Psychomyiidae

A catalogue of Japanese Trichoptera
Family Psychomyiidae Walker

Author: Takaaki TORII
E-mail: ttakaaki[AT]ideacon.co.jp
Last updated: 7 Feb. 2024

Three subfamilies, Eoneureclipsinae, Psychomyiinae and Tinodinae, belonging to this family have been recorded, and the genus Eoneureclipsis Kimmins was recently recorded from Japan.

Subfamily Eoneureclipsinae Mey

Genus Eoneureclipsis Kimmins

Four species belonging to this genus were described recently from Japan.

Eoneureclipsis montana Torii and Nishimoto 2011

Polycentropodidae Gen. sp.: Morita 1999b, 27-28, male, Honshu (Mie). Mentioned by Torii and Nishimoto (2011).

Morita 2001a, 220, Honshu (Mie).

Eoneureclipsis sp.: Katsuma 2006, 35, 43, male, Honshu (Ibaraki). Mentioned by Torii and Nishimoto (2011).

Torii and Hattori 2006, 35, 36 Honshu (Shizuoka). Mentioned by Torii and Nishimoto (2011).
Morita 2006, 197 Honshu (Mie) (as Eoneureeclipsis[!] sp.). Mentioned by Torii and Nishimoto (2011).
Morita 2008c, 92, Honshu (Mie). Mentioned by Torii and Nishimoto (2011).
Morita 2009, 139, Honshu (Mie). Mentioned by Torii and Nishimoto (2011).
Kawase and Morita 2010, 39, Honshu (Mie). Mentioned by Torii and Nishimoto.

Eoneureclipsis montana Torii and Nishimoto 2011, male, female, Honshu (Ibaraki, Shizuoka, Aichi, Mie) (montanus).

Morita 2011, 215, Honshu (Mie) (monatanus).
Katsuma 2012, 64, Honshu (Ibaraki) (montanus).
Tanida 2016, 86.
Torii and Nakamura 2016, 426-429, larva, Honshu (Nara). (montanus).
Nozaki 2018, 928, Honshu (Kanagawa).
Tanida 2018, 546, 549, 551, male, wings.

Type locality. Honshu (Mie), Japan.
Distribution. Japan (Honshu).

Eoneureclipsis shikokuensis Torii and Nishimoto 2011

Eoneureclipsis shikokuensis Torii and Nishimoto 2011, male, female, Shikoku (Tokushima, Kochi)

Nozaki 2016, 73, 336, 433, 450, adult, Shikoku (Tokushima).
Tanida 2016, 86.
Tanida 2018, 546, 551, male.

Type locality. Shikoku (Tokushima), Japan.
Distribution. Japan (Shikoku).

Eoneureclipsis okinawaensis Torii and Nishimoto 2011

Eoneureclipsis okinawaensis Torii and Nishimoto 2011, male, female, Okinawa (Okinawa-to)
Type locality. Okinawa (Okinawa-to), Japan.

Tanida 2016, 86.
Tanida 2018, 546, 551, male.

Distribution. Japan (Okinawa).

Eoneureclipsis yaeyamaensis Torii and Nishimoto 2011

Eoneureclipsis yaeyamaensis Torii and Nishimoto 2011, male, female, Okinawa (Ishigaki-jima, Iriomote-jima)

Tanida 2016, 86.
Tanida 2018, 546, 551, male.

Type locality. Okinawa (Ishigaki-jima), Japan.
Distribution. Japan (Okinawa).

Subfamily Psychomyiinae Walker

Three genera belonging to this subfamily have been recorded from Japan.

Genus Metalype Klapalek

One species belonging to this genus has been recorded from Japan.

Metalype uncatissima (Botosaneanu 1970)

Psychomyia uncatissima Botosaneanu 1970, 301-302, male, North Korea (Hamgjong).

Mey and Nozaki 2006, 24, Honshu (Nagano).

Metalype uncatissima: Morita 2001b, 338, Honshu (Mie).

