikari, Iburi), Honshu (Aomori, Okayama).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 503, 506, 511, male, larva, case.
Ito and Kuhara 2023, 61, Hokkaido (Soya).
Oxyethira sp.: Ohtaka et al. 2008, 60, Honshu (Aomori). Mentioned by Ito and Olah (2017).
Type locality. Iburi, Hokkaido, Japan.
Distribution. Japan (Hokkaido, Honshu).

Oxyethira angustella Martynov 1933

Oxyethira angustella Martynov, 1933, 139-140, venation of fore wing, general feature of antenna, palpus, head and thorax of female, Honshu (Shinano = Nagano).
Ito et al. 1993, 142.
Tanida 2016, 72.
Ito and Oláh 2017, 6-8, 23, male, female, Hokkaido (Nemuro, Kushiro, Ishikari, Iburi), Honshu (Yamagata, Nagano).
Ito 2018, 506, 509, male.
Nozaki et al. 2019, 168, 173, phenology, Honshu (Nagano).
Ito and Kuhara 2023, 61, Hokkaido (Soya).
Oxyethira kakida Oláh and Ito 2013, 30-31, male, Hokkaido (Nemuro, Kushiro, Tokachi, Ishikari), Honshu (Yamagata, Tokyo, Shizuoka). Synonymized by Ito and Oláh (2017).
Oxyethira josifovi: Nozaki and Tanida 2007, 246, Honshu (Shizuoka). Misidentification, mentioned by Oláh and Ito (2013).
Oxyethira datra: Malicky and Chantaramongkol 2007, 1030, Hokkaido (Ishikari). Misidentification, mentioned by Oláh and Ito (2013).
Type locality. Nagano, Honshu, Japan.
Distribution. Japan (Hokkaido, Honshu).

Oxyethira chitosea Oláh and Ito 2013

Oxyethira chitosea Oláh and Ito 2013, 38-39, male, Hokkaido (Kushiro, Ishikari).
Ito and Kuhara 2016, 15, Hokkaido (Okhotsk).
Tanida 2016, 72.
Ito 2017a, 3, Honshu (Fukui).
Ito and Oláh 2017, 8-10, 23, male, female, Hokkaido (Nemuro, Kushiro, Ishikari), Honshu (Fukui).
Ito 2018, 506, 509, male.
Type locality. Ishikari, Hokkaido, Japan.
Distribution. Japan (Hokkaido, Honshu).

Oxyethira hiroshima Oláh and Ito 2013

Oxyethira hiroshima Oláh and Ito 2013, 35-36, male, Honshu (Hiroshima).
Tanida 2016, 72.
Ito and Oláh 2017,10-12, 23, male, female, Honshu (Aomori, Shizuoka, Shiga, Okayama)
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 506, 509, male.
Morita 2018, 51, Honshu (Mie).
Type locality. Hiroshima, Honshu, Japan.
Distribution. Japan (Honshu).

Oxyethira mekunna Oláh and Ito 2013

Oxyethira mekunna Oláh and Ito 2013, 36-37, male, Hokkaido (Tokachi, Sorachi, Shiribeshi).
Tanida 2016, 72.
Ito and Oláh 2017, 12-14, 23, male, female, Hokkaido (Tokachi, Kamikawa, Sorachi, Shiribeshi).
Ito 2017e, 40, Hokkaido (Kamikawa).
Ito 2018, 506, 509, male.
Type locality. Shiribeshi, Hokkaido, Japan.
Distribution. Japan (Hokkaido).

Oxyethira miea Oláh and Ito 2013

Oxyethira miea Oláh and Ito 2013, 42-43, male, Honshu (Mie).
Tanida 2016, 72.
Ito and Oláh 2017, 14-16, 23, male, female, Honshu (Shizuoka, Aichi, Shiga).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 506, 509, male.
Type locality. Mie, Honshu, Japan.
Distribution. Japan (central Honshu), Korea.
Remarks. This species was recorded from Korea (Park et al. 2018).

Oxyethira okinawa Oláh and Ito 2013

Oxyethira okinawa Oláh and Ito 2013, 43-44, male, Okinawa (Okinawa-jima).
Tanida 2016, 72.
Kuhara and Ito 2017, 14, 16, Yakushima.
Ito and Oláh 2017, 16-18, 23, male, female, Yakushima, Amami-Oshima, Okinawa-jima.
Ito 2018, 506, 509, male.
Type locality. Okinawa-jima, Okinawa, Japan.
Distribution. Japan (Yakushima, Amami-Oshima, Okinawa-jima).

