lepidostomatidae

A catalogue of Japanese Trichoptera
Family Lepidostomatidae Ulmer

Author: Tomiko ITO
E-mail: tobikera[AT]siren.ocn.ne.jp
Last updated: 3 Jan. 2024

Two subfamilies, Lepidostomatinae Ulmer and Theliopsychinae Weaver, belonging to this family have been recorded from Japan.

Subfamily Lepidostomatinae Ulmer
A genus belonging to this subfamily has been recorded from Japan.

Genus Lepidostoma Rambur
Forty eight species belonging to this genus have been recorded from Japan.

Lepidostoma albardanum (Ulmer 1906)
Maniconeura albardana Ulmer 1906, 26-28, male.
Dinarthrodes albardanus: Fischer 1970, 12.

Kumanski and Weaver 1992, 154-156, male, female, new synonym.
Ito et al. 1993, 143-144.

Dinarthrodes kurentzovi Martynov 1935, 392-395, male. Synonymized by Kumanski and Weaver (1992).

Fischer 1970, 14.
Tani 1976, 392, Tsushima.
Tani 1977, 200, male.
Kobayashi 1985b, 17, Tsushima.
Ito 1989, 46-50, male, female, larva, case, habitat, Tsushima.

Goerodes kurentzovi: Ito 1992b, 104.
Goerodes albardanus: Ito 2001, 8.
Lepidostoma albardanum: Weaver 2002, 179.

Ito 2005a, 506, 508, 514, male, larva, case, habitat.
Ito 2005d, 199, 205.
Ito 2005d, 199.
Mey and Nozaki 2006, 24, Honshu (Nagano).
Tanida 2016, 105.
Ito 2018, 599, 603, 610, male, larva.

Type locality. Siberia, Russia.
Distribution. Japan (Honshu (central), Tsushima), Russia (Siberia, Khabarovsk, South Primorye, Sakhalin, South Kurils), Korea, China (Heilongjiang, Jilin).

Lepidostoma albicorne (Banks 1906)
Crunoecia albicornis Banks 1906, 109, head, scape, maxillary palpus of female, Kyushu (Buzen).

Tsuda 1942, 328.
Fischer 1970, 71.

Dinarthrodes albicorne: Weaver, 1985, 244-245, female.

Kumanski and Weaver 1992, 154-156, male, female.
Ito et al. 1993, 144.

Goerodes albicornis: Ito 1999a, 320-323, male, female, larva, case, habitat, Kyushu (Fukuoka).
Lepidostoma albicorne: Weaver 2002, 179.

Ito 2005a, 505, 507, 513, male, larva, case, habitat.
Ito 2005d, 199.
Tanida 2016, 105.
Ito 2018, 599, 601, 609, male, larva.
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka, Oita).

Type locality. Kyushu (Buzen=Fukuoka), Japan.
Distribution. Japan (Kyushu).

Lepidostoma amagiense Ito 2011
Lepidostoma amagiense Ito 2011, 168-170, male, female, habitat, Honshu (Shizuoka, Yamanashi).

Tanida 2016, 105.
Ito 2018, 599, 604, 611, male.

Type locality. Honshu (Shizuoka), Japan.
Distribution. Japan, Honshu (central).

Lepidostoma amamiense (Ito 1990)
Goerodes amamiensis Ito 1990a 369-371, male, female, larva, case, habitat, Amami-Oshima.

Ito 1999b, 499, variation of male, female, Amami-Oshima.

Lepidostoma amamiense: Weaver 2002, 179.

Ito 2005a, 506-507, 513, male, larva, case, habitat.
Ito 2005d, 199.
Shimura et al. 2015, 47, Amami-Oshima.
Tanida 2016, 105.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Amami-Oshima, Japan.
Distribution. Japan (Amami-Oshima).

Lepidostoma axis (Ito 1985)
Goerodes axis Ito 1985d, 512-515, male, female, pupa, larva, case, habitat, Honshu (Tottori).

Hatta and Nozaki 1991, 204, Honshu (Aichi).
Ito et al. 1993, 144.
Ito 1994, 88, Honshu (Hyogo, Tottori).

Lepidostoma axis: Weaver 2002, 180.

Ito 2005a, 505, 507, 513, male, larva, case, habitat.
Ito 2005d, 199-200.
Inazu and Nishida 2011, 198, male, Honshu (Hyogo).
Inazu 2013c, 74, Honshu (Hyogo).
Tanida 2016, 105.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Tottori), Japan.
Distribution. Japan (Honshu (central, west)).

Lepidostoma bipertitum (Kobayashi 1955)
Dinarthrodes bipertitus Kobayashi 1955, 70-72, pl. 14, male, wings of female, Honshu (Saitama) (bipertita).

Kobayashi 1959, 354, Kyushu (Fukuoka).
Fischer 1970, 12.
Tani 1971, 66-67, male, Honshu (Kyoto, Nara) (bipertita).
Kobayashi 1977, 2, Hokkaido (Tokachi) (bipertita).
Tani 1977, 200, male.
Tani 1978, 11, Honshu (Niigata).
Yamanouchi　1978，10，Hokkaido (Tokachi) (bipertita).
Kobayashi 1982, 96, Honshu (Kanagawa) (bipertita).
Tani et al. 1982, 34, Oki-no-shima.
Kobayashi 1984a, 18, 27, female, Hokkaido (Hiyama, Oshima) (bipertita).

Goerodes bipertitus: Ito 1983c, 269-275, habitat, Hokkaido (Rumoi).

Ito 1984c, 507-514.
Ito 1985b, 255-266, female, pupa, larva.
Kobayashi 1987, 22, Honshu (Shimane) (bipertita).
Nozaki 1988b, 42, Honshu (Kanagawa).
Morita 1989, 41, Honshu (Mie).
Ito and Asahina 1990, 21, Honshu (Tokyo).
Kugo and Nozaki, 1990 63, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Hatta and Nozaki, 1991 204, Honshu (Aichi).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 144-145.
Morita 1993, 7, Honshu (Mie).
Nishimoto and Nishimoto 1993, 87, Honshu (Aichi).
Hori et al. 1993, 389, Honshu (Aichi).
Tanida and Takemon 1993, 244, Honshu (Kyoto).
Kuranishi and Kuhara, 1994 1230, Hokkaido (Kushiro).
Moriya 1994, 80, Honshu (Kanagawa, Yamanashi).
Ito et al. 1997, 24, 31, Hokkaido (Oshima).
Kuhara 1997a, 61, Hokkaido (Shiribeshi).
Morita 1997b, 82, Honshu (Mie).
Morita 1999, 41, Honshu (Mie).
Moriya 1997, 69, Honshu (Kanagawa, Yamanashi).
Nishiharu 1997, 61, Honshu (Fukui).
Nozaki 1997a, 37, Honshu (Kanagawa).
Nozaki 1997b, 30, Honshu (Tokyo).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Kagaya et al. 1998, 64, 210, habitat, Honshu (Tokyo, Yamanashi).
Morita 1998a, 14, Honshu (Mie).
Morita 1998d, 4-5, Honshu (Mie).
Morita 1999b, 82, Honshu (Mie).
Nakano and Tanida 1999, 277, Honshu (Nara).
Ohkawa 1999, 40, 44, Hokkaido (Tokachi).
Morita 2001a, 221, Honshu (Aichi).
Morita 2001d, 339, Honshu (Mie).
Ito et al. 2002b, 25, Shikoku (Kagawa, Ehime, Kochi).
Nozaki and Nakamura 2002, 174, Honshu (Hiroshima).
Nakamura et al. 2003, 154, Honshu (Hiroshima).
Shinoda 2003, 145, Honshu (Yamanashi).
Kameyama et al. 2006, 95, Honshu (Hiroshima).
Kameyama et al. 2010, 112, Honshu (Shimane).

Lepidostoma bipertitum: Weaver 2002, 180.

Nozaki 2004, 1236, Honshu (Kanagawa).
Ito 2005a, 505, 508, 514, male, larva, case, habitat.
Ito 2005d, 200.
Nozaki and Nakamura 2005, 223, Honshu (Shimane).
Katsuma 2005b, 14, Honshu (Gunma).
Mey and Nozaki 2006, 24, Honshu (Shiga, Nagano).
Ishiwata et al. 2005, 110, 242-244, Honshu (Kanagawa).
Morita 2006, 200, Honshu (Mie).
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Katsuma 2007a, 18, Honshu (Tochigi).
Morita 2007a, 14, Honshu (Mie).
Nakamura and Hajiri 2007, 198, Honshu (Shimane).
Nakamura and Tsujimura 2007, 78, Honshu (Hiroshima).
Nozaki and Tanida 2007, 251, Honshu (Shizuoka).
Inazu 2008, 80, 94, flight period, Honshu (Hyogo).
Morita 2008a, 6, Honshu (Mie).
Tanaka 2009a, 46, male, Honshu (Akita).
Tanaka 2009b, 19, 22, male, Honshu (Akita).
Kobayashi 2010, 140, Honshu (Tochigi).
Mano 2010, 21, 23, Honshu (Aichi).
Kawase and Morita 2010, 43, Honshu (Mie, Shiga).
Ito et al. 2010, 62, Hokkaido (Shiribeshi, Hiyama, Oshima).
Tanaka 2010a, 25, Honshu (Akita).
Tanaka 2010b, 17, Honshu (Akita).
Inazu and Nishida 2011, 198, male, Honshu (Hyogo).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Katsuma 2011b, 68, Honshu (Ibaraki).
Katsuma 2011c, 69, Honshu (Ibaraki).
Morita 2011, 216, Honshu (Mie).
Nozaki 2011, 4, Honshu (Aichi).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 65, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Hayashi et al. 2013, 13, Oki-no-shima.
Ito 2013, 60, 83, Hokkaido (Kushiro, Tokachi).
Inazu 2013c, 74, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Ito and Nagasaka 2014, 11, Hokkaido (Sorachi).
Nishimoto and Nishimoto 2014, 65, Honshu (Aichi).
Tanaka 2014, 36, 40, male, Honshu (Akita).
Nozaki et al. 2014, 62, Honshu (Nagano).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Imasaka et al. 2016, 34, Kyushu (Fukuoka).
Katsuma, 2016, 48, Honshu (Fukushima).
Morita and Kawase 2016, 22, Honshu (Ishikawa).
Nozaki 2016, 79, 366-368, male, Shikoku (Kochi, Tokushima).
Tanida 2016, 105.
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Ito 2017a, 5, Honshu (Fukui).
Ito 2017b, 44, Hokkaido (Kamikawa).
Kawase et al. 2017, 33, Honshu (Shiga).
Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 599, 603, 610, male, larva.
Tanaka 2018b, 9, Honshu (Akita).
Nozaki et al. 2019, 170, phenology, Honshu (Nagano).
Yamamoto et al. 2019, 72, Shikoku (Ehime).
Morita 2020a, 18, Honshu (Mie).
Morita 2020b, 81, Honshu (Mie).
Taira 2020, 8, Honshu (Nara).
Abe and Ito 2021, 35, mite parasite, Honshu (Yamanashi).
Tezuka et al., 2021, 43, Honshu (Kanagawa).
Yamamoto and Nozaki 2021, 128, Shikoku (Ehime).
Tanaka 2022, 36, Honshu (Akita).
Ito and Kuhara 2023, 65, Hokkaido (Soya, Kamikawa).
Tanaka 2023, 42, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).