Nozaki and Nakamura 2002, 171-172, Honshu (Hiroshima).
Tanida 2005, 466, 468, male, wings.
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Morita 2007b, 14, Honshu (Mie).
Nozaki and Nakamura 2007, 94, Honshu (Hiroshima).
Morita 2008d, 17, Honshu (Mie).
Torii 2011, 8-11, larva, pupa, Honshu (Aichi).
Kawase and Morita 2014, 5, Honshu (Shiga).
Katsuma 2015, 91, Honshu (Fukushima).
Nozaki 2016, 333-334, 433, male, female, wings.
Tanida 2016, 84.
Torii and Nakamura 2016, 429, larva, Honshu (Aichi).
Nojima 2017, 116, Honshu (Okayama).
Oláh et al. 2018, 106, North Korea (Chondjin).
Tanida 2018, 545, 549-550, male, wings.
Kuhara et al. 2023, 17, Kyushu (Ôita).

Type locality. North Korea.
Distribution. Japan (Honshu, Kyushu), Russia (South Primorye), North Korea.
Remarks. Psychomyia uncatissima Botosaneanu 1970 was transferred to the genus Metalype Klapalek by Li and Morse (1997a).

Genus Paduniella Ulmer

Five species belonging to this genus have been recorded from Japan.

Paduniella amamiensis Nishimoto 2011

Paduniella sp.: Tanida 1997, 446 (in part). Mentioned by Nishimoto (2011).
Paduniella amamiensis Nishimoto 2011, 385-386, male, Amami-Oshima (Kagoshima).

Tanida 2016, 84.
Tanida 2018, 545, 551, male.
Katsuma 2020, 15, Kyushu (Oita).

Type locality. Amami-Oshima (Japan).
Distribution. Japan (Kyushu, Amami-Oshima).

Paduniella communis Li & Morse 1997

Paduniella communis Li & Morse 1997b, male, China (An-hui).

Nishimoto 2011, 381-384, male, wings, Okinawa (Ishigaki-jima, Iriomote-jima).
Tanida 2016, 84.
Tanida 2018, 545, 549, 551, male, wings.

Paduniella sp.: Tanida 1997, 446 (in part). Mentioned by Nishimoto (2011).
Type locality. China.
Distribution. Japan (Okinawa), Eastern China.

Paduniella horaiensis Nishimoto 2011

Paduniella horaiensis Nishimoto 2011, 388-390, male, Honshu (Aichi, Hiroshima, Kochi).

Tanida 2016, 84.
Torii and Nakamura 2016, 429, larva, Honshu (Aichi).
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 551, male.
Katsuma 2020, 15, Kyushu (Oita).

Type locality. Aichi (Japan).
Distribution. Japan (Honshu, Shikoku, Kyushu).

Paduniella tanidai Nishimoto 2011

Paduniella sp.: Tanida and Nishimoto 1992, 28, Honshu (Shiga). Mentioned by Nishimoto (2011).
Paduniella amurensis: Tanida et al. 1999, 391, Honshu (Shiga). Misidentification mentioned by Nishimoto (2011).

Nishimoto and Morita 2001, 74, Honshu (Aichi). Misidentification mentioned by Nishimoto (2011).
Nozaki and Nakamura 2005, 220, Honshu (Shimane). Misidentification mentioned by Nishimoto (2011).
Kameyama et al. 2009, 111, Honshu (Hiroshima).
Kawase et al. 2017, 28, Honshu (Shiga).
Kobayashi et al. 2017, 17, Honshu (Shiga, Kyoto).

Paduniella tanidai: Nishimoto 2011, 387-388, male, Honshu (Aichi, Gifu, Shiga, Nara, Kyoto, Osaka, Shimane, Tokushima).

Morita and Katsuma 2011, 98, Honshu (Mie).
Tanida 2016, 84.
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 551, male.
Tanaka 2022, 35, Honshu (Akita).
Kuhara et al. 2023, 17, Kyushu (Saga).

Type locality. Aichi (Japan).
Distribution. Japan (Honshu, Shikoku, Kyushu).

Paduniella uralensis Martynov 1914

Paduniella uralensis Martynov 1914, 5-10, male, Russia (Ural Mounatains, Lakes Ilmen and Suratkul).

Fischer 1962, 165.
Fischer 1972, 68.
Nishimoto 2011, 384-385, male, Honshu (Hyogo, Shimane, Yamaguchi).
Tanida 2016, 84.
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 551, male.
Katsuma 2020, 15, Kyushu (Oita).

Type locality. Russia.
Distribution. Japan (Honshu, Kyushu), Russia (European Russia, Soth Primorye), China.