Oxyethira ozea Oláh and Ito 2013

Oxyethira ozea Oláh and Ito 2013, 41-42, male, Honshu (Gunma).
Tanida 2016, 72.
Ito and Oláh 2017, 18-19, 23, male, Honshu (Gunma).
Ito 2018, 506, 509, male.
Type locality. Gunma, Honshu, Japan.
Distribution. Japan (central Honshu).

Oxyethira shumari Ito and Oláh 2017

Oxyethira shumari Ito amd Oláh 2017, 20-23, male, female, Hokkaido (Kamikawa).
Ito 2017e, 40, Hokkaido (Kamikawa).
Ito 2018, 506, 509, male.
Ito and Kuhara 2023, 61, Hokkaido (Kamikawa).
Type locality. Kamikawa, Hokkaido, Japan.
Distribution. Japan (Hokkaido).

Oxyethira tsuruga Ito and Oláh 2017

Oxyethira tsuruga Ito and Oláh 2017, 19-21, 23, male, female, Hokkaido (Kushiro), Honshu (Fukui).
Ito 2018, 506, 509, male.
Morita 2020b, 43, Honshu (Mie).
Type locality. Fukui, Honshu, Japan.
Distribution. Japan (Hokkaido, Honshu).

Genus Plethus Hagen

One species belonging to this genus has been recorded from Japan.

Plethus ukalegon Malicky and Chantaramongkol 2007

Plethus ukalegon Malicky and Chantaramongkol 2007, 1047, 1092, male, Taiwan.
Ito and Saito 2016, 466-476, male, female, pupa, larva, case, habitat, Ryukyu Islands (Iriomote-jima, Ishigaki-jima).
Ito 2018, 503, 506-507, 511, male.
Type locality. Taiwan.
Distribution. Japan (Ryukyu Islands: Iriomote-jima, Ishigaki-jima), Taiwan.

Genus Pseudoxyethira Schmid

Genus Pseudoxyethira Schmid was revised recently to replace Scelotrichia Ulmer 1951, because the latter genus name is preoccupied by Scelotrichia Reuter 1890, in Hemiptera (Zhou et al. 2016). Three species belonging to this genus have been recorded from Japan.

Pseudoxyethira ishiharai (Utsunomiya 1994)

Scelotrichia ishiharai Utsunomiya 1994, 345-348, male, female, larva, case, habitat, Shikoku (Kochi).
Ohkawa and Ito 2002, 449-458, 658-659, male, female, pupa, larva, case, feeding habit, habitat, Hokkaido (Ishikari, Shiribeshi), Honshu (Yamagata, Tokyo, Kanagawa, Nagano, Gifu, Shizuoka, Aichi), Shikoku (Ehime, Kochi), Kyushu (Nagasaki, Miyazaki).
Kuhara 2001, 18, Hokkaido (Shiribeshi).
Ito 2005, 442.
Katsuma 2006, 35, Honshu (Ibaraki).
Satake and Kuranishi 2007, 282, Izu Islands (Mikura-jima).
Ito et al. 2010, 57, Hokkaido (Hiyama, Oshima).
Kuhara 2011, 52, 72, flight season, Hokkaido (Sorachi).
Nozaki 2016, 321, adult.
Tanida 2016, 73.
Kuhara and Ito 2017, 14, 16, Yakushima.
Pseudoxyethira ishiharai: Ito 2017c, 195-196, 199, male, habitat, Hokkaido (Ishikari, Shiribeshi, Hiyama, Oshima), Honshu (Ishikawa, Yamanashi, Shizuoka, Shiga, Okayama, Shimane), Shikoku (Tokushima, Ehime), Kyushu (Nagasaki, Oita), Amami-Oshima, Okinawa-jima, Ishigaki-jima, Iriomote-jima.
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 507, 511, male.
Type locality. Kochi, Shikoku, Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Izu Islands, Yakushima, Amami-Oshima, Okinawa-jima, Ishigaki-jima, Iriomote-jima).

Pseudoxyethira thingana (Oláh 1989)

Scelotrichia thingana Oláh, 1989, 267, male, Vietnam.
Pseudoxyethira thingana: Zhou et al. 2016, 214-215, male, southern China.
Ito 2017c, 197-199, male, habitat, Honshu (Yamanashi, Gifu, Aichi, Shiga, Shimane), Kyushu (Nagasaki), Amami-Oshima, Ishigaki-jima, Iriomote-jima.
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Ito 2018, 503, 507, 511, male, larva, case.
Scelotrichia sp.: Kuhara and Ito 2017, 16, Yakushima.
Type locality. Vietnam.
Distribution. Japan (Honshu, Kyushu, Yakushima, Amami-Oshima, Ishigaki-jima, Iriomote-jima), Vietnam, China (Jiang-xi).