Type locality. Honshu (Saitama), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki-no-shima).

Lepidostoma complicatum (Kobayashi 1968)
Dinarthrodes complicatus Kobayashi 1968, 8, pl. 4, male, Honshu (Niigata) (complicata).

Tani 1971, 67-69, male, wings of female, Honshu (Nagano, Shiga, Nara), Shikoku (Tokushima), Kyushu (unknown locality) (complicata).
Yamanouchi 1976, 3, Hokkaido (Tokachi) (complicata).
Tani 1977, 200, male, Honshu (Nagano).
Ito 1978, 575-581, larva, case (complicata).
Tani 1978, 11, Honshu (Niigata) (complicata).
Ito 1980, 313-316, life cycle, Hokkaido (Okhotsuk, Ishikari).
Tanida 1982a, 18, 19, Honshu (Ishikawa).
Tanida 1982b, 361, Honshu (Gunma).
Kobayashi 1984a, 18, male, female, Hokkaido (Hiyama, Oshima).

Goerodes complicatus: Ito 1983c, 269-275, habitat, Hokkaido (Rumoi, Ishikari, Oshima).

Ito 1984b, 240-246, drift of larva, Hokkaido (Ishikari).
Ito 1984c, 507-514.
Ito 1985b, 263-264, female, pupa, larva.
Nozaki 1988b, 37, Honshu (Kanagawa).
Ito and Asahina 1990, 21, Honshu (Yamanashi).
Ito et al. 1993, 145.
Morita 1993, 7, Honshu (Mie).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Kuranishi and Kuhara 1994, 1230, Hokkaido (Kushiro).
Moriya 1994, 80-81, Honshu (Kanagawa, Yamanashi).
Itou and Muramatsu 1995, 42, Hokkaido (Okhotsuk).
Ito et al. 1997, 24, 28, 30, Hokkaido (Oshima).
Kuhara 1997a, 62, Hokkaido (Shiribeshi).
Moriya 1997, 37, Honshu (Kanagawa).
Nishiharu 1997, 61, Honshu (Fukui).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Ito et al. 1998, 11-14, 17, Hokkaido (Nemuro).
Kagaya et al. 1998, 64, 211, habitat, Honshu (Tokyo, Yamanashi).
Sekimoto et al. 1998, 361, Honshu (Hiroshima).
Ito et al. 1999, 24-27, Hokkaido (Kushiro).
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000b, 36, Hokkaido (Ishikari).
Kuhara et al. 2000, 256, Hokkaido (Ishikari, Iburi).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Ito et al. 2002b, 25, Shikoku (Ehime, Kochi).
Aoyagi et al. 2004, 26, Honshu (Kyoto).
Morita and Kawase 2004, 17, Honshu (Ishikawa).

Lepidostoma complicatum: Weaver 2002, 180.

Ishiwata et al. 2005, 110, 243, Honshu (Kanagawa).
Ito 2005a, 505, 508, 510, 514, male, larva, case, habitat.
Ito 2005b, 73-78, food.
Ito 2005d, 200.
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Kochi and Kagaya 2005, 656-667, food.
Kimura et al. 2006, 7, Honshu (Nagano).
Mey and Nozaki 2006, 24, Honshu (Nagano, Shiga).
Morita 2006, 200, Honshu (Mie).
Ito and Kosugi 2007, 54, Hokkaido (Kushiro).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Kuhara et al. 2007, 72, Hokkaido (Ishikari).
Morita 2008a, 6, Honshu (Mie).
Morita 2008c, 6, Honshu (Mie).
Morita and Kawase 2008, 55, Honshu (Ishikawa).
Muramatsu 2008, 7, Hokkaido (Okhotsuk).
Ito et al. 2009, 8-9, Hokkaido (Okhotsuk).
Ito 2010, 131, Hokkaido (Kushiro).
Tsuji et al. 2010, 6, Hokkaido (Okhotsuk).
Kawase and Morita 2010, 43, Honshu (Mie, Shiga).
Ito et al. 2010, 62, Hokkaido (Shiribeshi, Hiyama, Oshima).
Inazu and Nishida 2011, 198, male, Honshu (Hyogo).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Kuhara 2011, 53, 63, 74, flight season, Hokkaido (Sorachi. Ishikari, Hidaka).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 60, 83, Hokkaido (Nemuro, Okhotsk, Kushiro, Tokachi).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Inazu 2013c, 74, Honshu (Hyogo).
Yamamoto and Ito 2014, 14, 18-19, Shikoku (Ehime).
Ito and Nagasaka 2014, 11, Hokkaido (Sorachi).
Tanaka 2014, 37, 40, male, Honshu (Akita).
Ijima and Tsuchiya 2014, 31, Hokkaido (Kushiro).
Nozaki et al. 2014, 62, Honshu (Nagano).
Ito 2016b, 1-8, protogyny.
Nozaki 2016, 79, 367-368, male, Shikoku (Ehime. Kochi).
Tanida 2016, 105.
Yamamoto and Ito 2016, 41, 43, 44, Shikoku (Ehime).
Ito 2017b, 44, Hokkaido (Kamikawa).
Katsuma 2017, 41, Honshu (Ibaraki).
Nojima 2017, 121, Honshu (Okayama).
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima).
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 599, 603, 610, male, larva.
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Tanaka 2018a, 21, Honshu (Akita).
Tanaka 2018b, 9, Honshu (Akita).
Suzuki 2019a, 117, Honshu (Gifu).
Suzuki 2019b, 65, Honshu (Gifu).
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka, Oita).
Ito 2020, 17-22, spermatophore, Hokkaido (Ishikari).
Nakatani 2021, 72, Hokkaido (Kushiro).
Yamamoto and Nozaki 2021, 129, Shikoku (Ehime).
Yamamoto et al. 2021, 74, Shikoku (Ehime).
Tanaka 2022, 36, Honshu (Akita).
Ito and Kuhara 2023, 65, Hokkaido (Soya, Okhotsuk, Kamikawa).
Tanaka 2023, 42, Honshu (Akita).

Dinarthrodes japonica: Kobayashi 1977, 2, Hokkaido (Tokachi). Misidentification mentioned by Ito (2013).
Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).

Lepidostoma coreanum (Kumanski and Weaver 1992)
Indocrunoecia coreanum Kumanski and Weaver 1992, 163-165, male, female.
Dinarthrum coreanum: Ito 1998, 585-588, male, female, larva, case, habitat, Honshu (Miyagi, Yamagata, Kanagawa, Yamanashi, Shizuoka, Kyoto, Wakayama), Russia (Primorye).

Kawase and Morita 2001, 9, Honshu (Mie).
Ito et al. 2002b, 23, adult flight season, sex ratio, Shikoku (Ehime).
Morita 2002, 37, Honshu (Mie).
Morita 2003, 26, Honshu (Mie).

Lepidostoma coreanum: Weaver 2002, 180.

Ito 2005a, 504, 507, 512, male, larva, case, habitat.
Ito 2005d, 200.
Morita 2008d, 91-92, Honshu (Mie).
Kawase and Morita 2010, 43, Honshu (Mie).
Morita 2009, 140, Honshu (Mie).
Morita 2011, 216, Honshu (Mie).
Ito 2016a, 398-403, male, female, larva, case, Honshu (Shizuoka, Wakayama).
Tanida 2016, 106.
Ito 2018, 598, 601, 608, male, larva.

Lepidostomatidae gen. sp.: Nozaki 1997a, 37, Honshu (Kanagawa), mentioned by Nozaki (2004).
Type locality. North Korea.
Distribution. Japan (Honshu, Shikoku), Russia (South Primorye), North Korea.
Remarks. Descriptions of the female in Ito (1998) is wrong due to incorrect specimens.

Lepidostoma cornigera (Ulmer 1907)
Goerodes cornigera Ulmer 1907a, 37-40, male, wings of female, Japan (Tokyo?).

Ulmer 1907b, 105-106, wings of male and female.
Tsuda 1942, 328, Honshu (Kyoto).
Fischer 1970, 19 (corniger).
Malicky 1979, 101.
Ito 1990b, 789-790, male, Hokkaido (Ishikari) (corniger).
Ito et al. 1993, 145 (corniger).
Kuhara et al. 2000, 256, Hokkaido (Ishikari, Iburi) (corniger).
Ito 2001, 1-8, male, variation of male, female, pupa, larva, case, new synonym, Hokkaido (Ishikari, Iburi), Honshu (Akita, Yamagata, Aichi), Kyushu (Fukuoka), Tsushima, Yakushima (corniger).
Morita 2001a, 221, Honshu (Aichi) (corniger).
Morita 2001b, 175, Honshu (Mie) (corniger).

Crunoeciella toyotamaensis Kobayashi 1985b, 15-17, male. Synonymized by Ito (2001).
Goerodes toyotamaensis: Ito 1989, 53-55, male, female, larva, case, habitat, Tsushima.

Ito 1990a, 362, Yakushima.
Gyotoku and Nozaki 1992, 139-140, Kyushu (Fukuoka).
Ito et al. 1993, 146.
Nishimoto and Nishimoto 1993, 87, Honshu (Aichi).

Lepidostoma cornigera: Weaver 2002, 184 (cornigerum).

Ito 2005a, 505, 508, 515, male, larva, case, habitat (cornigerum).
Ito 2005d, 200 (cornigerum).
Tanaka 2010b, 17-19, male, Honshu (Akita) (cornigerum).
Morita 2011, 213, 216, Honshu (Mie) (cornigerum).
Morita 2012, 250, Honshu (Mie) (cornigerum).
Tanaka 2014, 37, 40, male, Honshu (Akita) (cornigerum).
Morita 2015, 17, Honshu (Mie) (cornigerum).
Tanida 2016, 106 (cornigerum).
Kuhara and Ito 2017, 14, 17, Yakushima.
Nojima 2017, 121, Honshu (Okayama).
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima) (cornigerum).
Ito 2018, 599, 604, 611, male, larva.
Tanaka 2018b, 9, Honshu (Akita).
Tanaka 2019, 12, Honshu (Akita) (cornigerum).
Shimura and Imasaka 2021, 5, Kagoshima (Koshiki-retto) (cornigerum).
Tanaka 2023, 42, Honshu (Akita).

Type locality. Japan (Tokyo?).
Distribution. Japan (Hokkaido, Honshu, Kyushu, Tsushima, Yakushima, Koshiki-retto), China (Fujian, Jiangxi).
Remarks. The species-group name "cornigera" should be treated as a noun in accordance with the International Code of Zoological Nomenclature, 1999, Article 31.2.2, and need not agree in gender with the generic name (Article 31.2.1, 34.2.1).

Lepidostoma crassicorne (Ulmer 1907)
Severinia crassicornis Ulmer 1907a, 35-37, male, wings and basal parts of female antenna, Japan.

Ulmer 1907b, 105, wings of male and female.

Neoseverinia carassicornis: Ulmer 1908, 343.