Genus Psychomyia Latreille

Twelve species belonging to this genus have been recorded from Japan. In addition, Akagi (1959) described Psychomyia sp. PA based on not only the larva but also adult male, but the specific status of this species has not yet been solved. Furthermore, an unnamed species accompanied with an alphabetic code was described for a larva, but it was not associated with adults. Tsuda (1942) pointed out that Psychomyia pussila (Fabricius, 1781) recorded by Iwata (1927) was a mis-identification. The genus Psychomyiella Ulmer was treated to the genus Psychomyia Latreille by Malicky (1993) and Li and Morse (1997a).

Psychomyia acutipennis (Ulmer 1908)

Psychomyiella acutipennis Ulmer 1908, 353-354, venation of fore and hind wings of female, Honshu (Utsubusa (Shizuoka?)).

Tsuda 1942a, 274-275, male, Honshu (Kyoto).
Tsuda 1942b, 263-264, Honshu (Kyoto).
Tsuda and Tatsumi 1951, 29, Honshu (Niigata).
Fischer 1962, 231.
Fischer 1972, 93.
Kobayashi 1984b, 4, Sado.
Gyotoku and Nozaki 1991, 73, Kyushu (Fukuoka).
Morita 1996b, 301, Honshu (Mie).
Moriya 1997, 61. Honshu (Kanagawa).
Nishimoto and Morita 2001, 76, Honshu (Aichi).

Psychomyia acutipennis: Nozaki and Nakamura 2002, 172, Honshu (Hiroshima).

Nozaki 2004, 1231, Honshu (Kanagawa).
Kawase et al. 2004, 112, Honshu (Ibaraki).
Ishiwata et al. 2005, 242-244, Honshu (Kanagawa).
Tanida 2005, 466, 468, male, wings.
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Nakamura et al. 2006, 101, Honshu (Hiroshima).
Katsuma 2007, 17, Honshu (Tochigi).
Nozaki and Nakamura 2007, 94, Honshu (Hiroshima).
Nozaki and Tanida 2007, 247, Honshu (Shizuoka).
Inazu 2008, 74, Honshu (Hyogo).
Morita 2008a, 5, Honshu (Mie).
Kameyama et al. 2009, 111, Honshu (Hiroshima).
Mano 2010, 21-23, Honshu (Aichi).
Katsuma 2010, 85, Honshu (Ibaraki).
Kawase and Morita 2010, 40, Honshu (Mie).
Inazu and Nishida 2011, 186, male, Honshu (Hyogo).
Katsuma 2011a, 49, Honshu (Ibaraki).
Kawase and Hayashi 2011b, 172, Honshu (Shimane).
Morita and Katsuma 2011, 98, Honshu (Mie).
Katsuma 2014, 45, Honshu (Ibaraki).
Tanida 2016, 84.
Kawase et al. 2017, 28, Honshu (Shiga).
Tanida 2018, 545, 550, male.
Morita 2019, 22, Honshu (Mie).
Nozaki et al. 2019, Honshu (Nagano).
Katsuma 2020, 15, Kyushu (Oita).
Morita 2020a, 43, Honshu (Mie).
Morita 2020c, 81, Honshu (Mie).
Yashiro et al. 2020, 185, Koshiki-jima.

Type locality. Honshu (Shizuoka?), Japan.
Distribution. Japan (Honshu, Kyushu, Koshiki-jima).
Remarks. Katsuma (2016) suggested that it is necessary to review the taxonomy of this species.

Psychomyia armata Schmid 1964

Psychomyia armata Schmid 1964, 829-831, male, Honshu (Miyagi).

Yamanouchi 1987, 35, Hokkaido (Tokachi).
Tanida 2005, 466, 468, male.
Ito and Kosugi 2007, 52, female, Hokkaido (Kushiro).
Katsuma 2007, 17, Honshu (Tochigi).
Katsuma 2011c, 69, Honshu (Ibaraki).
Ito 2013, 54, Hokkaido (Tokachi, Kushiro).
Ijima and Tsuchiya 2014, 28, Hokkaido (Okhotsk, Kushiro).
Katsuma 2014, 45, Honshu (Ibaraki).
Ito and Kuhara 2016, 16, Hokkaido (Kushiro).
Katsuma 2016, 47, Honshu (Ibaraki).
Tanaka 2016, 2, 4, male, Honshu (Akita).
Tanida 2016, 85.
Ito 2017b, 41, Hokkaido (Kamikawa).
Tanida 2018, 545, 550, male, female.
Nozaki et al. 2019, 169, Honshu (Nagano).
Kuhara et al. 2023, 17, Honshu (Yamanashi).