Pseudoxyethira funatsuki Ito 2017

Pseudoxyethira funatsuki Ito 2017c, 197-199, male, habitat, Ishigaki-jima, Iriomote-jima.
Ito 2018, 507, 511, male.
Type locality. Iriomote-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Ishigaki-jima, Iriomote-jima).

Genus Stactobia McLachlan

Thirteen species belonging to this genus have been recorded from Japan.

Stactobia campire Ito 2017

Stactobia campire Ito 2017d, 225-227, 230-231, male, final and early instar larvae, case, habitat, Iriomote-jima.
Ito 2018, 503, 508, 510, male.
Type locality. Iriomote-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Iriomote-jima).

Stactobia chichibu Ito 2017

Stactobia chichibu Ito 2017d, 219-222, 231, male, Honshu (Saitama, Yamanashi, Shizuoka), Shikoku (Tokushima, Kochi).
Ito 2018, 503, 508, 511, male.
Type locality. Saitama, Honshu, Japan.
Distribution. Japan (Honshu, Shikoku).

Stactobia distinguenda Botosaneanu and Nozaki 1996

Stactobia distinguenda Botosaneanu and Nozaki 1996, 60-62, male, habitat, Honshu (Gifu).
Tanida 2016, 73.
Ito 2017d, 219-220, male, Hokkaido (Ishikari), Honshu (Shizuoka, Shiga).
Ito 2018, 508, 510, male.
Ito and Shimura 2019, 35-37, larva, case, Kyushu (Nagasaki).
Ito 2023b, 3-4, adult appearance period, Honshu (Shizuoka).
Type locality. Gifu, Honshu, Japan.
Distribution. Japan (Hokkaido, Honshu, Kyushu).

Stactobia gunma Ito 2017

Stactobia gunma Ito 2017d, 216-218, 229, male, Honshu (Gunma, Tochigi, Kanagawa, Yamanashi, Nagano, Gifu, Shizuoka, Nara).
Ito 2018, 507, 510, male.
Type locality. Gunma, Honshu, Japan.
Distribution. Japan (Honshu).

Stactobia hattorii Botosaneanu and Nozaki 1996

Stactobia hattorii Botosaneanu and Nozaki 1996, 56-58, male, Honshu (Shizuoka).
Tanida 2016, 73.
Ito 2017d, 215-216, 229, male, Honshu (Shizuoka, Kyoto, Hyogo, Okayama).
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 507, 510, male.
Type locality. Shizuoka, Honshu, Japan.
Distribution. Japan (Honshu).
Remarks. Botosaneanu and Nozaki (1996) said that this species belonged to the S. martynovi Species Group Schmid 1983.

Stactobia inexpectata Botosaneanu and Nozaki 1996

Stactobia inexpectata Botosaneanu and Nozaki 1996, 55-56, male, Honshu (Kanagawa, Yamanashi).
Nozaki 1997, 33, 38, Honshu (Kanagawa).
Kagaya et al. 1998, 43, 123, Honshu (Tokyo, Yamanashi).
Nozaki 2004, 1234, Honshu (Kanagawa).
Ito et al. 2010, 57, Hokkaido (Hiyama, Oshima).
Moriya and Kubota 2012, 135, Honshu (Kanagawa).
Tanida 2016, 73.
Ito 2017d, 213-215, 229, male, larva, case, Hokkaido (Ishikari, Shiribeshi, Oshima), Honshu (Iwate, Miyagi, Niigata, Nagano, Shizuoka, Aichi, Mie, Hyogo, Shimane, Okayama, Hiroshima), Shikoku (Tokushima, Ehime, Kochi), Kyushu (Oita, Miyazaki, Kagoshima), Yakushima.
Kuhara and Ito 2017, 14, 16, Yakushima.
Nojima 2017, 112, Honshu (Okayama).
Ito 2018, 507, 510, male, case.
Ito 2023b, 3, Honshu (Aomori), Shikoku (Ehime).
Stactobia sp.: Kagaya et al. (1993), Honshu (Yamanashi), mentioned by Botosaneanu and Nozaki (1996).
Type locality. Kanagawa, Honshu, Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Yakushima).