Tsuda 1936, 403-405, larva, case, Honshu (Nagano, Kyoto).
Tsuda 1942, 328, Honshu (Nagano, Kyoto).
Tsuda 1943, 163, Honshu (Kyoto).
Tsuda 195１, 68, larva, case.
Tsuda 1959, 151, larva, case.
Fischer 1970, 26-27.
Tani 1971, male, wings of female, Honshu (Tochigi, Kyoto, Nara).
Tani 1977, 200, male.
Tani 1978, 12, Honshu (Niigata).
Tani et al. 1982, 34, Oki-no-shima.
Ito 1983a, 207-214, geographical variation of male, female, pupa.
Ito 1983b, 322-326, larva, case.
Kobayashi 1984c, 6, Sado.
Tanida 1985, 201, larva, case.
Ohtsuka and Tanida 1986, 6, Kyushu (Kumamoto).
Yamanouchi 1987, 36, Hokkaido (Tokachi).
Kobayashi 1987, 21, Honshu (Shimane).
Morita 1989, 41, Honshu (Mie).
Morita 1990, 31, Honshu (Mie).
Hatta and Nozaki 1991, 2004, Honshu (Aichi).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 146-147.
Morita 1993, 7, Honshu (Mie).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Kuhara and Ito 1994, 21, 23, Hokkaido (Ishikari).
Kuranishi and Kuhara 1994, 1230, Hokkaido (Kushiro).
Moriya 1994, 82, Honshu (Kanagawa, Yamanashi).
Itoet al. 1997, 24, 28, Hokkaido (Oshima).
Kuhara 1997b, 96, Hokkaido (Shiribeshi).
Moriya 1997, 70, Honshu (Kanagawa, Yamanashi).
Nishiharu 1997, 61, Honshu (Fukui).
Nozaki 1997a, 30, Honshu (Tokyo).
Nozaki 1997b, 37, Honshu (Kanagawa).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Ito et al. 1998, 11, 13, Hokkaido (Nemuro).
Kagaya et al. 1998, 67-68, 219, habitat, Honshu (Tokyo, Yamanashi).
Morita 1998a, 13, Honshu (Mie).
Nishimoto and Nishimoto 1998, 151, Honshu (Aichi).
Nozaki 1998b, 42, Honshu (Kanagawa).
Sekimoto et al. 1998, 361, Honshu (Hiroshima).
Kushiro Insect Lover’s Society 1999, 219, Hokkaido (Nemuro).
Morita 1999c, 89, Honshu (Mie).
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 2000a, 27-28, Hokkaido (Soya).
Ito et al. 2000b, 37, Hokkaido (Ishikari).
Maruyama and Takai 2000, 109-113, larva, case, Honshu (Nara).
Kawase and Morita 2001, 9, Honshu (Ishikawa).
Kuhara and Sato 2001, 65-67, Rishiri.
Morita 2001a, 221, Honshu (Aichi).
Morita 2001d, 339, Honshu (Mie).
Ito et al. 2002b, 28-29, flight season, Shikoku (Ehime, Kochi).
Nozaki and Nakamura 2002, 174, Honshu (Hiroshima).
Uchida 2002, 78, Honshu (Tochigi).
Morita 2003, 26, Honshu (Mie).
Nakamura et al. 2003, 154, Honshu (Hiroshima).
Aoyagi et al. 2004, 26, Honshu (Kyoto).
Kawase et al. 2004, 115-116, Honshu (Ibaraki).
Kameyama and Nakamura 2006, 190, Honshu (Tottori).

Lepidostoma crassicorne: Weaver 2002, 180.

Yoshitake 2004, 28, Honshu (Okayama).
Ito 2005a, 505, 509-511, 515, male, larva, case, habitat.
Ito 2005d, 200-201.
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Nozaki 2005, 110, adult, larva, case.
Mey and Nozaki 2006, 24, Honshu (Nagano).
Morita 2006, 200, Honshu (Mie).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Torii and Hattori 2006, 38, Honshu (Shizuoka).
Ito and Kosugi 2007, 54, Hokkaido (Kushiro).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Kuhara et al. 2007, 72, Hokkaido (Ishikari).
Morita 2007a, 14, Honshu (Mie).
Morita 2007b, 97, Honshu (Mie).
Inazu 2008, 80, 94, flight period, Honshu (Hyogo).
Morita 2008c, 6-7, Honshu (Shiga).
Muramatsu 2008, 8, Hokkaido (Okhotsuk).
Tanida 2008, pl. 93, 252, male, Honshu (Ishikawa).
Kobayashi 2009, 63, Honshu (Tochigi).
Morita 2009, 140, Honshu (Mie).
Tanaka 2009b, 19, 22, male, Honshu (Akita).
Kawase and Hayashi 2010b, 86, Honshu (Shimane).
Katsuma 2010, 85, Honshu (Ibaraki).
Nakamura 2010, 239, Honshu (Hiroshima).
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Ito et al. 2010, 62, Hokkaido (Shiribeshi, Oshima).
Tanaka 2010b, 17, Honshu (Akita).
Ito and Nakatani 2011, 147, Hokkaido (Kushiro).
Inazu and Nishida 2011, 198, male, larva, case, Honshu (Hyogo).
Katsuma 2011a, 49-51, Honshu (Ibaraki).
Katsuma 2011b, 68, Honshu (Ibaraki).
Kuhara 2011, 53, 63, 74, flight season, Hokkaido (Sorachi, Ishikari, Hidaka).
Watanabe and Mitsuhashi 2011, case material selection, Honshu (Hyogo).
Tanaka and Okuma 2011, 24, Honshu (Akita).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 60, 83, Hokkaido (Nemuro, Okhotsk, Kushiro, Tokachi).
Enju et al. 2013, 89, larva, case.
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Inazu 2013a, 46, Honshu (Hyogo).
Inazu 2013b, 57, Honshu (Hyogo).
Inazu 2013c, 74, Honshu (Hyogo).
Ijima and Tsuchiya 2014, 31, Hokkaido (Kushiro).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nozaki et al. 2014, 62, Honshu (Nagano).
Tanaka 2014, 37, 41, male, Honshu (Akita).
Yamamoto and Ito 2014, 14, 18-19, Shikoku (Ehime).
Yamamoto and Ito 2015, 33, Shikoku (Ehime).
Watanabe 2015, 19-27, egg mass and oviposition site, Honshu (Hyogo).
Horiguchi 2016, 413, Honshu (Tokyo).
Nozaki 2016, 79, 367-368, male, Shikoku (Kochi).
Tanida 2016, 106.
Morita and Kawase 2016, 22, Honshu (Ishikawa).
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Katsuma 2017, 41, Honshu (Ibaraki).
Kawase et al. 2017, 33, Honshu (Shiga).
Nojima 2017, 121, Honshu (Okayama).
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 599, 604, 607, 611, male, larva, case.
Tsuruda 2018, 27, 34, adult, Honshu (Tokyo).
Nakatani 2019, 49, Hokkaido (Kushiro).
Suzuki 2019, 119, Honshu (Gifu).
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka).
Abe and Ito 2021, 35, mite parasite, Hokkaido (Ishikari).
Yamamoto et al. 2021, 74, Shikoku (Ehime).
Ito and Kuhara 2023, 65, Hokkaido (Soya, Kamikawa).
Kawase and Kato 2023, 8, Honshu (Niigata).
Tanaka 2023, 43, Honshu (Akita).

Ito 1992b, 100-102, male, larva, case, habitat, Hong Kong.

Lepidostoma doligung: Weaver 2002, 184

Shimura 2010, 50, 54, adult, Yonaguni-jima.
Tanida 2016, 106.
Ito 2018, 600, 604, 611, male, larva.

Type locality. India (Andaman Islands).
Distribution. Ryukyu (Yonaguni-jima), India, Taiwan, Hong Kong.

Lepidostoma ebenacanthum (Ito 1992)
Goerodes ebenacanthus Ito 1992a, 339-342, male, female, larva, case, habitat, Iriomote-jima.

Ito et al. 1993, 145.
Ito 1999b, 500, Ishigaki-jima, Iriomote-jima.

Lepidostoma ebenacanthum: Weaver 2002, 180.

Ito 2005a, 506, 508, 514, male, larva, case, habitat.
Ito 2005d, 200 (ebenacanthus).
Tanida 2016, 106.
Ito 2018, 599, 603, 610, male, larva.

Type locality. Iriomote-jima, Japan.
Distribution. Japan (Ishigaki-jima, Iriomote-jima), Taiwan.

Lepidostoma elongatum (Martynov 1935)
Dinarthrodes elongatus Martynov 1935, 379-384, male, forewing of female, Ussuri, Russia (elongata).

Tani 1976, 393, Tsushima (elongata).
Tani 1977, 200, male (elongata).
Kobayashi 1985b, 17, Tsushima (elongata).
Ito 1989, 50-53, male, female, larva, case, habitat, Tsushima.
Kumanski and Weaver 1992, 157-159, male, female.
Ito et al. 1993, 144.

Goerodes elongatus: Ito 1985b, female.

Ito 1992b, 103-104.
Ito 2001, 8.

Lepidostoma elongatum: Weaver 2002, 180.

Ito 2005a, 506, 508, 514, male, larva, case, habitat.
Ito 2005d, 201.
Tanida 2016, 106.
Ito 2018, 599, 604, 611, male, larva.

Type locality. Ussuri, Russia.
Distribution. Japan (Tsushima), Russia (Siberia, Khabarovsk, Sakhalin, Kurils), Korea, China (Heilongjianga, Jilin).

Lepidostoma emarginatum (Ito 1985)
Goerodes emarginatus Ito 1985d, 507-512, male, female, pupa, larva, case, habitat, Honshu (Iwate, Tochigi).

Ito et al. 1993, 145, 212.
Kagaya et al. 1998, 65, Honshu (Yamanashi).

Lepidostoma emarginatum: Weaver 2002, 180.

Ito 2005a, 504, 507, 513, male, larva, case, habitat.
Ito 2005d, 201.
Tanida 2016, 106.
Ito 2018, 599, 601, 609, male, larva.
Tanaka 2023, 43, 47, male, Honshu (Akita).

Ito 2018, 598, 601, 608, male.
Nojima 2022, 38, Honshu (Okayama).

Type locality. Honshu (Yamanashi), Japan.
Distribution. Japan (Honshu (central, western)).

Lepidostoma hirtum (Fabricius 1775)
Phrygania hirta Fabricus, 1775, 308, England.
Lepidostoma hirtum: McLachlan 1880, 274-276, male, female.

Hickin 1967, 447-449, larva, case.
Fischer 1970, 43-57.
Fischer 1973, 162.
Ito et al. 1993, 146.
Weaver 2002, 184.
Ito 2005a, 505, 509, 512, male, larva, case, habitat.
Ito 2005d, 201.
Nozaki and Nakamura 2005, 223, Honshu (Shimane).
Nakamura and Hajiri 2007, 199, Honshu (Shimane).
Nakamura and Tsujimura 2007, 78, Honshu (Hiroshima).
Inazu and Nishida 2011, 199, male, Honshu (Hyogo).
Tanida 2016, 106-107.
Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 599, 604, 611, male, larva.

Ayabeopsyche nipponica Tsuda 1942a, 332-333, male, Honshu (Kyoto). Synonymized by Kumanski and Weaver (1992).

Tsuda 1942b, 263-264, Honshu (Kyoto).
Fishcher 1970, 70.
Tani 1971, 49-50, male, wings of female, Honshu (Kyoto).

Goerodes nipponica: Tani and Nakamura, 1997, 65, Honshu (Hiroshima).

Kameyama et al. 2006, 96, Honshu (Hiroshima).
Kameyama et al. 2010, 112-113, Honshu (Shimane).

Type locality. England.
Distribution. Japan (Honshu (west)), Palaearctic.