Type locality. Honshu (Miyagi), Japan.
Distribution. Japan (Hokkaido, Honshu), Russia (Sakhalin).

Psychomyia billnis (Kobayashi 1987)

Psychomyiella billnis Kobayashi 1987, 15, 30, Pl. 1-3, male, Honshu (Shimane).

Morita 1999a, 15, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).

Psychomyia billnis: Nozaki and Nakamura 2002, 172, Honshu (Hiroshima).

Tanida 2005, 466.
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Nozaki and Nakamura 2007, 94, Honshu (Hiroshima).
Inazu 2008, 74, Honshu (Hyogo).
Kameyama et al. 2009, 111, Honshu (Hiroshima).
Kawase and Hayashi 2011b, 172, Honshu (Shimane).
Kawase and Hayashi 2011c, 191-192, Honshu (Shimane).
Morita and Katsuma 2011, 98, Honshu (Mie).
Tanida 2016, 85.
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 550, male.
Katsuma 2020, 15, Kyushu (Oita).
Kuhara et al. 2023, 17, Kyushu (Ôita).

Type locality. Honshu (Shimane), Japan.
Distribution. Japan (Honshu, Kyushu).

Psychomyia curvicacumen Nishimoto 2020

Psychomyia curvicacumen Nishimoto 2020, 107-108, male, Honshu (Fukui, Gifu, Shiga).
Type locality. Honshu (Fukui), Japan.
Distribution. Japan (Honshu).

Psychomyia flavida Hagen 1861

Psychomyia flavida Hagen 1861, 294, Canada, America (Washington).

Fischer 1962, 220.
Fischer 1972, 89.
Ito et al. 2000b, 21-22, male, Hokkaido (Souya).
Tanida 2005, 466, 468, male.
Tanida 2016, 85.
Tanida 2018, 545, 550, male.

Type locality. Canada (precise locality unknown).
Distribution. Japan (Hokkaido), Russia (Siberia, South Primorye), Mongolia, Canada, North America.

Psychomyia incisa Nishimoto 2020

Psychomyia sp.: Torii and Hattori 2006, 36, Honshu (Shizuoka). Mentioned by Nishimoto (2020).
Psychomyia incisa Nishimoto 2020, 108-109, male, Honshu (Shizuoka, Aichi).
Type locality. Honshu (Shizuoka), Japan.
Distribution. Japan (Honshu).

Psychomyia morisitai Tsuda 1942

Psychomyia morisitai Tsuda 1942a, 273-274, male, Honshu (Nagano), Shikoku (Tokushima).

Kobayashi 1959, 348, Honshu (Gunma).
Fischer 1972, 90.
Gyotoku and Nozaki 1991, 73, Kyushu (fukuoka).
Kuranishi and Kuhara 1994, 1226, Hokkaido (Kushiro).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Kuhara 1997c, 58, Hokkaido (Shiribeshi).
Moriya 1997, 60, Honshu (Kanagawa).
Nozaki 1997, 35, Honshu (Kanagawa).
Kagaya et al. 1998, 26, 53, 166, Honshu (Tokyo).
Morita 1998, 13, Honshu (Mie).
Ito et al. 1999, 27, Hokkaido (Kushiro).
Ito et al. 2000a, 21, Hokkaido (Eniwa).
Kuhara et al. 2000, 244-245, Hokkaido (Iburi).
Kuhara 2001, 15, Hokkaido (Shiribeshi).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 172, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Ito et al. 2004, 10, Hokkaido (Kushiro).
Nozaki 2004, 1231, Honshu (Kanagawa).
Ishiwata et al. 2005, 242-244, Honshu (Kanagawa).
Nagayasu et al. 2005, 52, Hokkaido (Ishikari).
Tanida 2005, 466, 468, male.
Ito and Kosugi 2007, 53, Hokkaido (Kushiro).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Nozaki and Tanida 2007, 247, Honshu (Shizuoka).
Inazu 2008, 74, Honshu (Hyogo).
Morita 2008a, 5, Honshu (Mie).
Aoyagi & Iwasaki 2009, 101 Honshu (Osaka).
Aoyagi & Iwasaki 2010, 37, Honshu (Osaka)
Ito et al. 2010, 59, Hokkaido (Oshima).
Kawase and Morita 2010, 40, Honshu (Mie).
Inazu and Nishida 2011, 187, male, Honshu (Hyogo).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Katsuma 2011b, 67, Honshu (Ibaraki).
Morita 2011, 215, Honshu (Mie).
Katsuma 2012, 64, Honshu (Ibaraki).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Hayashi et al. 2013, 13, Oki (Dogo).
Ito 2013, 54, Hokkaido (Tokachi, Kushiro).
Ijima and Tsuchiya 2014, 28, Hokkaido (Okhotsk, Kushiro).
Kawase and Morita 2014, 5, Honshu (Shiga).
Shimura et al. 2014, 44, Amami-oshima.
Katsuma 2015, 91, Honshu (Ibaraki).
Ito and Kuhara 2016, Hokkaido (Okhotsk, Kamikawa).
Malicky 2016, 25-26, male, Taiwan (Ilan, Taipei).
Nakatani 2016, 240, Hokkaido (Shibetsu).
Nozaki 2016, 73, 335, 433, 450, male, female, Shikoku (Kochi).
Tanida 2016, 85.
Ito 2017a, 4, Honshu (Fukui).
Ito 2017b, 41, Hokkaido (Kamikawa).
Kawase et al. 2017, 28, Honshu (Shiga).
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 550, male.
Nakatani and Ito 2019, 162, Hokkaido (Kushiro).
Morita 2020a, 43, Honshu (Mie).
Morita 2020b, 45, Honshu (Mie).
Yamamoto 2023, 32, Shikoku (Ehime).