Stactobia japonica Iwata 1930

Stactobia japonica Iwata 1930, 59, larva, case, Honshu (Gifu, Nagano, Nara).
Fischer 1961, 109.
Tetsukawa 1965, 39-42, larva, case.
Botosaneanu 1990, 47-48.
Ito et al. 1993, 142.
Botosaneanu and Nozaki 1996, 54-55, 60, male, habitat, Honshu (Kanagawa, Nagano, Gifu, Shizuoka).
Nozaki 1997, 33, 38, Honshu (Kanagawa).
Nozaki 2004, 1234, Honshu (Kanagawa).
Moriya and Kubota 2012, 135, Honshu (Kanagawa).
Tanida 2016, 73.
Ito 2017d, 208-213, 229, male, larva, case, Honshu (Iwate, Miyagi, Tochigi, Saitama, Kanagawa, Toyama, Ishikawa, Yamanashi, Nagano, Gifu, Shizuoka, Aichi, Nara, Okayama), Shikoku (Tokushima, Ehime), Kyushu (Fukuoka, Nagasaki, Oita, Miyazaki).
Nojima 2017, 112-113, Honshu (Okayama).
Ito 2018, 503, 507, 510, male, case.
Ito 2023b, 3-4, adult appearance oeriod, Honshu (Akita, Shizuoka).
Type locality. Gifu, Honshu, Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).
Remarks. Botosaneanu and Nozaki (1996) said that this species belonged to the S. martynovi Species Group Schmid 1983.

Stactobia kanagawa Ito 2017

Stactobia kanagawa Ito 2017d, 208, 210, 229, male, Honshu (Kanagawa, Gifu, Shizuoka, Hyogo), Kyushu (Oita).
Ito 2018, 507, 508, 510, male.
Type locality. Kanagawa, Honshu, Japan.
Distribution. Japan (Honshu, Kyushu).

Stactobia makartschenkoi Botosaneanu and Levanidova 1988

Stactobia makartschenkoi Botosaneanu and Levanidova 1988, 169-172, male, larva, case, Far East Russia (Kunashir).
Botosaneanu 1990, 47-48.
Botosaneanu and Nozaki 1996, 55, 61, habitat, Hokkaido (Ishikari), Honshu (Niigata, Shizuoka).
Kuhara et al. 2007, 68, Hokkaido (Ishikari).
Ito et al. 2010, 57, Hokkaido (Shiribeshi, Hiyama, Oshima).
Ito 2013a, 51, 80, Hokkaido (Nemuro, Okhotsk).
Tanida 2016, 73.
Ito 2017d, 203-208, 229, male, final and early instar larvae, case, life cycle, Hokkaido (Tokachi, Kamikawa, Ishikari, Shiribeshi), Honshu (Iwate, Miyagi, Yamagata, Saitama, Niigata, Toyama, Nagano, Gifu, Hyogo, Tottori, , Okayama), Shikoku (Ehime), Kyushu (Fukuoka, Nagasaki, Oita, Kagoshima).
Nojima 2017, 113, Honshu (Okayama).
Ito 2018, 503, 507, 510, male, larva, case.
Ito 2023b, 3, Honshu (Akita).
Type locality. South Kurils, Russia.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (South Kurils), Mongolia.
Remarks. Botosaneanu and Nozaki (1996) said that this species belonged to the S. martynovi Species Group Schmid 1983.

Stactobia nishimotoi Botosaneanu and Nozaki 1996

Stactobia nishimotoi Botosaneanu and Nozaki 1996, 58-61, male, Honshu (Aichi).
Tanida 2016, 73.
Ito 2017d, 218-219, 229, male, Honshu (Miyagi, Hyogo, Shimane), Shikoku (Kochi), Kyushu (Oita, Kagoshima), Yakushima.
Ito 2018, 508, 510, male.
Type locality. Aichi, Honshu, Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu, Yakushima), Korea.
Remarks. Botosaneanu and Nozaki (1996) said that this species belonged to the S. martynovi Species Group Schmid 1983. This species was recorded from Korea (Park et al. 2018).

Stactobia semele Malicky and Chantaramongkol 2007

Stactobia semele Malicky and Chantaramongkol 2007, 1044, 1090, male, Taiwan.
Ito 2017d, 228, 230-232, male, larva, case, Iriomote-jima.
Ito 2018, 508, 510, male.
Type locality. Taiwan.
Distribution. Japan (Ryukyu Islands: Iriomote-jima), Taiwan.