Lepidostoma hiurai (Tani 1971)
Crunoeciella hiurai Tani, 1971, 53-54, male, wings of female, Hokkaido (Rumoi).

Tani 1977, 200, male.
Yamanouchi 1978, 10, Hokkaido (Tokachi).

“Crunoeciella” hiurai: Malicky 1979, 100-101.
Goerodes hiurai: Ito 1983c, 269-275, habitat.

Ito 1984c, 507-514.
Ito 1985a, 12-16, 23, female, pupa, larva, case, habitat, life cycle.
Ito 1990b, 791, Hokkaido (Kushiro, Tokachi, Iburi).
Itou 1989, 56, Hokkaido (Kushiro).
Ito et al. 1993, 145.
Nishimoto 1993, 22, Hokkaido (Kushiro).
Itou 1995, 167, Hokkaido (Kushiro).
Ohkawa 1999, 40-44, Hokkaido (Tokachi).
Ito et al. 1998, 11-13, 16, Hokkaido (Nemuro).
Ito et al. 1999, 24-25, Hokkaido (Kushiro).
Ito et al. 2000a, 27-30, Hokkaido (Soya).
Kuhara et al. 2000, 256-257, Hokkaido (Ishikari, Iburi).
Kuhara 2001, 19, Hokkaido (Shiribeshi).

Lepidostoma hiurai: Weaver 2002, 185.

Ito et al. 2004, 15, Hokkaido (Kushiro).
Ito 2005a, 505, 508, 515, male, larva, case, habitat.
Ito 2005d, 201.
Ito and Kosugi 2007, 54, Hokkaido (Kushiro).
Ito et al. 2007, 149, Hokkaido (Kushiro).
Nakano et al. 2007. 40, Hokkaido (Nemuro).
Muramatsu 2008, 8, Hokkaido (Okhotsuk).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Kuhara 2011, 53, 74, flight season, Hokkaido (Sorachi).
Ito 2013, 60-61, 83, Hokkaido (Nemuro, Okhotsk, Kushiro, Tokachi).
Nakatani 2016, 241, Hokkaido (Nemuro).
Tanida 2016, 106.
Ito 2017b, 44, Hokkaido (Kamikawa).
Ito 2018, 599, 603, 611, male, larva.
Ito and Kuhara 2023, 65, Hokkaido (Soya, Kamikawa).

Type locality. Hokkaido (Rumoi), Japan.
Distribution. Japan (Hokkaido), Russia (Khabarovsk, Sakhalin, South Kurils).

Lepidostoma hokurikuense (Ito 1994)
Goerodes hokurikuensis Ito 1994, 79-82, male, female, larva, case, habitat, Honshu (Ishikawa, Fukui, Tottori).
Lepidostoma hokurikuense: Weaver 2002, 181.

Ito 2005a, 504, 507, 513, male, larva, case, habitat.
Ito 2005d, 201.
Morita 2008c, 6-7, Honshu (Shiga).
Ito 2009, 49-51, male, female, Honshu (Ishikawa, Fukui, Yamanashi, Nagano, Gifu, Mie, Shiga, Kyoto, Hyogo, Tottori).
Kawase and Morita 2010, 34, 44, Honshu (Mie).
Inazu and Nishida 2011, 199, male, female, Honshu (Hyogo).
Inazu 2013c, 74, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nozaki et al. 2014, 62, Honshu (Nagano).
Tanida 2016, 107.
Kawase et al. 2017, 33, Honshu (Shiga).
Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Fukui), Japan.
Distribution. Japan (Honshu (central, west)).

Lepidostoma iriomotense (Ito 1999)
Goerinella arcuatus: Ito 1992a, 337-339, male, female, larva, case, Okinawa (Iriomote). Misidentification, mentioned by Ito (1999a).

Ito et al. 1993, 144.

Goerodes iriomotensis Ito 1999a, 328-331, male, female, larva, case, habitat, Okinawa (Iriomote-jima).
Lepidostoma iriomotense: Weaver 2002, 185.

Ito 2005a, 506, 509, 515, male, larva, case, habitat.
Ito 2005d, 201.
Tanida 2016, 107.
Ito 2018, 600, 604, 611, male, larva.

Type locality. Iriomote-jima, Japan.
Distribution. Japan (Iriomote-jima).

Lepidostoma ishigakiense Ito 2016
Lepidostoma ishigakiense Ito 2016a, 415-417, male, habitat, Ryukyu Islands (Ishigaki-jima).

Ito 2018, 599, 601, 608, male.

Type locality. Ryukyu (Ishigaki-jima), Japan.
Distribution. Japan (Ryukyu (south)).

Lepidostoma itoae (Kumanski & Weaver 1992)
Dinarthrodes itoae Kumanski and Weaver 1992, 161-163, figs. 15-18, male, northern Korea.
Lepidostoma itoae: Weaver 2002, 181.

Ito and Yamamoto 2012, 7-13, male, female, pupa, larva, habitat, Honshu (Mie), Shikoku (Ehime).
Yamamoto 2013, 32-34, adult, pupa, larva, habitat, Shikoku (Ehime).
Ito 2016b, 1-8, emergence pattern.
Tanida 2016, 107.
Ito 2018, 598, 601, 608, male, larva.
Ito 2020, 17-22, spermatophore, Shikoku (Ehime).
Yamamoto 2021, 147-149, adult, larva, habitat, Shikoku (Ehime).

Type locality. Korea.
Distribution. Japan (Honshu, Shikoku), Korea.

Lepidostoma japonicum (Tsuda 1936)
Atomyiella japonica Tsuda 1936, 400-403, male, larva, case, Honshu (Kyoto).

Tsuda 1937, 258-259, larva, case, habitat, Honshu (Tochigi, Kanagawa).

Dinarthrodes japonicus: Tsuda, 1941, 105, Honshu (Nagano) (japonica).

Tsuda 1942a, 329, Honshu (Kanagawa, Nagano, Shizuoka, Shiga, Kyoto) (japonica).
Tsuda 1942b, 263-264, Honshu (Kyoto) (japonica).
Tsuda 1943, 163, 168, 170, Honshu (Kyoto) (japonica).
Tsuda 1950, 428, adult (japonica).
Tsuda 1959, 152, larva, case (japonica).
Ishihara et al. 1953, 38, Shikoku (Ehime) (japonica).
Kobayashi 1955, 72, male (japonica)
Chihara 1956, 84-87, pupa (japonica).
Kobayashi 1959, 354, Kyushu (Fukuoka) (japonica).
Kuwada 1965, 166, adult, Shikoku (Ehime) (japonica).
Fischer 1970, 13-14.
Tani 1971, 64-66, male, wings of female, Honshu (Niigata, Shiga, Kyoto, Osaka, Hyogo, Nara), Fukue-jima? (based on female) (japonica).
Tani 1977, male, Honshu (Shiga).
Ito 1978, 583, larva, case (japonica).
Tani 1978, 11, Honshu (Niigata).
Tani 1980a, 28, larva, case.
Tani 1980b, 107-108, larva, case.
Tani et al. 1982, 34, Oki-no-shima.
Tanida 1982, 18, Honshu (Ishikawa).
Kobayashi 1982, 96, Honshu (Kanagawa) (japonica).
Kobayashi 1984c, 6, Sado (japonica).
Kadowaki 1998, 7, Honshu (Shimane).

Goerodes japonicus: Ito 1983c, 269-275, habitat.

Ito 1984c, 507-514.
Tanida 1985, 201, larva, case.
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Ito 1985b, 266, female, pupa, larva.
Kobayashi 1987, 21-22, Honshu (Shimane), Oki-no-shima.
Nozaki 1987, 42-44, pl. 7, adult, Honshu (Kanagawa).
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 42, Honshu (Kanagawa).
Morita 1989, 41, Honshu (Mie).
Ito 1990a, 361-362, Yakushima.
Ito and Asahina 1990, 21, Honshu (Tokyo, Shizuoka).
Kugo and Nozaki 1990, 63, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Nozaki and Gyotoku 1990, 17, Kyushu (Fukuoka).
Hatta and Nozaki 1991, 204, Honshu (Aichi).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 145.
Morita 1993, 7, Honshu (Mie).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 81, Honshu (Kanagawa, Yamanashi).
Kuranishi and Ohtaka 1996, 15, Honshu (Aomori).
Morita 1996, 303, Honshu (Mie).
Ito et al. 1997, 24, Hokkaido (Oshima).
Kuhara 1997a, 62, Hokkaido (Shiribeshi).
Moriya 1997, 69, Honshu (Kanagawa, Yamanashi).
Nishiharu 1997, 61, Honshu (Fukui).
Nozaki 1997a, 37, Honshu (Kanagawa).
Nozaki 1997b, 31, Honshu (Tokyo).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Kagaya et al. 1998, 65, 213, habitat, Honshu (Tokyo, Yamanashi).
Morita 1998a, 13, Honshu (Mie).
Morita 1998d, 5, Honshu (Mie).
Nishimoto and Nishimoto 1998, 151, Honshu (Aichi).
Nakano and Tanida 1999, 277.
Kawase and Morita 2001, 9, Honahu (Ishikawa).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Morita 2001a, 221, Honshu (Aichi).
Morita 2001b, 175, Honshu (Mie).
Morita 2001d, 339, Honshu (Mie).
Ito et al. 2002b, 25-26, adult flight season, Shikoku (Kagawa, Ehime).
Nozaki and Nakamura 2002, 174, Honshu (Hiroshima).
Uchida 2002, 78, Honshu (Tochigi).
Nakamura et al. 2003, 154, Honshu (Hiroshima).
Kawase et al. 2004, 114-115, Honshu (Ibaraki).
Shinoda 2003, 145, Honshu (Yamanashi).
Kameyama and Nakamura 2006, 190, Honshu (Tottori).

Lepidostoma japonicum: Weaver 2002, 181.