Type locality. Honshu (Nagano), Shikoku (Tokushima), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki, Amami-oshima), Russia (Sakhalin), Taiwan.

Psychomyia nipponica Tsuda 1942

Psychomyia nipponica (nomen nudum): Tsuda 1941, 104, 107, Honshu (Nagano).
Psychomyia nipponica Tsuda 1942a, 272-273, male, Honshu (Kyoto, Nagano).

Tsuda 1942b, 263-264, Honshu (Kyoto).
Fischer 1972, 90.
Morita 1996a, 50, Honshu (Mie).
Moriya 1997, 61, Honshu (Kanagawa).
Kagaya et al. 1998, 26, 53, 167, Honshu (Tokyo).
Nakano and Tanida 1999, 276, Honshu (Nara).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 338-339, Honshu (Mie).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Nozaki and Nakamura 2002, 172, Honshu (Hiroshima).
Nakamura and Umemori 2003, 59, Honshu (Hiroshima).
Nozaki 2004, 1231, Honshu (Kanagawa).
Ishiwata et al. 2005, 242-244, Honshu (Kanagawa).
Tanida 2005, 466, 468, male.
Kameyama et al. 2006, 93, Honshu (Hiroshima).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Katsuma 2007, 17, Honshu (Tochigi).
Nozaki and Tanida 2007, 247, Honshu (Shizuoka).
Morita 2008d, 17, Honshu (Mie).
Mano 2010, 22-23, Honshu (Aichi).
Ito et al. 2010, 59, male, female, Hokkaido (Oshima).
Kawase and Morita 2010, 40, Honshu (Mie).
Morita 2011, 215, Honshu (Mie).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Katsuma 2015, 91, Honshu (Fukushima).
Katsuma 2016, 47, Honshu (Ibaraki).
Tanida 2016, 85.
Kawase et al. 2017, 28, Honshu (Shiga).
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 545, 550, male.
Morita 2019, 66, Honshu (Mie).
Morita 2020b, 45, Honshu (Mie).
Nishimoto 2020, 104-106, male, Hokkaido (Oshima), Honshu (Niigata, Yamanashi, Gifu, Shizuoka, Aichi, Mie, Nara).
Morita 2020c, 81, Honshu (Mie).
Tanaka 2022, 35, Honshu (Akita).
Kuhara et al. 2023, 17, Kyushu (Kumamoto).

Psychomyiella nipponica Tsuda and Tatsumi 1951, 29, Honshu (Niigata).
Type locality. Honshu (Kyoto, Nagano), Japan.
Distribution. Japan (Hokkaido, Honshu, Kyushu).

Psychomyia pseudonipponica Nishimoto 2020

Psychomyia pseudonipponica Nishimoto 2020, 106-107, male, Shikoku (Tokushima, Ehime, Kochi).

Yamamoto 2023, 32, Shikoku (Ehime).

Type locality. Shikoku (Kochi), Japan.
Distribution. Japan (Shikoku).

Psychomyia quadridentata Nishimoto 2020

Psychomyia quadridentata Nishimoto 2020, 110, male, Honshu (Mie).