Stactobia urauchi Ito 2017

Stactobia urauchi Ito 2017d, 224-225, 231, male, larva, case, Ishigaki-jima, Iriomote-jima, Yonaguni-jima.
Ito 2018, 508, 510, male.
Type locality. Iriomote-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Ishigaki-jima, Iriomote-jima, Yonaguni-jima).

Stactobia yona Ito 2017

Stactobia yona Ito 2017d, 2222-223, 231, male, larva, case, Amami-Oshima, Okinawa-jima.
Ito 2018, 508, 510, male.
Type locality. Okinawa-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Amami-Oshima, Okinawa-jima).

Genus Stactobiella Martynov

Five species belonging to this genus have been recorded from Japan.

Stactobiellaa aichi Ito 2020

Stactobiella aichi Ito 2020, 565-566, fig. 3, male, habitat, Honshu (Aichi, Shizuoka, Hyogo).
Kawase and Kato 2023, 7, Honshu (Niigata),
Type locality. Aichi, Honshu, Japan.
Distribution. Japan (central Honshu).

Stactobiella amami Ito 2020

Stactobiella amami Ito 2020, 567-568, fig. 4, male, habitat, Amami-o-shima.
Type locality. Amami-o-shima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Amami-o-shima).

Stactobiella biramosa Martynov 1929

Stactobiella biramosa Martynov 1929, 297, male, Russia (Siberia).
Ito 2020, 567-568, male, female, Hokkaido (Rumoi).
Ito and Kuhara 2023, 61, Hokkaido (Rumoi).
Type locality. South Siberia, Russia.
Distribution. Japan (Hokkaido), Russia (Siberia, Khabarovsk, Primorye, Magadan, Sakhalin), China (Hei-long-jiang), Mongolia (Arkhangai).

Stactobiella kumejima Ito 2020

Stactobia urauchi Ito 2020, 567-569, fig. 5, male, female, habitat, Kume-jima, Okinawa-jima.
Type locality. Kume-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Kume-jima, Okinawa-jima).

Stactobiella tshistjakovi (Arefina and Morse, 2002)

Stactobia tshistjakovi Arefina and Morse 2002, 103-104, figs. 16-18, male, Russian Far East (South Primorye).
Stactobiella tshistjakovi: Arefina 2004, 209.
Ito et al. 2010, 57, Hokkaido (Shiribeshi, Oshima).
Ito 2013a, 51, 80, Hokkaido (Kushiro).
Tanida 2016, 73.
Ito 2017e, 40, Hokkaido (Kamikawa).
Ito 2018, 508, 511, male.
Ito 2020, 562-563, male, female, Hokkaido (Kamikawa, Kushiro, Ishikari, Shiribeshi, Oshima), Honshu (Iwate, Shiga).
Ito 2022, 138-141, larva, case, habitat, Hokkaido (Kamikawa).
Ito and Kuhara 2023, 61, Hokkaido (Kamikawa).
Stactobiella sp.: Kuhara 2001, 18, Hokkaido (Shiribeshi). Mentioned by Ito et al. (2010).
Type locality. South Primorye, Russian Far East.
Distribution. Japan (Hokkaido, Honshu), Russia (South Primorye, Sakhalin).

Genus Ugandatrichia Mosely

Two species belonging to this genus have been recorded from Japan.

Ugandatrichia nakijinensis Ito 2012

Ugandatrichia nakijinensis Ito 2012 in Ito and Ohkawa 2012, 49-54, figs. 1-4, 5, male, female, pupa, larva, case, habitat, Okinawa-jima, Amami-Oshima.
Tanida 2016, 74.
Ito 2018, 503, 508, 511, male, larva, case.
Type locality. Okinawa-jima, Ryukyu Islands, Japan.
Distribution. Japan (Ryukyu Islands: Okinawa-jima, Amami-Oshima).

Ugandatrichia taiwanensis Hsu and Chen 2002

Ugandatrichia taiwanensis Hsu and Chen 2002, 74-79, figs. 1-15, male, female, pupa, larva, case, habitat, Taiwan.
Ito and Ohkawa 2012, 54-56, male, female, pupa, larva, case, habitat, Japan (Ishigaki-jima, Iriomote-jima).
Tanida 2016, 74.
Ito 2018, 508, 511, male.
Type locality. Taiwan.
Distribution. Japan (Ryukyu Islands: Ishigaki-jima, Iriomote-jima), Taiwan.

References

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