Ishiwata et al. 2005, 110, 243-244, Honshu (Kanagawa).
Ito 2005a, 505, 508, 514, male, larva, case, habitat.
Ito 2005d, 201-202.
Nozaki and Nakamura 2005, 223, Honshu (Hiroshima).
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Nozaki 2005, 110, adult, larva, case.
Kimura et al. 2006, 7, Honshu (Nagano).
Tani 2006, 107, larva, case.
Mey and Nozaki 2006, 24, Honshu (Nagano, Gifu).
Morita 2006, 200, Honshu (Mie).
Torii and Hattori 2006, 38-39, Honshu (Shizuoka).
Morita 2007a, 14, Honshu (Mie).
Nozaki and Tanida 2007, 251, Honshu (Shizuoka).
Satake and Kuranishi 2007, 283, Izu Islands (Mikura-jima).
Shinoda 2007, 97, Honshu (Yamanashi).
Inazu 2008, 80-81, 94, flight period, Honshu (Hyogo).
Morita 2008a, 6, Honshu (Mie).
Morita 2008c, 6-7, Honshu (Shiga).
Morita 2008d, 92, Honshu (Mie).
Tanida 2008, pl. 93, 252, male, Shikoku (Ehime).
Aoyagi and Iwasaki 2009, 101, 102, 104, 106, Honshu (Osaka).
Morita 2009, 140-141, Honshu (Mie).
Tanaka 2009b, 19, 22, male, Honshu (Akita).
Kawase and Hayashi 2010b, 86, Honshu (Shimane).
Kawase et al. 2010, 107, Honshu (Shimane).
Kobayashi 2010, 141, Honshu (Tochigi).
Katsuma 2010, 85, Honshu (Ibaraki).
Mano 2010, 21, 23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Ito et al. 2010, 62, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kawase and Hayashi, 2011, 192, Honshu (Shimane).
Inazu and Nishida 2011, 199, male, female, Honshu (Hyogo).
Katsuma 2011a, 49-51, Honshu (Ibaraki).
Katsuma 2011b, 68, Honshu (Ibaraki).
Morita 2011, 216, Honshu (Mie).
Tsukiji 2011, 141, male.
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Morita 2012, 251, Honshu (Mie).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Enju et al. 2013, 85, male.
Inazu 2013c, 74, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Nishimoto and Nishimoto 2014, 66, male, Honshu (Aichi).
Tanaka 2014, 36, 40, male, Honshu (Akita).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Shimura 2015, male, 54, 63, Fukue-jima.
Horiguchi 2016, 413, Honshu (Tokyo).
Imasaka et al. 2016, 34, Kyushu (Fukuoka).
Nozaki, 2016, 79, 367-368, male, Shikoku (Kochi).
Tanida 2016, 107.
Uchifune 2016, adult, 200-201, Honshu (Kanagawa).
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Ito 2017, 5, Honshu (Fukui).
Kuhara and Ito 2017, 14, 17, Yakushima.
Kawase et al. 2017, 34, Honshu (Shiga).
Nojima 2017, 121, Honshu (Okayama).
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima).
Ito 2018, 599, 602, 610, male, larva.
Tanaka 2018b, 9, Honshu (Akita).
Nozaki et al. 2019, 170, phenology, Honshu (Nagano).
Yamamoto et al. 2019, 73, Shikoku (Ehime).
Morita 2020b, 81, Honshu (Mie).
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka).
Yashiro et al. 2020, 185, Kyushu (Kagoshima, Koshiki-jima-retto).
Abe and Ito 2021, 35, mite parasite, Honshu (Niigata).
Tezuka et al. 2021, Honshu (Kanagawa).
Shimura and Imasaka 2021, 5, Kagoshima (Koshiki-retto).
Yamamoto and Nozaki 2021, 129, Shikoku (Ehime).
Tanaka 2022, 36, Honshu (Akita).
Kawase and Kato 2023, 8, Honshu (Niigata).
Tanaka 2023, 43, Honshu (Akita).
Yamamoto 2023, 33, Shikoku (Ehime).

Lepidostoma hirtum Rambur: Iwata 1927a, 261-262, larva; Iwata 1927b, 216, larva. Misidentification of the larvae of L. crassicorne, L. emarginatum or L. japonicum, mentioned by Ito (1990b).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Hokkaido (south), Honshu, Shikoku, Kyushu, Sado, Izu, Oki-no-shima, Fukue-jima (Goto Retto), Koshiki-retto, Yakushima).
Remarks. The records based on female alone are questionable, since the females of L. japonicum, L. complicatum and L. kasugaense are very similar and difficult to distinguish each other (Ito, 1985b, 1999a).

Lepidostoma kanbaranum (Kobayashi 1968)
Crunoeciella kanbarana Kobayashi 1968, 10-11, pl. 6, male, Honshu (Niigata).
Goerodes kanbaranus: Ito 1990b, 786-789, male, female, larva, case, habitat, Honshu (Niigata).

Ito et al. 1993, 145.

Lepidostoma kanbaranum: Weaver 2002, 185.

Ito 2005a, 505, 509, 515, male, larva, case, habitat.
Ito 2005d, 202.
Nozaki et al. 2016, 329, Honshu (Niigata).
Tanida 2016, 107.
Ito 2018, 599, 603, 611, male, larva.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Honshu (central)).

Lepidostoma kantoense (Ito 1994)
Goerodes kantoensis Ito 1994, 84-87, male, female, larva, case, habitat, Honshu (Tochigi, Tokyo,Yamanashi).

Moriya 1997, 69, Honshu (Kanagawa).
Nozaki 1997a, 37, Honshu (Kanagawa).
Kagaya et al. 1998, 66, Honshu (Yamanashi).

Lepidostoma kantoense: Weaver 2002, 181.

Ito 2005a, 505, 507, 513, male, larva, case, habitat.
Ito 2005d, 202.
Moriya and Kubota 2013, 136, Honshu (Kanagawa).
Tanida 2016, 107.
Katsuma 2017, 37, 41, Honshu (Ibaraki).
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Yamanashi), Japan.
Distribution. Japan (Honshu (central)).

Lepidostoma kasugaense (Tani 1971)
Dinarthrodes kasugaensis Tani 1971, 63-64, male, Honshu (Shiga, Hyogo, Nara).

Tani 1977, 200, male.
Ito et al. 1993, 144.
Moriya 1994, 82, Honshu (Kanagawa, Yamanashi).
Nozaki 1997b, 30, Honshu (Tokyo).
Kagaya et al. 1998, 64, 208, habitat, Honshu (Tokyo, Yamanashi).

Goerodes kasugaensis: Tanida and Takemon 1993, 243, Honshu (Kyoto).

Ito 1999a, 323-325, male, female, larva, case, habitat, Honshu (Iwate, Toyama, Yamanshi, Gifu, Shizuoka, Aichi, Mie, Wakayama), Kyushu (Oita, Kagoshima).
Ito et al. 2002b, 26, adult flight season, Shikoku (Kagawa, Ehime, Kochi).
Aoyagi et al. 2004, 26, Honshu (Kyoto).

Lepidostoma kasugaense: Weaver 2002, 181.

Ito 2005a, 505, 508, 514, male, larva, case, habitat.
Ito 2005d, 202.
Katsuma 2005a, 7, Honshu (Ibaraki).
Morita 2006, 200, Honshu (Mie).
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Kimura et al. 2007, 167, Honshu (Nagano).
Morita 2007a, 14, Honshu (Mie).
Inazu 2008, 81, 94, flight period, Honshu (Hyogo).
Morita 2008a, 6, Honshu (Mie).
Aoyagi and Iwasaki 2009, 101, 102, 104, 106, 107, Honshu (Osaka).
Morita 2009, 141, Honshu (Mie).
Kobayashi 2010, 141, Honshu (Tochigi).
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Inazu and Nishida 2011, 199, male, female, Honshu (Hyogo).
Morita 2011, 216, Honshu (Mie).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Inazu 2013c, 74, Honshu (Hyogo).
Nishimoto and Nishimoto 2014, 66, Honshu (Aichi).
Horiguchi 2016, 413, Honshu (Tokyo).
Tanida 2016, 107-108.
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Ito 2017, 5, Honshu (Fukui).
Kawase et al. 2017, 34, Honshu (Shiga).
Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 599, 603, 610, male, larva.
Morita 2020b, 81, Honshu (Mie).

Type locality. Honshu (Nara), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu).

Lepidostoma kojimai (Tani 1971)
Paradinarthrodes kojimai Tani 1971, 61, male, wings of female, Honshu (Saitama).

Tani 1977, 200, male.
Ito and Asahina 1990, 20, male, larva, case, habitat, Honshu (Shizuoka).

Goerodes kojimai: Ito 1990b, 783-786, male, female, larva, case, habitat, Honshu (Shizuoka).

Ito et al. 1993, 145.

Lepidostoma kojimai: Weaver 2002, 185.

Ito 2005a, 505, 508, 514, male, larva, case, habitat.
Ito 2005d, 202.
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2007c, 18, Honshu (Iwate).
Nozaki and Tanida 2007, 251, Honshu (Shizuoka).
Tanida and Nozaki 2010, 89, adult, Honshu (Shizuoka).
Inazu and Nishida 2011, 200, male, Honshu (Hyogo).
Katsuma 2011b, 68, Honshu (Ibaraki).
Katsuma 2012, 66, Honshu ( Ibaraki).
Kuranishi and Kikuchi 2014, 363, Honshu (Yamagata).
Tanaka 2014, 34, 40, male, Honshu (Akita).
Katsuma 2015, 91, Honshu (Ibaraki).
Tanida 2016, 108.
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima).
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 599, 603, 610, male, larva.
Tanida et al. 2018, 30-31, Honshu (Tottori).
Nozaki et al. 2019, 170, 174, phenology, Honshu (Nagano).
Tanaka 2019, 12, Honshu (Akita).
Nojima 2022, 38, Honshu (Okayama).
Tanaka 2023, 43, Honshu (Akita).

Type locality. Honshu (Saitama), Japan.
Distribution. Japan, Honshu (central).

Lepidostoma konosense Ito 2011
Lepidostoma konosense Ito 2011, 167-168, male, female, larva, case, habitat, Shikoku (Tokushima, Ehime).

Tanida 2016, 108.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Shikoku (Tokushima), Japan.
Distribution. Japan (Shikoku).

Lepidostoma kumanoense (Ito 1994)
Goerodes kumanoensis Ito 1994, 82-84, male, female, larva, case, habitat, Honshu (Mie).

Morita 1999a, 27, Honshu (Mie).
Morita 2001d, 339, Honshu (Mie).

Lepidostoma kumanoense: Weaver 2002, 181.

Ito 2005a, 505, 507, 513, male, larva, case, habitat.
Ito 2005d, 202.
Morita 2006, 197, 200, Honshu (Mie).
Morita 2007a, 14, Honshu (Mie).
Morita 2008d, 92, Honshu (Mie).
Morita 2009, 141, Honshu (Mie)
Kawase and Morita 2010, 34, 44, Honshu (Mie, Shiga).
Morita 2011, 216, Honshu (Mie).
Tanida 2016, 108.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Mie), Japan.
Distribution. Japan (Honshu (central)).

Lepidostoma kunigamiense (Ito 1999)
Goerodes kunigamiensis Ito 1999b, 494-499, male, female, larva, case, habitat, Okinawa-jima.
Lepidostoma kunigamiense: Weaver 2002, 181.

Ito 2005a, 506-507, 513, male, larva, case, habitat.
Ito 2005d, 202.
Tanida 2016, 108.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Okinawa-jima, Japan.
Distribution. Japan (Okinawa-jima).

Lepidostoma laeve (Ito 1984)
Dinarthrum laeve Ito 1984a, 9-12, male, female, pupa, larva, case, habitat, food, Hokkaido (Ishikari, Oshima), Honshu (Iwate).

Ito 1987, 271-272, variation of case, Hokkaido (Ishikari).
Ito et al. 1993, 144.

Lepidostoma laeve: Weaver 2002, 181.

Ito 2005a, 504, 507, 512, male, larva, case, habitat.
Ito 2005d, 202.
Muramatsu 2008, 7, Hokkaido (Okhotsuk).
Ito et al. 2010, 62, Hokkaido (Oshima).
Ito 2013, 61, 83, Hokkaido (Okhotsk).
Tanida 2016, 108.
Ito 2018, 598, 601, 608, male, larva.
Nojima 2022, 38-39, Honshu (Okayama).
Ito and Kuhara 2023, 65, Hokkaido (Soya).

Type locality. Hokkaido, Japan.
Distribution. Japan (Hokkaido, Honshu).

Lepidostoma mennokiense (Ito 2011)
Lepidostoma mennokiense Ito 2011, 167-168, male, female, larva, case, habitat, Honshu (Kanagawa, Shizuoka, Aichi).

Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Tanida 2016, 108.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Aichi), Japan.
Distribution. Japan (Honshu (central)).

Lepidostoma nanseiense (Ito 1990)
Goerodes nanseiensis Ito 1990a, 366-369, male, female, larva, case, habitat, Amami-Oshima.

Ito 1992a, 334-335, variation of male, Okinawa-jima.
Ito et al. 1993, 145.
Ito 1999b, 499, Amami-Oshima, Okinawa-jima.

Lepidostoma nanseiense: Weaver 2002, 185.

Ito 2005a, 506, 509, 515, male, larva, case, habitat.
Ito 2005d, 202, 205.
Shimura et al. 2015, 47, Amami-Oshima.
Tanida 2016, 108.
Ito 2018, 600, 604, 611, male, larva.

Type locality. Amami-Oshima, Japan.
Distribution. Japan (Amami-Oshima, Okinawa-jima).

Lepidostoma naraense (Tani 1971)
Crunobiodes naraensis Tani, 1971, 58-60, male, wings of female, Honshu (Nara), Kyushu (Fukuoka).

Tani 1977, 200, male.

Goerodes naraensis: Ito 1984c, 507-514.

Ito 1985c, 199-210, geographical variation of male, female, larva, case, habitat, food, life cycle, Hokkaido (Ishikari, Shiribeshi), Honshu (Iwate, Tochigi, Gunma, Niigata, Nagano, Shiga, Hyogo).
Morita 1989, 41, Honshu (Mie).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 145.
Morita 1993, 7, Honshu (Mie).
Ito 1994, 87-88, Hokkaido (Nemuro), Honshu (Akita, Yamagata, Tokyo, Kanagawa, Yamanashi, Wakayama, Tottori, Hiroshima), Kyushu (Fukuoka, Oita, Kumamoto).
Morita 1994a, 47, Honshu (Mie).
Morita 1994b, 98, Honshu (Mie).
Moriya 1994, 82, Honshu (Kanagawa).
Morita 1996, 303, Honshu (Mie).
Moriya 1997, 70, Honshu (Kanagawa).
Nozaki 1997a, 37, Honshu (Kanagawa).
Ito et al. 1998, 11, 17, Hokkaido (Nemuro).
Kagaya et al. 1998, 66, 215, habitat, Honshu (Tokyo, Yamanashi).
Ito et al. 2000b, 37, Hokkaido (Ishikari).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Ito et al. 2002b, 25-26, flight period, sex ratio, Shikoku (Kagawa, Ehime, Kochi).
Morita 2003, 26, Honshu (Mie).

Lepidostoma naraense: Weaver 2002, 182.

Ito 2005a, 504, 507, 511, 513, male, larva, case, habitat.
Ito 2005d, 202-203.
Nagayasu et al. 2005, 56, Hokkaido (Ishikari).
Morita 2006, 200, Honshu (Mie).
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Morita 2007a, 14, Honshu (Mie).
Inazu 2008, 81, 94, flight period, Honshu (Hyogo).
Morita 2008a, 6, Honshu (Mie).
Morita 2008d, 92-93, Honshu (Mie).
Morita 2009, 141, Honshu (Mie)
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Ito et al. 2010, 62, Hokkaido (Oshima).
Inazu and Nishida 2011, 200, male, Honshu (Hyogo).
Morita 2011, 216, Honshu (Mie).
Kuhara 2011, 53, 63, 74, flight season, Hokkaido (Sorachi, Ishikari).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 61, 83, Hokkaido (Nemuro, Tokachi).
Yamamoto and Ito, 2014, 14, 18-21, Shikoku (Ehie).
Ito and Nagasaka 2014, 11-12, Hokkaido (Sorachi).
Nishimoto and Nishimoto 2014, 66, Honshu (Aichi).
Nozaki 2016, 79, 367-368, male, Shikoku (Kochi).
Tanida 2016, 108.
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Katsuma 2017, 41, Honshu (Ibaraki).
Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 599, 601, 607, 609, male, larva, case.
Shimura and Imasaka 2018, 172, Kyushu (Fukuoka, Oita).
Shimura and Imasaka 2020, 14, Kyushu (Fukuoka, Oita).
Taira 2020, 8, Honshu (Nara).
Yamamoto et al. 2021, 74, Shikoku (Ehime).
Ito and Kuhara 2023, 65, Hokkaido (Soya, Kamikawa).

Type locality. Honshu (Nara), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (South Kurils).

Lepidostoma niigataense Ito 2016
Lepidostoma niigataense Ito 2016a, 403-407, male, female, habitat, Honshu (Miyagi, Yamagata, Kanagawa, Niigata, Aichi).

Nojima 2017, 121, Honshu (Okayama).
Ito 2018, 598, 601, 608, male.
Kawase and Kato 2023, 8, Honshu (Niigata).

Type locality. Honshu (Niigata), Japan.
Distribution. Japan, Honshu.

Lepidostoma orientale (Tsuda 1942)
Crunoeciella orientalis Tsuda 1942a, 329-331, male, wings of female, Honshu (Shiga, Kyoto).

Tsuda 1942b, 263-264, Honshu (Kyoto).
Tsuda 1943, 166, Honshu (Kyoto).
Tsuda and Tatsumi 1951, 30, Honshu (Niigata).
Kobayashi 1959, 354, Kyushu (Fukuoka).
Fischer 1970, 82.
Tani 1971, 51-52, male, wings of female, Honshu (Shiga, Nara), Shikoku (Tokushima), Tsushima.
Tani 1976, 393, Tsushima.
Tani 1977, 200, male.
Kobayashi 1984c, 6, Sado.
Kadowaki 1998, 7, Honshu (Shimane).

“Crunoeciella” orientalis: Malicky 1979, 100-101.
Goerodes orientalis: Ito 1984c, 507-514.

Ito 1985a, 16-19, female, pupa, larva, case, habitat, Honshu (Kyoto).
Ito 1989, 57, Tsushima.
Ito 1990b, 790, Honshu (Tokyo, Shizuoka).
Ito and Asahina 1990, 20-21, Honshu (Shizuoka).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 146.
Ito 1994, 91, Honshu (Wakayama, Okayama).
Nishiharu 1997, 61, Honshu (Fukui).
Tani and Nakamura 1997, 65, Honshu (Hiroshima).
Morita 1998b, 73, Honshu (Mie).
Morita 2001c, 175, Honshu (Mie).
Ito et al. 2002b, 25-26, Shikoku (Ehime).
Nozaki and Nakamura 2002, 175, Honshu (Hiroshima).
Kawase et al. 2004, 115, Honshu (Ibaraki).

Lepidostoma orientale: Weaver 2002, 186.

Uenishi and Yoshitake 2004, 18, Honshu (Okayama).
Ito 2005a, 505, 508, 515, male, larva, case, habitat.
Ito 2005d, 203.
Nozaki and Nakamura 2005, 223, Honshu (Hiroshima).
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Morita 2006, 200, Honshu (Mie).
Nakamura et al. 2006, 102, Honshu (Hiroshima).
Katsuma 2007a, 18, Honshu (Tochigi).
Morita 2008a, 6, Honshu (Mie).
Nakamura and Hajiri 2007, 199, Honshu (Shimane).
Nozaki and Tanida 2007, 252, Honshu (Shizuoka).
Katsuma 2008, 58, Honshu (Ibaraki).
Morita 2008a, 6, Honshu (Mie).
Kawase and Hayashi 2010a, 80, Honshu (Shimane).
Mano 2010, 21-23, Honshu (Aichi).
Nakamura 2010, 239, Honshu (Hiroshima).
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Kawase and Hayashi, 2011, 192, Honshu (Shimane).
Inazu and Nishida 2011, 200, male, female, Honshu (Hyogo).
Morita 2011, 216-217, Honshu (Mie).
Inazu 2012, 9, Honshu (Hyogo).
Morita 2012, 251, Honshu (Mie).
Inazu 2013b, 57, Honshu (Hyogo).
Yamamoto 2014, 32-33, adult, Shikoku (Ehime).
Yamamoto and Ito 2015, 33, Shikoku (Ehime).
Katsuma 2015, 91, Honshu (Ibaraki).
Horiguchi 2016, 413, Honshu (Tokyo).
Nozaki, 2016, 80, 367-368, male, Shikoku (Kochi).
Tanida 2016, 108-109.
Yamamoto and Ito 2016, 41, 43, 44, Shikoku (Ehime).
Ito 2017, 5, Honshu (Fukui).
Kawase et al. 2017, 34, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga).
Nojima 2017, 121-122, Honshu (Okayama).
Saito et al. 2017, 184, Honshu (Chiba).
Ito 2018, 599, 603, 611, male, larva.
Nozaki et al. 2019, 170, phenology, Honshu (Nagano).
Yamamoto et al. 2019, 73, Shikoku (Ehime).
Katsuma 2020, 15, Kyushu (Oita).
Tezuka et al. 2021, Honshu (Kanagawa).
Shimura and Imasaka 2021, 5, Kagoshima (Koshiki-retto).
Yamamoto and Nozaki 2021, 129, 132, adult appearance months, Shikoku (Ehime).
Kuhara et al. 2023, 18-19, Kyushu (Kumamoto, Kagoshima).
Yamamoto 2023, 33, Shikoku (Ehime).

Type locality. Honshu (Shiga, Kyoto), Japan.
Distribution. Japan (Honshu (central, west), Shikoku, Kyushu, Sado, Tsushima, Koshiki-retto), Russia (continental part of Far East), China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan).

Lepidostoma pseudemarginatum Ito 2011
Lepidostoma pseudemarginatum Ito 2011, 159-61, male, female, larva, case, habitat, Honshu (Niigata, Hyogo).

Inazu 2013c, 74-75, Honshu (Hyogo).
Tanida 2016, 109.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan, Honshu (central, west).

Lepidostoma robustum (Ito 1984)
Dinarthrum robustum Ito 1984a, 1-9, male, female, pupa, larva, case, habitat, food, life cycle, Hokkaido (Ishikari, Oshima), Honshu (Miyagi, Niigata).

Ito 1987, 271-272, case, Hokkaido (Ishikari).
Ito et al. 1993, 144.
Kuhara and Ito 1994, 21, 23, Hokkaido (Ishikari).
Ohkawa 1999, 40, 44, Hokkaido (Tokachi).
Ito et al. 2000b, 36, Hokkaido (Ishikari).
Kuhara et al. 2000, 256, Hokkaido (Ishikari).

Lepidostoma robustum: Weaver 2002, 182.

Ito 2005a, 504, 507, 512-513, male, larva, case, habitat.
Ito 2005d, 203.
Nagayasu et al. 2005, 56, Hokkaido (Ishikari).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Kuhara et al. 2007, 72, Hokkaido (Ishikari).
Kuhara 2011, 53, 74, flight season, Hokkaido (Sorachi, Ishikari).
Ito 2013, 61, 83, Hokkaido (Kushiro, Tokachi).
Ito and Nagasaka 2014, 12, Hokkaido (Sorachi).
Tanaka 2016, 2, 5, male, Honshu (Akita).
Tanida 2016, 109.
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 598, 600, 607, 608, male, larva, case.
Tanaka 2022, 36, Honshu (Akita).
Ito and Kuhara 2023, 65-66, Hokkaido (Soya, Kamikawa).

Type locality. Hokkaido (Oshima), Japan.
Distribution. Japan (Hokkaido, Honshu (north, central)).