Nojima 2022, 38, Honshu (Okayama).

Type locality. Honshu (Mie), Japan.
Distribution. Japan (Honshu).

Psychomyia serrata Nishimoto 2020

Psychomyia serrata Nishimoto 2020, 109-1108, male, Honshu (Wakayama), Shikoku (Tokushima, Kochi).
Type locality. Shikoku (Tokushima), Japan.
Distribution. Japan (Honshu, Shikoku).

Psychomyia usuguronis (Matsumura 1931)

Hydroptila usuguronis Matsumura 1931, 1136, female, Hokkaido, Honshu. Transferred to the genus Psychomyia by Ito et al. (2011).

Tanida 2016, 85.

Type locality. Hokkaido (Sapporo?), Japan.
Distribution. Japan (Hokkaido, Honshu).

Psychomyia sp. PA

Psychomyia sp. PA: Akagi 1959, 40, larva, male genitalia, Honshu (Kyoto).

Tsuda and Akagi 1962, 130-131, larva.

Remarks. Although Tanida (1985) treated this species as the same species as Melanotrichia kibuneana (Tsuda 1942), but male genitalia illustrated by Akagi (1959) are apparently different from those of M. kibuneana.

Psychomyia sp. PB

Psychomyia sp. PB: Akagi 1959, 41, larva, Honshu (Kyoto).

Tsuda 1962, 130-131, larva.
Tanida 1985, 175, larva.
Tanida 2016, 85.

Subfamily Tinodinae Li & Morse

Two genera belonging to this subfamily have been recorded from Japan.

Genus Lype McLachlan

One species belonging to this genus has been recorded from Japan.

Lype excisa Mey 1991

Lype excisa Mey 1991, 194-195, male, Russia (Sakhalin).

Kuranishi et al. 1992, 448, Hokkaido (Kushiro).
Kuhara and Ito 1994, 19, 23, Hokkaido (Ishikari).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Kuhara 1997a, 30, Hokkaido (Shiribeshi).
Kuhara 1997b, 94, Hokkaido (Shiribeshi).
Kuhara 1997c, 58, Hokkaido (Shiribeshi).
Nakajima et al. 1997, 13, 18-19, Hokkaido (Oshima).
Ito et al. 1998, 14, Hokkaido (Nemuro).
Ito et al. 1999, 27, Hokkaido (Kushiro).
Ito et al. 2000b, 21, Hokkaido (Souya).
Kuhara et al. 2000, 244, Hokkaido (Iburi).
Kuhara 2001, 15, Hokkaido (Shiribeshi).
Nishimoto and Morita 2001, 76, Honshu (Aichi).
Ito et al. 2004, 10, Hokkaido (Kushiro).
Tanida 2005, 466.
Ito et al. 2007, 148, Hokkaido (Kushiro).
Ito and Kosugi 2007, 52, Hokkaido (Kushiro).
Nakano et al. 2007, 40, Hokkaido (Nemuro).
Inazu 2008, 73, Honshu (Hyogo).
Ito et al. 2010, 59, Hokkaido (Oshima).
Inazu and Nishida 2011, 186, male, female, Honshu (Hyogo).
Ito and Kubo 2011, 7, Hokkaido (Tokachi).
Kuhara 2011, 51, 56, Hokkaido (Ishikari, Sorachi, Hidaka).
Inazu 2012, 6, Honshu (Hyogo).
Katsuma 2012, 64, Honshu (Ibaraki).
Ito 2013, 54, Hokkaido (Abashiri, Tokachi, Kushiro).
Ijima and Tsuchiya 2014, 28, Hokkaido (Kushiro).
Morita 2015, 17, Honshu (Mie).
Nozaki 2016, 333-334, 433, male, female, wings.
Tanida 2016, 85.
Kuhara and Ito 2017, 15, Yaku-shima.
Ito 2017b, 41, Hokkaido (Kamikawa).
Nojima 2017, 116, Honshu (Okayama).
Tanida 2018, 546, 550, male.
Nakatani and Ito 2019, 162, Hokkaido (Kushiro).
Yamamoto et al. 2021, 72, Shikoku (Ehime).
Kawase and Kato 2023, 6, Honshu (Niigata).

Type locality. Sakhalin, Russia.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Yaku-shima), Russia (Sakhalin, Kuriles).

Genus Tinodes Curtis

Five species belonging to this genus have been recorded from Japan.

Tinodes aonensis Kobayashi 1984

Tinodes aonensis Kobayahsi 1984a, 2-3, male, Honshu (Kanagawa).

Hatta and Nozaki 1991, 199, Honshu (Gifu) (as Tinodes aonesis).
Morita 1996b, 301, Honshu (Mie).
Ito et al. 1997, 23, 27, Hokkaido (Oshima).
Moriya 1997, 61, Honshu (Kanagawa).
Morita 2001b, 339, Honshu (Mie).
Nozaki 2004, 1231, Honshu (Kanagawa).
Torii and Hattori 2006, 36, male, Honshu (Shizuoka).
Morita 2007b, 14, Honshu (Mie) (as Tinodes aoensis).
Aoyagi and Iwasaki 2009, 101, Honshu (Osaka).
Ito et al. 2010, 59, Hokkaido (Oshima).
Kawase and Morita 2010, 40, Honshu (Mie).
Tanida 2016, 85.
Nojima 2017, 117, Honshu (Okayama).
Tanaka 2018, 8, 11, male, Honshu (Akita).
Tanida 2018, 546 (as Tinodes aoensis).

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Hokkaido, Honshu).

Tinodes ashigaranis Kobayashi 1971

Tinodes ashigaranis Kobayashi 1971a, 3-4, Pl. 3, male, Honshu (Kanagawa).

Kobayashi 1971b, 29-30, Honshu (Kanagawa).
Morita 1996b, 301, Honshu (Mie) (ashigaraensis).
Nozaki 1997, 35, Honshu (Kanagawa).
Kagaya et al. 1998, 26, 54, 169, Honshu (Tokyo).
Morita 2001a, 220, Honshu (Mie).
Morita 2001b, 339, Honshu (Mie) (ashigaraensis).
Morita 2003, 25, Honshu (Mie).
Kawase et al. 2004, 112, Honshu (Ibaraki).
Nozaki 2004, 1231, Honshu (Kanagawa).
Mey and Nozaki 2006, 24, Honshu (Nagano).
Torii and Hattori 2006, 36, male, Honshu (Shizuoka).
Morita 2007b, 14, Honshu (Mie) (ashigaraensis).
Morita 2008b, 6, Honshu (Mie).
Morita 2009, 139, Honshu (Mie).
Ito et al. 2010, 59, Hokkaido (Oshima).
Kawase and Morita 2010, 40, Honshu (Mie).
Inazu and Nishida 2011, 187, male, Honshu (Hyogo).
Nozaki 2016, 73, 335-336, 433, 450, male, Shikoku (Kochi).
Tanida 2016, 86.
Tanida 2018, 546.
Shimura and Imasaka 2020, 13, Kyushu (Fukuoka).
Yamamoto et al. 2021, 73, Shikoku (Ehime).
Nojima 2022, 38, Honshu (Okayama).

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).

Tinodes higashiyamanus Tsuda 1942

Tinodes higashiyamanus Tsuda 1942a, 271-272, male, Honshu (Kyoto) (higashiyamana).