Lepidostoma ryukyuense (Ito 1992)
Goerodes ryukyuensis Ito 1992a, 335-337, male, female, larva, case, habitat, Okinawa-jima.

Ito et al. 1993, 146.
Ito 1999b, 499-500, Okinawa-jima.

Lepidostoma ryukyuense: Weaver 2002, 182.

Ito 2005a, 506, 508, 514, male, larva, case, habitat.
Ito 2005d, 203.
Tanida 2016, 109.
Ito 2018, 599, 603, 610, male, larva.

Type locality. Okinawa-jima, Japan.
Distribution. Japan (Okinawa-jima).

Lepidostoma satoi (Kobayashi 1968)
Dinarthrodes satoi Kobayashi 1968, 9-10, pl. 5, male, Honshu (Niigata).

Ito 1978, 581-582, larva, case.
Ito 1980, 313-316, life cycle, Hokkaido (Ishikari).
Tanabe 1980, 65, adult, Hokkaido (Ishikari).

Gen. satoi: Tani, 1978, 10-11, male, Honshu (Niigata).
Goerodes satoi: Ito 1983c, 269-273, habitat, Hokkaido (Rumoi, Ishikari, Oshima).

Ito 1984b, 240-246, drift of larva, Hokkaido (Ishikari).
Ito 1984c, 507-514.
Ito 1985b, 263-264, female, pupa, larva.
Nozaki 1987, 37, Honshu (Kanagawa).
Kagaya et al. 1998, 66, Honshu (Tokyo, Yamanashi).
Nozaki 1988a, 46, Honshu (Kanagawa).
Nozaki 1988b, 42, Honshu (Kanagawa).
Ito and Asahina 1990, 21, Honshu (Tokyo).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Kuhara 1992, 46, Hokkaido (Shiribeshi).
Ito et al. 1993, 146.
Morita 1993, 7, Honshu (Mie).
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Moriya 1994, 81, Honshu (Kanagawa, Yamanashi).
Itou and Muramatsu 1995, 42, Hokkaido (Okhotsuk).
Ito et al. 1997, 24, 26, Hokkaido (Oshima).
Moriya 1997, 37, Honshu (Kanagawa).
Ito et al. 1998, 11-14, Hokkaido (Nemuro).
Kagaya et al. 1998, 4-5, 217, habitat, Honshu (Tokyo, Yamanashi).
Ito et al. 1999, 24, 27, Hokkaido (Kushiro).
Ito et al. 2000a, 27-28, Hokkaido (Soya).
Ito et al. 2000b, 36, Hokkaido (Ishikari).
Kuhara et al. 2000, 256, Hokkaido (Iburi).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Ito et al. 2002b, 25, flight season, sex ratio, Shikoku (Ehime, Kochi).
Uchida 2002, 78, Honshu (Tochigi).
Kawase et al. 2004, 115, Honshu (Ibaraki).
Morita and Kawase 2004, 16, Honshu (Ishikawa).

Lepidostoma satoi: Weaver 2002, 182.

Uenishi and Yoshitake 2004, 28, Honshu (Okayama).
Ito 2005a, 505, 508, 510, 514, male, larva, case, habitat.
Ito 2005c, 189-197, food.
Ito 2005d, 203.
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Nishio 2006, 41-42, emergence, copulation, Honshu (Nagano).
Kuhara et al. 2006, 45-47, Hokkaido (Ishikari).
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Ito 2007, 155-161, emergence period.
Ito et al. 2007, 149, Hokkaido (Kushiro).
Kuhara et al. 2007, 72, Hokkaido (Ishikari).
Inazu 2008, 81, 94, flight period, Honshu (Hyogo).
Morita 2008a, 6, Honshu (Mie).
Morita 2008b, 17, Honshu (Mie).
Morita 2008c, 6, Honshu (Shiga).
Muramatsu 2008, 8, Hokkaido (Okhotsk).
Aoyagi and Iwasaki 2009, 101, 102, 104, 107, Honshu (Osaka).
Morita 2009, 141, Honshu (Mie)
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Ito et al. 2010, 63, Hokkaido (Shiribeshi, Oshima).
Inazu and Nishida 2011, 200, male, Honshu (Hyogo).
Katsuma 2011a, 49, Honshu (Ibaraki).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Kuhara 2011, 53, 63, 74, flight season, Hokkaido (Sorachi, Ishikari, Hidaka).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 61, 83, Hokkaido (Nemuro, Okhotsk, Kushiro, Tokachi).
Kimura and Hirabayashi 2013, 185, Honshu (Nagano).
Inazu 2013a, 46, Honshu (Hyogo).
Inazu 2013c, 75, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Yamamoto and Ito 2014, 14, 18, Shikoku (Ehime).
Ito and Nagasaka 2014, 12, Hokkaido (Sorachi).
Tanaka 2014, 37, 40, male, Honshu (Akita).
Nozaki et al. 2014, 62, Honshu (Nagano).
Ito 2016b, 1-8, protogyny.
Nozaki, 2016, 80, 368, male, Shikoku (Kochi).
Tanaka 2016, 2, Honshu (Akita).
Tanida 2016, 109.
Hayashi et al. 2017, 80, Kyushu (Miyazaki).
Ito 2017b, 44, Hokkaido (Kamikawa).
Katsuma 2017, 41, Honshu (Ibaraki).
Kawase et al. 2017, 34, Honshu (Shiga).
Nojima 2017, 122, Honshu (Okayama).
Ito 2018, 59, 603, 610, male, larva.
Nozaki et al. 2019, 170, 174, phenology, Honshu (Nagano).
Tanaka 2019, 12, Honshu (Akita).
Ito 2020, 17-22, spermatophore, Hokkaido (Ishikari).
Shimura and Imasaka 2020, 15, Kyushu (Fukuoka, Oita).
Yamamoto et al. 2021, 75, Shikoku (Ehime).
Tanaka 2022, 36, Honshu (Akita).
Ito and Kuhara 2023, 66, Hokkaido (Soya, Okhotsuk, Kamikawa, Rumoi).
Tanaka 2023, 43, Honshu (Akita).

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Hokkaido, Honshu Shikoku, Kyushu), Russia (South Kuriles).

Lepidostoma semicirculare (Ito 1994)
Goerodes semicircularis Ito 1994, 88-90, male, female, larva, case, habitat, Honshu (Aomori, Akita, Niigata).
Lepidostoma semicirculare: Weaver 2002, 186.

Ito 2005a, 505, 509, 515, male, larva, case, habitat.
Ito 2005d, 203.
Katsuma 2005a, 7-8, Honshu (Ibaraki).
Katsuma 2007b, 18, Honshu (Ibaraki).
Katsuma 2011b, 68, Honshu (Ibaraki).
Tanaka 2014, 36, 41, male. Honshu (Akita).
Tanida 2016, 109.
Aoya 2018, 5, male, Honshu (Akita).
Ito 2018, 599, 604, 611, male, larva.
Tanaka 2023, 43, Honshu (Akita).

Type locality. Honshu (Akita), Japan.
Distribution. Japan (Honshu (north, central)).

Lepidostoma spathulatum (Ito 1989)
Goerodes spathulatus Ito 1989, 55-57, male, female, larva, case, habitat, Tsushima.

Ito et al. 1993, 146.

Lepidostoma spathulatum: Weaver 2002, 186.

Ito 2005a, 506-507, 513, male, larva, case, habitat.
Ito 2005d, 203.
Tanida 2016, 109.
Ito 2018, 599, 603, 610, male, larva.

Type locality. Tsushima, Japan.
Distribution. Japan (Tsushima).

Lepidostoma speculiferum (Matsumura 1907)
Pycnocentria speculifera Matsumura 1907, 191, adult.
Dinarthrodes nukabiraensis Kobayashi 1964, 88-90, male. Synonymized by Ito (1999a).

Tani 1971, 69-70, male, Honshu (Kyoto).
Kobayashi 1977, 2, Hokkaido (Tokachi).
Tani 1977, 200, male.
Tani 1978, 11, Honshu (Niigata).
Yamanouchi 1976, 3, Hokkaido (Tokachi).
Ito 1978, 582-583, larva, case.
Yamanouchi 1978, 10, Hokkaido (Tokachi).
Kadowaki 1998, 7, Honshu (Shimane).

Goerodes nukabiraensis: Ito 1983c, 269-275, habitat, life cycle, Hokkaido (Rumoi, Ishikari, Oshima).

Ito 1984b, 240-246, drift of larva, Hokkaido (Ishikari).
Ito 1984c, 507-514.
Ito 1985b, 264, 268, female, pupa, larva.
Kobayashi 1987, 22, Honshu (Shimane).
Nozaki 1988b, 42, Honshu (Kanagawa).
Ito et al. 1993, 146.
Nishimoto 1993, 22, Hokkaido (Kushiro).
Kuranishi and Kuhara 1994, 1230, Hokkaido (Kushiro).
Moriya 1994, 81, Honshu (Kanagawa, Yamanashi).
Nakajima et al. 1997, 14, 18-19, 28, 30, 31, Hokkaido (Oshima).
Ito et al. 1997, 24, Hokkaido (Oshima).
Kuhara 1997a, 62, Hokkaido (Shiribeshi).
Moriya 1997, 70, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 37, Honshu (Kanagawa).
Ito et al. 1998, 11, Hokkaido (Nemuro).
Kagaya et al. 1998, 66, 216, habitat, Honshu (Tokyo, Yamanashi).
Ito et al. 1999, 24-27, Hokkaido (Kushiro).
Ohkawa 1999, 40, 44, Hokkaido (Tokachi).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Kameyama and Nakamura 2006, 190, Honshu (Tottori).

Goerodes speculifer: Ito 1999a, 326-328, variation of male genitalia, female, new synonym, Hokkaido (Nemuro, Okhotsuk, Kushiro, Tokachi, Rumoi, Kamikawa, Sorachi, Ishikari, Iburi, Oshima), Honshu (Aomori, Iwate, Akita, Fukushima, Tochigi, Toyama, Niigata, Yamanashi, Nagano, Tottori, Shimane, Okayama), Kyushu (Oita, Kumamoto).

Ito et al. 2000b, 37, Hokkaido (Ishikari).
Kuhara et al. 2000, 257, Hokkaido (Iburi).
Uchida 2002, 78, Honshu (Tochigi).
Nagayasu et al. 2003, 110, Hokkaido (Ishikari).
Shinoda 2003, 145, Honshu (Yamanashi).

Lepidostoma speculiferum: Weaver 2002, 183.

Ito 2005a, 505, 508, 514, male, larva, case, habitat.
Ito 2005d, 203.
Katsuma 2005a, 7-8, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Gunma).
Nagayasu et al. 2005, 57, Hokkaido (Ishikari).
Mey and Nozaki 2006, 25, Honshu (Nagano).
Ito and Kosugi 2007, 54, Hokkaido (Kushiro).
Nakano et al. 2007. 40, Hokkaido (Nemuro).
Muramatsu 2008, 7, Hokkaido (Okhotsuk).
Ito et al. 2009, 9, Hokkaido (Okhotsuk).
Ito 2010, 131, Hokkaido (Kushiro).
Kobayashi 2010, 141, Honshu (Tochigi).
Tsuji et al. 2010, 6, Hokkaido (Okhotsk).
Ito et al. 2010, 63, Hokkaido (Shiribeshi, Hiyama, Oshima).
Tanaka 2010a, 25, 27-28, male, Honshu (Akita).
Tanaka 2010b, 17-19, male, Honshu (Akita).
Inazu and Nishida 2011, 200, male, Honshu (Hyogo).
Ito and Kubo 2011, 8, Hokkaido (Tokachi).
Katsuma 2011b, 68, Honshu (Ibaraki).
Kuhara 2011, 53, 63, 74, flight season, Hokkaido (Sorachi, Ishikari).
Katsuma 2012, 66, Honshu (Ibaraki).
Moriya and Kubota 2012, 136, Honshu (Kanagawa).
Ito 2013, 61, 83, Hokkaido (Nemuro, Okhotsuk, Kushiro, Tokachi).
Inazu 2013c, 75, Honshu (Hyogo).
Ijima and Tsuchiya 2014, 31, Hokkaido (Okhotsuk, Kushiro, Tokachi).
Ito and Nagasaka 2014, 12, Hokkaido (Sorachi).
Kawase and Morita 2014, 6, Honshu (Shiga).
Kuranishi and Kikuchi 2014, 363, Honshu (Yamagata).
Tanaka 2014, 36-37, 40, male, Honshu (Akita).
Nozaki et al. 2014, 62, Honshu (Nagano).
Katsuma 2015, 91, Honshu (Fukushima, Ibaraki).
Nozaki, 2016, 80, 368, male, Honshu (Okayama).
Tanida 2016, 109.
Ito 2017a, 5, Honshu (Fukui).
Ito 2017b, 44, Hokkaido (Kamikawa).
Nojima 2017, 122, Honshu (Okayama).
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima).
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 599, 602-603, 610, male, larva.
Nakatani and Ito 2019, 163, Hokkaido (Kushiro).
Abe and Ito 2021, 35, mite parasite, Hokkaido (Kushiro).
Nakatani 2021, 72, Hokkaido (Kushiro).
Tanaka 2022, 36, Honshu (Akita).
Aoya and Tanaka 2023, 27, Honshu (Akita).
Ito and Kuhara 2023, 66, Hokkaido (Soya, Okhotsuk, Kamikawa, Rumoi).
Tanaka 2023, 43, Honshu (Akita).

Lepidostoma japonicum: Ijima and Tsuchiya 2014, 31, Hokkaido (Okhotsk, Kushiro, Tokachi). Misidentification, mentioned by Ito and Kuhara (2016).
Type locality. Japan.
Distribution. Japan (Hokkaido, Honshu, Kyushu).

Lepidostoma stellatum (Ito 1984)
Dinarthrum stellatum Ito 1984a, 12-15, male, female, pupa, larva, case, habitat, food, life cycle, Hokkaido (Tokachi, Rumoi, Sorachi, Ishikari, Shiribeshi).

Ito et al. 1993, 144.
Ito and Minakawa 1995, 667-668, variation of male, Hokkaido (Sorachi).
Ito et al. 1998, 11, 16, Hokkaido (Nemuro).
Kushiro Insect Lover’s Society 1999, 219, Hokkaido (Nemuro).
Ito et al. 2000b, 36, Hokkaido (Ishikari).
Kuhara et al. 2000, 256, Hokkaido (Ishikari).
Nagayasu et al. 2003, 110, Hokkaido (Ishikari).

Lepidostoma stellatum: Weaver 2002, 183.

Ito 2005a, 504, 506, 512, male, larva, case, habitat.
Ito et al. 2007, 149, Hokkaido (Kushiro).
Ito 2005d, 204.
Ito et al. 2010, 63, Hokkaido (Shiribeshi, Oshima).
Ito and Nakatani 2011, 52, 147, adult, larva, Hokkaido (Kushiro).
Ito 2013, 61, 83, Hokkaido (Nemuro, Kushiro, Tokachi).
Ijima and Tsuchiya 2014, 31, Hokkaido (Okhotsuk).
Tanida 2016, 109.
Ito 2018, 598, 601, 608, male, larva.
Abe and Ito 2021, 35, mite parasite, Hokkaido (Ishikari).
Ito and Kuhara 2023, 65, Hokkaido (Soya).

Type locality. Hokkaido (Tokachi), Japan.
Distribution. Japan (Hokkaido), Russia (Sakhalin, South Kurils).
Remarks. Type locality described by Tanida (2016) is incorrect.

Lepidostoma tsudai (Tani 1971)
Yamatopsyche tsudai Tani, 1971, 55-58, male, wings of female, Honshu (Fukui, Nagano, Shiga, Nara).

Tani 1977, 200, male, Honshu (Shiga).
Tani 1978, 12, Honshu (Niigata).
Kadowaki 1998, 7, Honshu (Shimane).

Goerodes tsudai: Ito 1984b, 507-514.

Ito 1985a, 19-20, female, larva, case, habitat, food, Honshu (Niigata, Okayama).
Togashi and Tanida 1985, 69, Honshu (Ishikawa).
Ito 1990b, 791, Honshu (Shizuoka, Niigata, Fukui).
Ito and Asahina 1990, 21, Honshu (Shizuoka).
Gyotoku and Nozaki 1992, 139, Kyushu (Fukuoka).
Ito et al. 1993, 146.
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Ito 1994, 90-91, Honshu (Ishikawa, Tottori, Shimane, Okayama), Kyushu (Oita).
Nishiharu 1997, 61, Honshu (Fukui).
Tani and Nakamura 1997, 65, Honshu (Hiroshima).
Morita 1998c, 97, Honshu (Mie).
Sekimoto et al. 1998, 361, Honshu (Hiroshima).
Morita 2000, 59, Honshu (Mie).
Morita 2001b, 175, Honshu (Mie).
Aoyagi et al. 2004, 26, Honshu (Kyoto).
Kawase et al. 2004, 115, Honshu (Ibaraki).
Morita and Kawase 2004, 16, Honshu (Ishikawa).
Kameyama and Nakamura 2006, 190, Honshu (Tottori).
Sasaji et al. 2013, 274, Honshu (Fukui).

Lepidostoma tsudai: Weaver 2002, 187.

Ito 2005a, 505, 509, 5135, male, larva, case, habitat.
Ito 2005d, 204.
Nozaki and Nakamura 2005, 223, Honshu (Hiroshima).
Morita 2006, 200, Honshu (Mie).
Nozaki and Tanida 2007, 252, Honshu (Shizuoka).
Inazu 2008, 82, 94, flight period, Honshu (Hyogo).
Morita 2009, 141, Honshu (Mie)
Katsuma 2010, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 44, Honshu (Mie, Shiga).
Inazu and Nishida 2011, 201, male, female, Honshu (Hyogo).
Inazu 2012, 9, Honshu (Hyogo).
Katsuma 2012, 66, Honshu (Ibaraki).
Morita 2012, 251, Honshu (Mie).
Hayashi et al. 2013, 13, Oki-no-shima.
Inazu 2013a, 46, Honshu (Hyogo).
Inazu 2013b, 57, Honshu (Hyogo).
Inazu 2013c, 75, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Tanaka 2014, 37, 40, male, Honshu (Akita).
Nozaki et al. 2014, 62, Honshu (Nagano).
Nozaki, 2016, 80, 368-369, male, Honshu (Matsumoto).
Nozaki et al. 2016, 329, Honshu (Niigata).
Tanida 2016, 109.
Ito 2017a, 5, Honshu (Fukui).
Katsuma 2017, 41, Honshu (Ibaraki).
Kawase et al. 2017, 34, Honshu (Shiga).
Ohira and Tsutsumi 2017, 25, Honshu (Fukushima).
Tanaka 2017, 3, Honshu (Akita).
Ito 2018, 599, 604, 611, male, larva.
Tsuruda 2018, 27, adult appearnce period, Honshu (Tokyo).
Kawase and Kato 2023, 8, Honshu (Niigata).
Tanaka 2023, 43, Honshu (Akita).

Type locality. Honshu (Nara), Japan.
Distribution. Japan (Honshu (central to west), Kyushu, Oki-no-shima).

Lepidostoma yakushimaense (Ito 1990)
Dinarthrum yakushimaense Ito 1990a, 362-366, male, female, larva, case, habitat, Yakushima (yakushimaensis).

Ito et al. 1993, 144 (yakushimaensis).

Lepidostoma yakushimaense: Weaver 2002, 183.

Ito 2005a, 506-507, 512, male, larva, case, habitat.
Ito 2005d, 204.
Tanida 2016, 110.
Ito 2018, 600-601, 608, male, larva.

Type locality. Yakushima, Japan.
Distribution. Japan (Yakushima).

Lepidostoma yosakoiense Ito 2011
Leoidostoma yosakoiensis Ito 2011, 165-167, male, female, larva, case, habitat, Shikoku (Kochi, Ehime).

Yamamoto 2014, 14, 18-20, Shikoku (Ehime).
Tanida 2016, 111.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Kochi), Japan.
Distribution. Japan (Shikoku).

Lepidostoma yunotaniense Ito 2011
Lepidostoma yunotaniense Ito 2011, 167-168, male, female, larva, case, habitat, Honshu (Tochigi, Niigata).

Tanida 2016, 111.
Ito 2018, 599, 602, 609, male, larva.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Honshu (central)).

Lepidostoma yuwanense Ito 2016
Lepidostoma yuwanense Ito 2016a, 412-414, male, female, habitat, Ryukyu Islands (Amami-Oshima, Okinawa-jima)

Ito 2018, 598, 601, 608, male.

Type locality. Ryukyu (Amami-Oshima), Japan.
Distribution. Japan (Ryukyu (central)).

Lasiocephala basalis: Iwata 1927a, b. See Brachycentridae.

Subfamily Theliopsychinae Weaver
A genus belonging to this subfamily has been recorded from Japan.

Genus Zephyropsyche Weaver
Two species belonging to this genus have been recorded from Japan.

Zephyropsyche monticola Ito, Kagaya and Hattori 2002
Zephyropsyche monticola Ito et al. 2002a, 122-128, male, female, pupa, larva, case, habitat, Honshu (Saitama, Niigata, Yamanashi, Shizuoka, Tottori).

Ito 2005a, 505, 509, 516, male, larva, case, habitat.
Ito 2005d, 204.
Inazu 2008, 80, 94, flight period, Honshu (Hyogo).
Inazu and Nishida 2011, 201, male, female, Honshu (Hyogo).
Inazu 2012, 9, Honshu (Hyogo).
Kawase and Morita 2014, 6, Honshu (Shiga).
Tanida 2016, 110.
Nojima 2017, 122, Honshu (Okayama).
Ito 2018, 599, 605, 612, male, larva, case.

Type locality. Honshu (Niigata), Japan.
Distribution. Japan (Honshu (central, west)).

Zephyropsyche odamiyamensis Ito and Yamamoto 2002
Zephyropsyche odamiyamensis Ito et al. 2002a, 128-130, male, female, pupa, larva, case, habitat, food, Shikoku (Ehime).

Ito et al. 2002b, 29, larva, case, habitat, flight season, sex ratio, Shikoku (Ehime, Kochi).
Ito 2005a, 505, 509, 516, male, larva, case, habitat.
Ito 2005d, 204.
Nozaki 2016, 80, 369, male, Shikoku (Kochi).
Tanida 2016, 110.
Ito 2018, 599, 605, 612, male, larva.

Type locality. Shikoku (Ehime), Japan.
Distribution. Japan (Shikoku).

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