Tsuda and Tatsumi 1951, 29, Honshu (Niigata) (higashiyamana).
Fischer 1972, 78 (higashiyamana).
Tanida 1985, 175 (higashiyamana).
Morita 1995, 82, male, Honshu (Mie) (higashiyamana).
Akagi 1970, 26, 27, larva, Honshu (Okayama) (higashiyamana).
Gyotoku and Nozaki 1991, 73, Kyushu (Fukuoka) (higashiyamana).
Ito et al. 1997, 23, 27, Hokkaido (Oshima) (higashiyamana).
Nozaki 1997, 35, Honshu (Kanagawa) (higashiyamana).
Kagaya et al. 1998, 26, 54, 170, Honshu (Tokyo, Yamanashi) (higashiyamana).
Morita 2000, 60, Honshu (Mie) (higashiyamana).
Morita 2001b, 339, Honshu (Mie) (higashiyamana).
Nozaki 2004, 1231, Honshu (Kanagawa) (higashiyamana).
Kawase et al. 2004, 112, Honshu (Ibaraki) (higashiyamana).
Tanida 2005, 466, 468, male (higashiyamana).
Mey and Nozaki 2006, 24, Honshu (Nagano) (higashiyamana).
Torii and Hattori 2006, 36-37, Honshu (Shizuoka) (higashiyamana).
Nozaki and Tanida 2007, 247, Honshu (Shizuoka) (higashiyamana).
Morita 2008b, 6, Honshu (Mie) (higashiyamana).
Morita 2008c, 92, Honshu (Mie) (higashiyamana).
Aoyagi and Iwasaki 2009, 101, Honshu (Osaka) (higashiyamana).
Aoyagi and Iwasaki 2010, 37, Honshu (Osaka).
Ito et al. 2010, 59, Hokkaido (Oshima).
Kawase and Morita 2010, 40, Honshu (Mie).
Mano 2010, 22-23, Honshu (Aichi) (higashiyamana).
Nozaki 2010, 216, Honshu (Kanagawa).
Shimura 2010, 49, Yonaguni-to (Okinawa).
Inazu and Nishida 2011, 187, male, Honshu (Hyogo) (higashiyamana).
Morita 2011, 215, Honshu (Mie) (higashiyamana).
Inazu 2012, 6, Honshu (Hyogo) (higashiyamana).
Moriya and Kubota 2012, 133, Honshu (Kanagawa).
Hayashi et al. 2013, 13, Oki (Dogo) (higashiyamana).
Nozaki 2016, 73, 336, 433, 450, male, Shikoku (Ehime).
Kuhara and Ito 2017, 15, Yaku-shima.
Kawase et al. 2017, 28, Honshu (Shiga).
Hayashi et al. 2017, 79, Kyushu (Miyazaki).
Ito 2017b, 41, Hokkaido (Kamikawa).
Nojima 2017, 117, Honshu (Okayama).
Oláh et al. 2018, 106, North Korea (Kumgangsan).
Nozaki 2018, 928, Honshu (Kanagawa).
Tanida 2018, 546, 550, male.
Nozaki et al. 2019, 168-169, Honshu (Nagano).
Shimura and Imasaka 2020, 13, Kyushu (Fukuoka).
Shimura and Imasaka 2021, 4, Koshiki-jima.
Yamamoto et al. 2021, 73, Shikoku (Ehime).

Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Koshiki-jima, Yaku-shima, Yonaguni), North Korea.

Tinodes miyakonis Tsuda 1942

Tinodes miyakonis Tsuda 1942a, 270-271, male, Honshu (Kyoto).

Tsuda 1942b, 263-264, Honshu (Kyoto).
Fischer 1972, 79.
Kobayashi 1985, 10, Tsushima.
Gyotoku and Nozaki 1991, 73, Kyushu (Fukuoka).
Nozaki and Nakamura 2002, 172, Honshu (Hiroshima).
Nakamura et al. 2003, 153, Honshu (Hiroshima).
Tanida 2005, 466, 468, male, wings.
Morita 2007a, 82, Honshu(Mie)
Aoyagi and Iwasaki 2010, 37, Honshu (Osaka).
Kawase and Morita 2010, 40, Honshu (Mie).
Nakamura 2010, 239, Honshu (Hiroshima).
Inazu and Nishida 2011, 187, male, Honshu (Hyogo).
Kawase and Hayashi 2011a, 80, Honshu (Shimane).
Kawase and Hayashi 2011b, 172, Honshu (Shimane).
Kawase and Hayashi 2011c, 192, Honshu (Shimane).
Morita 2011, 215, Honshu (Mie).
Inazu 2013a, 56, Honshu (Hyogo).
Inazu 2013b, 71, Honshu (Hyogo).
Tanida 2016, 86.
Nojima 2017, 117, Honshu (Okayama).
Tanida 2018, 546, 549-550, male, wings.
Yamamoto et al. 2019, 71, Shikoku (Ehime).
Shimura and Imasaka 2021, 4, Koshiki-jima.
Yamamoto and Nozaki 2021, 128, Shikoku (Ehime).
Kuhara et al. 2023, 17, Kyushu (Kagoshima).

Tinodes sauteri Tsuda 1939, 209-211, larva, case, Honshu (Kyoto). Misidentification mentioned by Tsuda (1942c).
Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu, Shikoku, Kyushu, Tsushima, Koshiki-jima).

Tinodes sauteri Ulmer 1908

Tinodes sauteri Ulmer 1908, 352-353, male, Honshu (Kanagawa).

Tsuda 1942a, 270.
Fischer 1962, 202.
Fischer 1972, 83.
Tanida 2016, 86.
Tanida 2018, 546.

Type locality. Honshu (Kanagawa), Japan.
Distribution. Japan (Honshu).

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